treatments-xml/data/03/94/87/039487AECA17FFB2FF67FCE2FD0DF8B7.xml
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<mods:title id="4FB99293667C673976571C2E897650EB">New data on geometroid moths (Lepidoptera: Geometroidea: Uraniidae and Geometridae) from Sakhalin and Moneron islands with notes on their taxonomy distribution and ecology</mods:title>
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<mods:namePart id="69BF23E3AD7F8098F73DD968BE6E0F24">Beljaev, Еvgeniy A.</mods:namePart>
<mods:affiliation id="0F3271EDB317F1518EBCE2771A37FCF6">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, Vladivostok, Russia.</mods:affiliation>
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<mods:namePart id="B03C6F26CCE189360DC31A239681E246">Titova, Olga L.</mods:namePart>
<mods:affiliation id="3596FAAC8D97CEB5D3BEE0D7E334AE83">Sakhalin Territory Department of Hydrometeorology and Environmental Monitoring, Sakhalinskaya oblast, Yuzhno-Sakhalinsk, Russia.</mods:affiliation>
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<subSubSection id="C3276533CA17FFB2FF67FCE2FD84FCE5" box="[151,518,782,809]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8B8236B8CA17FFB2FF67FCE2FD84FCE5" blockId="16.[151,518,782,845]" box="[151,518,782,809]" pageId="16" pageNumber="17">
<heading id="D0CA81D4CA17FFB2FF67FCE2FD84FCE5" bold="true" box="[151,518,782,809]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<taxonomicName id="4C3D4D3BCA17FFB2FF67FCE2FD84FCE5" ID-CoL="93RK3" authority="(Butler)" baseAuthorityName="Butler" baseAuthorityYear="1878" box="[151,518,782,809]" class="Insecta" family="Geometridae" genus="Trichopteryx" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hemana">
<emphasis id="B949EAAACA17FFB2FF67FCE2FD84FCE5" bold="true" box="[151,518,782,809]" pageId="16" pageNumber="17">
<emphasis id="B949EAAACA17FFB2FF67FCE2FE1EFCE4" bold="true" box="[151,412,782,809]" italics="true" pageId="16" pageNumber="17">Trichopteryx hemana</emphasis>
(Butler)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3276533CA17FFB2FF67FCDEFD86FC81" box="[151,516,818,845]" pageId="16" pageNumber="17" type="description">
<paragraph id="8B8236B8CA17FFB2FF67FCDEFD86FC81" blockId="16.[151,518,782,845]" box="[151,516,818,845]" pageId="16" pageNumber="17">
(
<figureCitation id="13062A3DCA17FFB2FF6FFCDEFEA2FC80" box="[159,288,818,845]" captionStart="FIGURES 2257" captionStartId="14.[151,264,186,210]" captionTargetId="figure-17@13.[151,1435,237,1723]" captionTargetPageId="13" captionText="FIGURES 2257. Larentiinae and Sterrhinae from Sakhalin. 22—Leptostegna tenerata, ♀, Sokhonda mount.; 23—Tyloptera bella, ♂, Sokhonda mount.; 24—Acasis exviretata, ♀, Yasnomorskoe; 25—A. viretata, ♀, Yasnomorskoe; 26—Trichopteryx carpinata, ♀, Novoaleksandrovsk; 27—T. fastuosa, ♀, S Kholmsk; 2829—T. hemana, S Kholmsk: 28—♂, 29—♀; 30—T. terranea, ♂, Novoaleksandrovsk; 31—Xanthorhoe hortensiaria, ♂, S Kholmsk; 32—Photoscotosia atrostrigata, ♀, S Kholmsk (a—upper side; b—underside); 33—Idiotephria evanescens, ♂, S Kholmsk; 34—Pennithera comis, ♀, S Kholmsk; 35—Eulithis achatinellaria, ♂, Krasnaya Tym; 36—E. testata, ♂, Sakhalin, Okhinskii district, 40 km S of Neftegorsk, near Nutovo mountain, ~ 52º37ʹ N 143º07ʹ E, 30.VIII.2000, A.V. Propletkin legit.; 37—Gandaritis evanescens, ♂, Sokhonda mount.; 38—G. pyraliata, ♂, S Kholmsk; 39—G. whitelyi, ♂, Sokhonda mount.; 40—Ecliptopera capitata capitulata, ♀, Yasnomorskoe; 41—E. pryeri, ♂, S Kholmsk; 42—E. silaceata leuca, ♂, Yasnomorskoe; 43—Dysstroma citrata, ♂, S Kholmsk; 44—Lampropteryx minna, ♀, S Kholmsk; 45—Operophtera peninsularis, ♂, Leonidovo; 46—Laciniodes denigrata, ♂, Yasnomorskoe; 47—Hydrelia shioyana, ♀, Kholmsk; 48—Venusia phasma, ♂, S