198 lines
22 KiB
XML
198 lines
22 KiB
XML
<document id="8EE8ABB9F9A6CB17880AFE7A11F26B8C" ID-CLB-Dataset="5563" ID-DOI="10.11646/zootaxa.4990.3.1" ID-GBIF-Dataset="6a6dd89d-6b74-47ed-ba8e-71b526355b69" ID-ISSN="1175-5326" ID-Zenodo-Dep="5026777" ID-ZooBank="70610FEE-4497-4804-817C-CEC2D66DDBFE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624542810433" checkinUser="plazi" docAuthor="Maxwell, Stephen J., Rymer, Tasmin L. & Congdon, Bradley C." docDate="2021" docId="039B8783874AFFE1FF1EF6A613CEFE1A" docLanguage="en" docName="zootaxa.4990.3.1.pdf" docOrigin="Zootaxa 4990 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Seraphs squamosus" docType="treatment" docVersion="6" lastPageNumber="418" masterDocId="FFA2FFFB875AFFF0FF89F03A132EFF92" masterDocTitle="Resolving phylogenetic and classical nomenclature: A revision of Seraphsidae Jung, 1974 (Gastropoda: Neostromboidae)" masterLastPageNumber="453" masterPageNumber="401" pageNumber="417" updateTime="1699020302559" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7B6F6CE2A33FA888DD25279886247B22">Resolving phylogenetic and classical nomenclature: A revision of Seraphsidae Jung, 1974 (Gastropoda: Neostromboidae)</mods:title>
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<mods:namePart id="C0AE041B9BA8E1B836A5C48C397B1B3C">Maxwell, Stephen J.</mods:namePart>
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<mods:affiliation id="A99F234F622D724862AD7D0313B721C4">College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. & Stephen. maxwell @ my. jcu. edu. au; https: // orcid. org / 0000 - 0002 - 4328 - 6171</mods:affiliation>
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<mods:namePart id="C530580CF9886F53A65C41A82EC8BCB9">Rymer, Tasmin L.</mods:namePart>
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<mods:affiliation id="EA4D934B5B340C8223A54872FFF51BAE">College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. & Centre for Tropical Environmental and Sustainability Sciences, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. & Tamsin. rymer @ jcu. edu. au; https: // orcid. org / 0000 - 0002 - 9963 - 6345</mods:affiliation>
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<mods:namePart id="33A8D69881EB2346E0648D5D09382E61">Congdon, Bradley C.</mods:namePart>
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<mods:affiliation id="5889A9BFBB93E75E1F5896ABCB30152D">College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. & Centre for Tropical Environmental and Sustainability Sciences, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. & brad. congdon @ jcu. edu. au; https: // orcid. org / 0000 - 0002 - 8751 - 0892</mods:affiliation>
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<mods:nameIdentifier id="BC18203A9796E959344015ABAE658B73" type="email">brad.congdon@jcu.edu.au</mods:nameIdentifier>
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<treatment id="039B8783874AFFE1FF1EF6A613CEFE1A" ID-DOI="http://doi.org/10.5281/zenodo.5026810" ID-GBIF-Taxon="183411034" ID-Zenodo-Dep="5026810" LSID="urn:lsid:plazi:treatment:039B8783874AFFE1FF1EF6A613CEFE1A" httpUri="http://treatment.plazi.org/id/039B8783874AFFE1FF1EF6A613CEFE1A" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417">
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<subSubSection id="C328651E874AFFE0FF1EF6A6116AF924" box="[151,580,1692,1719]" pageId="16" pageNumber="417" type="nomenclature">
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<heading id="D0C581F9874AFFE0FF1EF6A6116AF924" bold="true" box="[151,580,1692,1719]" fontSize="11" level="1" pageId="16" pageNumber="417" reason="1">
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<taxonomicName id="4C324D16874AFFE0FF1EF6A6116AF924" ID-CoL="cb157e9c-d109-41d3-b2cd-6a05d401e182" authority="(Martin, 1914)" baseAuthorityName="Martin" baseAuthorityYear="1914" box="[151,580,1692,1719]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="417" phylum="Mollusca" rank="species" species="squamosus">
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<emphasis id="B946EA87874AFFE0FF1EF6A6116AF924" bold="true" box="[151,580,1692,1719]" pageId="16" pageNumber="417">
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<emphasis id="B946EA87874AFFE0FF1EF6A612A8F925" bold="true" box="[151,390,1692,1719]" italics="true" pageId="16" pageNumber="417">Seraphs squamosus</emphasis>