Kholmsk; 49—V. semistrigata, ♀, S Kholmsk; 50—Perizoma alchemillata, ♂, S Kholmsk; 51—Gymnoscelis esakii, ♂, S Kholmsk, photo in nature by O.L. Titova; 52—Pasiphila chloerata, ♂, Yasnomorskoe; 53—Eupithecia absinthiata, ♂, Yuzhno-Sakhalinsk; 54—Idaea foedata, ♀, Razdolnoe; 55—I. imbecilla, ♂, Yasnomorskoe; 56—I. pallidata, ♂, Tymovskoe; 57—Timandra rectistrigaria, ♂, Zhitnitsa river. Scale—10 mm." figureDoi="http://doi.org/10.5281/zenodo.10211084" httpUri="https://zenodo.org/record/10211084/files/figure.png" pageId="16" pageNumber="17">Figs 28, 29</figureCitation>
,
<figureCitation id="13062A3DCA17FFB2FEDCFCDEFE7EFC81" box="[300,508,818,844]" captionStart="FIGURES 5862" captionStartId="27.[151,264,1809,1833]" captionTargetBox="[161,1417,194,1777]" captionTargetId="figure-17@27.[157,1430,181,1785]" captionTargetPageId="27" captionText="FIGURES 5862. Genitalia of Geometridae from Sakhalin. 58—Abraxas niphonibia, 5960—Acasis exviretata, 61—Trichopteryx hemana, form A, 62—T. hemana, form B. 58, 60— female genitalia, 59, 61, 62—male genitalia (a—genital segment, b—aedeagus). Scale—1 mm." figureDoi="http://doi.org/10.5281/zenodo.10147417" httpUri="https://zenodo.org/record/10147417/files/figure.png" pageId="16" pageNumber="17">61a, 61b, 62a, 62b</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3276533CA17FFB2FF67FC9BFC1AFC63" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8B8236B8CA17FFB2FF67FC9BFCA4FC42" blockId="16.[151,920,887,942]" box="[151,806,887,911]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB2D1196CA17FFB2FF67FC9BFCA4FC42" box="[151,806,887,911]" pageId="16" pageNumber="17">
<taxonomicName id="4C3D4D3BCA17FFB2FF67FC9BFEEDFC42" ID-CoL="93RK3" baseAuthorityName="Butler" baseAuthorityYear="1878" box="[151,367,887,911]" class="Insecta" family="Geometridae" genus="Trichopteryx" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hemana">
<emphasis id="B949EAAACA17FFB2FF67FC9BFEEDFC42" box="[151,367,887,911]" italics="true" pageId="16" pageNumber="17">Trichopteryx hemana</emphasis>
</taxonomicName>
:
<treatmentCitation id="0A9C10A9CA17FFB2FE8DFC9BFD2DFC42" author="Beljaev, E. A. &amp; Mironov, V. G." box="[381,687,887,911]" page="257" pageId="16" pageNumber="17" year="2019">
<bibRefCitation id="EFAC4B49CA17FFB2FE8DFC9BFD2DFC42" author="Beljaev, E. A. &amp; Mironov, V. G." box="[381,687,887,911]" pageId="16" pageNumber="17" pagination="235 - 281" refId="ref24431" refString="Beljaev, E. A. &amp; Mironov, V. G. (2019) Geometridae. In: Sinev, S. Yu. (Ed.), Catalogue of the Lepidoptera of Russia. 2 nd Edition. Zoological Institute RAS, St. Petersburg, pp. 235 - 281. [in Russian]" type="book chapter" year="2019">Beljaev &amp; Mironov 2019: 257</bibRefCitation>
</treatmentCitation>
(
<collectingRegion id="49F9F85ACA17FFB2FD4DFC9BFC9EFC42" box="[701,796,887,911]" country="Russia" name="Sakhalin" pageId="16" pageNumber="17">Sakhalin</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8236B8CA17FFB2FF67FC7AFC1AFC63" blockId="16.[151,920,887,942]" box="[151,920,917,942]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB2D1196CA17FFB2FF67FC7AFC1AFC63" box="[151,920,917,942]" pageId="16" pageNumber="17">
<taxonomicName id="4C3D4D3BCA17FFB2FF67FC7AFEE5FC60" ID-CoL="58J3V" authorityName="Yazaki" authorityYear="1978" box="[151,359,918,942]" class="Insecta" family="Geometridae" genus="Trichopteryx" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="incerta">
<emphasis id="B949EAAACA17FFB2FF67FC7AFEE5FC60" box="[151,359,918,942]" italics="true" pageId="16" pageNumber="17">Trichopteryx incerta</emphasis>
</taxonomicName>
, nec Yazaki:
<treatmentCitation id="0A9C10A9CA17FFB2FE1FFC7AFCA3FC60" author="Beljaev, E. A. &amp; Mironov, V. G." box="[495,801,917,942]" page="257" pageId="16" pageNumber="17" year="2019">