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(
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<bibRefCitation id="EFA34B64874AFFE0FE1CF6A61115F924" author="Martin, K." box="[405,571,1692,1718]" pageId="16" pageNumber="417" pagination="107 - 178" refId="ref38146" refString="Martin, K. (1914) Des Obereocans von Nanggulan, auf Java. Sammlungen des Geologischen Reishs-Museums in Leiden Neue Folge, 2, 107 - 178. https: // doi. org / 10.5962 / bhl. title. 50429" type="journal article" year="1914">Martin, 1914</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C328651E874AFFE0FF1EF6FB1227F94E" box="[151,265,1729,1756]" pageId="16" pageNumber="417" type="description">
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<paragraph id="8B8D3695874AFFE0FF1EF6FB1227F94E" blockId="16.[151,580,1692,1756]" box="[151,265,1729,1756]" pageId="16" pageNumber="417">
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<figureCitation id="13092A10874AFFE0FF1EF6FB1227F94E" box="[151,265,1729,1756]" captionStart="FIGURE 5" captionStartId="9.[151,250,1632,1656]" captionTargetBox="[169,1422,191,1598]" captionTargetId="figure-19@9.[151,1435,181,1609]" captionTargetPageId="9" captionText="FIGURE 5. Seraphs Montfort, 1810. A. Seraphs minus (Vincent, 1913), length 7.1 mm (Jung 1974, pl. 6, figs. 9, 10). B. Seraphs naricus (Vredenberg, 1925), holotype, GSI 12614, length 38.6 mm (Jung 1974, pl. 6, figs. 20–22). C. Seraphs olivaceus (Cossmann, 1889), MNHN A28577, Collection de Paleontologie, Paris, length 32 mm (Caze et al. 2010, figs. 6H, I). D. Seraphs peterjungi Caze et al., 2010, holotype, MNHN A28578, Collection de Paleontologie, length 16 mm (Caze et al. 2010, figs. 15J, K, N). E. Seraphs placiferus (Bayan, 1870), lectotype, MNHN G2100, length 33.7 mm (Jung 1974, pl. 7, figs. 5–7). F. Seraphs sopita (Brander, 1766), lectotype, NHMUK GG21010, length 26.4 mm (Jung 1974, pl. 1, figs. 1–3). G. Seraphs squamosus (Martin, 1914), Lectotype, RGM, Leiden St10344, length 37.1 mm (Jung 1974, pl. 6, figs. 17–19). H. Seraphs striatus (von Koenen, 1889), holotype, length 38.5 mm (von Koenen 1889, pl. I, figs. 7A–C). I. Seraphs subconvolutus (d’Orbigny, 1852), lectotype, MNHN UBT65-2-120, length 28 mm (Caze et al. 2010, figs. 14A, B). J. Seraphs volutatus (Brander, 1766), MNHN A28776, Collection de Paleontologie, Paris, length 17 mm (Caze et al. 2010, figs. 6A–C)." figureDoi="http://doi.org/10.5281/zenodo.5026789" httpUri="https://zenodo.org/record/5026789/files/figure.png" pageId="16" pageNumber="417">Figure 5G</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C328651E874AFFE0FF1EF73F1790F8AE" pageId="16" pageNumber="417" type="reference_group">
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<paragraph id="8B8D3695874AFFE0FF1EF73F1790F8AE" blockId="16.[151,1436,1797,1852]" pageId="16" pageNumber="417">
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<treatmentCitationGroup id="AB2211BB874AFFE0FF1EF73F1790F8AE" pageId="16" pageNumber="417">
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||
1914
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<taxonomicName id="4C324D16874AFFE0FF58F73F1157F88F" ID-CoL="1b37f155-80e4-4cad-9221-08be03493f29" authority="Martin" authorityName="Martin" authorityYear="1914" box="[209,633,1797,1821]" class="Gastropoda" family="Seraphsidae" genus="Terebellum" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="417" phylum="Mollusca" rank="species" species="squamosum" subGenus="Seraphs">
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<emphasis id="B946EA87874AFFE0FF58F73F126FF88F" box="[209,321,1797,1821]" italics="true" pageId="16" pageNumber="417">Terebellum</emphasis>
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(
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<emphasis id="B946EA87874AFFE0FED8F73F128DF88F" box="[337,419,1797,1821]" italics="true" pageId="16" pageNumber="417">Seraphs</emphasis>
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)
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<emphasis id="B946EA87874AFFE0FE3BF73C1104F88F" box="[434,554,1798,1821]" italics="true" pageId="16" pageNumber="417">squamosum</emphasis>
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Martin
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</taxonomicName>
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, p. 159, pl. 5, figs. 125, 126.
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<treatmentCitation id="0A931084874AFFE0FC2EF73C177AF88F" author="Jung, P." box="[935,1108,1797,1822]" page="25" pageId="16" pageNumber="417" year="1974">
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<bibRefCitation id="EFA34B64874AFFE0FC2EF73C173DF88F" author="Jung, P." box="[935,1043,1797,1822]" pageId="16" pageNumber="417" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung 1974</bibRefCitation>
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, p. 25
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</treatmentCitation>
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. =
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||
<taxonomicName id="4C324D16874AFFE0FBFDF73F13CCF8AE" ID-CoL="1b37f155-80e4-4cad-9221-08be03493f29" authority="Martin" authorityName="Martin" authorityYear="1914" class="Gastropoda" family="Seraphsidae" genus="Terebellum" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="417" phylum="Mollusca" rank="species" species="squamosum">
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<emphasis id="B946EA87874AFFE0FBFDF73F164DF88F" box="[1140,1379,1797,1821]" italics="true" pageId="16" pageNumber="417">Terebellum squamosum</emphasis>
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Martin
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</taxonomicName>
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—
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<treatmentCitation id="0A931084874AFFE0FF75F71E1288F8AE" author="Jung, P." box="[252,422,1828,1852]" page="25" pageId="16" pageNumber="417" year="1974">
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<bibRefCitation id="EFA34B64874AFFE0FF75F71E1248F8AE" author="Jung, P." box="[252,358,1828,1852]" pageId="16" pageNumber="417" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung 1974</bibRefCitation>
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, p. 25
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||
</treatmentCitation>
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||
. =
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||
<taxonomicName id="4C324D16874AFFE0FE4CF71E106EF8AE" ID-CoL="cb157e9c-d109-41d3-b2cd-6a05d401e182" authority="Martin" authorityName="Martin" box="[453,832,1828,1852]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="417" phylum="Mollusca" rank="species" species="squamosus" subGenus="Seraphs">
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<emphasis id="B946EA87874AFFE0FE4CF71E1139F8AE" box="[453,535,1828,1852]" italics="true" pageId="16" pageNumber="417">Seraphs</emphasis>
|
||
(
|
||
<emphasis id="B946EA87874AFFE0FDACF71E1159F8AE" box="[549,631,1828,1852]" italics="true" pageId="16" pageNumber="417">Seraphs</emphasis>
|
||
)
|
||
<emphasis id="B946EA87874AFFE0FD0FF71F11DBF8AE" box="[646,757,1829,1852]" italics="true" pageId="16" pageNumber="417">squamosus</emphasis>
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||
Martin
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||
</taxonomicName>
|
||
<emphasis id="B946EA87874AFFE0FCC9F71F1079F8AE" box="[832,855,1829,1852]" italics="true" pageId="16" pageNumber="417">—</emphasis>
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<treatmentCitation id="0A931084874AFFE0FCDEF71E172EF8AE" author="Jung, P." box="[855,1024,1828,1852]" page="25" pageId="16" pageNumber="417" year="1974">
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<bibRefCitation id="EFA34B64874AFFE0FCDEF71E10ECF8AE" author="Jung, P." box="[855,962,1828,1852]" pageId="16" pageNumber="417" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung 1974</bibRefCitation>
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||
, p. 25
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||
</treatmentCitation>
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||
, pl. 6, figs. 17–19.