<bibRefCitation id="EFAC4B49CA17FFB2FE1FFC7AFCA3FC60" author="Beljaev, E. A. &amp; Mironov, V. G." box="[495,801,917,942]" pageId="16" pageNumber="17" pagination="235 - 281" refId="ref24431" refString="Beljaev, E. A. &amp; Mironov, V. G. (2019) Geometridae. In: Sinev, S. Yu. (Ed.), Catalogue of the Lepidoptera of Russia. 2 nd Edition. Zoological Institute RAS, St. Petersburg, pp. 235 - 281. [in Russian]" type="book chapter" year="2019">Beljaev &amp; Mironov 2019: 257</bibRefCitation>
</treatmentCitation>
(
<collectingRegion id="49F9F85ACA17FFB2FCDFFC7AFC0CFC63" box="[815,910,918,942]" country="Russia" name="Sakhalin" pageId="16" pageNumber="17">Sakhalin</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3276533CA17FFB2FF67FC30FBD2FBF3" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8B8236B8CA17FFB2FF67FC30FBD2FBF3" blockId="16.[151,1437,988,1914]" pageId="16" pageNumber="17">
<emphasis id="B949EAAACA17FFB2FF67FC30FEFDFC3B" bold="true" box="[151,383,988,1014]" pageId="16" pageNumber="17">Material examined.</emphasis>
1 ♂, “Saghalien
<collectingRegion id="49F9F85ACA17FFB2FDCAFC30FDF1FC3B" box="[570,627,988,1014]" country="United Kingdom" name="Kent" pageId="16" pageNumber="17">Cent</collectingRegion>
. Exp. Sta., Takinosawa [
<quantity id="4CC59B5DCA17FFB2FC8EFC30FC44FC3B" box="[894,966,988,1014]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="17" unit="km" value="20.0">20 km</quantity>
NW of Uzhno-Sakhalinsk, railway station Pereval,
<geoCoordinate id="EE09507FCA17FFB2FF0AFBEDFED8FBD1" box="[250,346,1025,1052]" degrees="47" direction="north" minutes="03" orientation="latitude" pageId="16" pageNumber="17" precision="925" value="47.05">47°03N</geoCoordinate>
,
<geoCoordinate id="EE09507FCA17FFB2FE95FBEDFE52FBD1" box="[357,464,1025,1052]" degrees="142" direction="east" minutes="31" orientation="longitude" pageId="16" pageNumber="17" precision="925" value="142.51666">142°31E</geoCoordinate>
]
<date id="FF831078CA17FFB2FE2FFBECFDE1FBD7" box="[479,611,1024,1050]" pageId="16" pageNumber="17" value="1934-06-13">13.VI.1934</date>
”;
<collectionCode id="ED2CAE7DCA17FFB2FD8AFBECFD08FBD7" box="[634,650,1024,1050]" pageId="16" pageNumber="17">S</collectionCode>
Kholmsk: 1 ♂*,
<specimenCount id="9D3BFD31CA17FFB2FCBDFBECFCF5FBD7" box="[845,887,1024,1050]" count="1" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
,
<date id="FF831078CA17FFB2FC76FBECFC79FBD7" box="[902,1019,1024,1050]" pageId="16" pageNumber="17" value="2016-05-22">22.V.2016</date>
; 1 ♂,
<specimenCount id="9D3BFD31CA17FFB2FBB0FBECFBE9FBD7" box="[1088,1131,1024,1050]" count="1" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
,
<date id="FF831078CA17FFB2FB89FBECFB7EFBD7" box="[1145,1276,1024,1050]" pageId="16" pageNumber="17" value="2017-06-14">14.VI.2017</date>
; 2 ♂,
<specimenCount id="9D3BFD31CA17FFB2FAB0FBECFAE9FBD7" box="[1344,1387,1024,1050]" count="2" pageId="16" pageNumber="17" type="female">2 ♀</specimenCount>
, 23, 31.V,
<date id="FF831078CA17FFB2FF29FBC8FED9FBF3" box="[217,347,1060,1086]" pageId="16" pageNumber="17" value="2018-06-01">01.VI.2018</date>
; 3 ♂,
<specimenCount id="9D3BFD31CA17FFB2FE6FFBC8FE4BFBF3" box="[415,457,1060,1086]" count="1" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
, 16, 25,
<date id="FF831078CA17FFB2FDDBFBC8FD23FBF3" box="[555,673,1060,1086]" pageId="16" pageNumber="17" value="2019-05-30">30.V.2019</date>
; 1 ♂,
<specimenCount id="9D3BFD31CA17FFB2FD14FBC8FC8CFBF3" box="[740,782,1060,1086]" count="1" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
,
<date id="FF831078CA17FFB2FCECFBC8FC10FBF3" box="[796,914,1060,1086]" pageId="16" pageNumber="17" value="2021-05-24">24.V.2