|
||
</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C328651E874AFFE0FF1EF7501096F85F" pageId="16" pageNumber="417" type="materials_examined">
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<paragraph id="8B8D3695874AFFE0FF1EF7501276F83A" blockId="16.[151,1436,1898,2033]" pageId="16" pageNumber="417">
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<emphasis id="B946EA87874AFFE0FF1EF750126AF816" bold="true" box="[151,324,1898,1924]" pageId="16" pageNumber="417">Type material.</emphasis>
|
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<typeStatus id="54898837874AFFE0FEC2F7501295F816" box="[331,443,1898,1924]" pageId="16" pageNumber="417" type="lectotype">Lectotype</typeStatus>
|
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—
|
||
<collectionCode id="ED23AE50874AFFE0FE5EF7501139F816" box="[471,535,1898,1924]" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="417" type="Museum">RGM</collectionCode>
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||
St10344 (
|
||
<bibRefCitation id="EFA34B64874AFFE0FD01F750102FF816" author="Jung, P." box="[648,769,1898,1924]" pageId="16" pageNumber="417" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung 1974</bibRefCitation>
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). The type measures
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<quantity id="4CCA9B70874AFFE0FC63F7501760F817" box="[1002,1102,1898,1925]" metricMagnitude="-2" metricUnit="m" metricValue="3.71" pageId="16" pageNumber="417" unit="mm" value="37.1">37.1 mm</quantity>
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in length with a width of
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<quantity id="4CCA9B70874AFFE0FAE2F75013EDF83B" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="16" pageNumber="417" unit="mm" value="13.1">13.1 mm</quantity>
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(
|
||
<bibRefCitation id="EFA34B64874AFFE0FF58F7B41265F83A" author="Jung, P." box="[209,331,1934,1960]" pageId="16" pageNumber="417" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung 1974</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8B8D3695874AFFE0FF4EF7881096F85F" blockId="16.[151,1436,1898,2033]" box="[199,952,1970,1997]" pageId="16" pageNumber="417">
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<materialsCitation id="3B5A3CC8874AFFE0FF4EF788109AF85F" ID-GBIF-Occurrence="3320081305" box="[199,948,1970,1997]" county="Kai Puru" location="Djokjakarta" municipality="Naggulan" pageId="16" pageNumber="417" specimenCount="1" stateProvince="Jawa Timur" typeStatus="holotype">
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<emphasis id="B946EA87874AFFE0FF4EF7881248F85E" bold="true" box="[199,358,1970,1996]" pageId="16" pageNumber="417">
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<typeStatus id="54898837874AFFE0FF4EF788122CF85E" box="[199,258,1970,1996]" pageId="16" pageNumber="417">Type</typeStatus>
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||
locality.
|
||
</emphasis>
|
||
<collectingCounty id="62EC4E19874AFFE0FEE5F78812FAF85F" box="[364,468,1970,1997]" pageId="16" pageNumber="417">Kai Puru</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BE9ACEF874AFFE0FE56F7881161F85E" box="[479,591,1970,1996]" pageId="16" pageNumber="417">Naggulan</collectingMunicipality>
|
||
,
|
||
<location id="8EED604E874AFFE0FDD3F78811CFF85E" LSID="urn:lsid:plazi:treatment:039B8783874AFFE1FF1EF6A613CEFE1A:8EED604E874AFFE0FDD3F78811CFF85E" box="[602,737,1970,1996]" county="Kai Puru" municipality="Naggulan" name="Djokjakarta" pageId="16" pageNumber="417" stateProvince="Jawa Timur">Djokjakarta</location>
|
||
,
|
||
<collectingRegion id="49F6F877874AFFE0FD65F7891030F85F" box="[748,798,1971,1997]" country="Indonesia" name="Jawa Timur" pageId="16" pageNumber="417">Java</collectingRegion>
|
||
[Priabonian]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C328651E874AFFE1FF4EF7EC179CFE8F" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417" type="diagnosis">
|
||
<paragraph id="8B8D3695874AFFE1FF4EF7EC179CFE8F" blockId="16.[151,1436,1898,2033]" lastBlockId="17.[151,1436,150,392]" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417">
|
||
<emphasis id="B946EA87874AFFE0FF4EF7EC126CF862" bold="true" box="[199,322,2006,2032]" pageId="16" pageNumber="417">Diagnosis.</emphasis>
|
||
The shell has a stout but acute teleoconch. The greatest diameter of the shell occurs at the adapical end of the aperture. The body whorl is slightly concave at the lower half. The columella is bent to the left and backward. The callus of the labrum reaches the apex. The labrum is slightly thickened and terminates well above the columella. As a juvenile, the shell has a slight spiral groove near the apex. The adults have distinctive sculpturing on the lower half of the body whorl. The growth lines of the shell form a reticulate spiral pattern.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C328651E874BFFE1FF4EF11C164EFED2" box="[199,1376,294,321]" pageId="17" pageNumber="418" type="distribution">
|
||
<paragraph id="8B8D3695874BFFE1FF4EF11C164EFED2" blockId="17.[151,1436,150,392]" box="[199,1376,294,321]" pageId="17" pageNumber="418">
|
||
<emphasis id="B946EA87874BFFE1FF4EF11C124FFED2" bold="true" box="[199,353,294,320]" pageId="17" pageNumber="418">Distribution.</emphasis>
|
||
EOCENE—Priabonian—
|
||
<emphasis id="B946EA87874BFFE1FD0EF11C11DBFED2" box="[647,757,294,320]" italics="true" pageId="17" pageNumber="418">
|
||
<collectingCountry id="F3257605874BFFE1FD0EF11C11DBFED2" box="[647,757,294,320]" name="Indonesia" pageId="17" pageNumber="418">Indonesia</collectingCountry>
|
||
</emphasis>
|
||
Kali Puru, Nanggulan,
|
||
<collectingRegion id="49F6F877874BFFE1FB89F11D171CFED3" box="[1024,1074,295,321]" country="Indonesia" name="Jawa Timur" pageId="17" pageNumber="418">Java</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EFA34B64874BFFE1FBC9F11C17FEFED2" author="Martin, K." box="[1088,1232,294,320]" pageId="17" pageNumber="418" pagination="107 - 178" refId="ref38146" refString="Martin, K. (1914) Des Obereocans von Nanggulan, auf Java. Sammlungen des Geologischen Reishs-Museums in Leiden Neue Folge, 2, 107 - 178. https: // doi. org / 10.5962 / bhl. title. 50429" type="journal article" year="1914">Martin 1914</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA34B64874BFFE1FB55F11D167AFED2" author="Jung, P." box="[1244,1364,294,321]" pageId="17" pageNumber="418" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung 1974</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C328651E874BFFE1FF4EF17013CEFE1A" pageId="17" pageNumber="418" type="discussion">
|
||
<paragraph id="8B8D3695874BFFE1FF4EF17013CEFE1A" blockId="17.[151,1436,150,392]" pageId="17" pageNumber="418">
|
||
<emphasis id="B946EA87874BFFE1FF4EF1701215FEF6" bold="true" box="[199,315,330,356]" pageId="17" pageNumber="418">Remarks.</emphasis>
|
||
<taxonomicName id="4C324D16874BFFE1FECAF1701131FEF6" box="[323,543,330,356]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="17" pageNumber="418" phylum="Mollusca" rank="species" species="squamosus">
|
||
<emphasis id="B946EA87874BFFE1FECAF1701131FEF6" box="[323,543,330,356]" italics="true" pageId="17" pageNumber="418">Seraphs squamosus</emphasis>
|
||
</taxonomicName>
|
||
is uniquely shaped among the
|
||
<taxonomicName id="4C324D16874BFFE1FCF4F17010F9FEF6" authorityName="Montfort" authorityYear="1810" box="[893,983,330,356]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="17" pageNumber="418" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B946EA87874BFFE1FCF4F17010F9FEF6" box="[893,983,330,356]" italics="true" pageId="17" pageNumber="418">Seraphs</emphasis>
|
||
</taxonomicName>
|
||
with its triangular spire and broad body whorl.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |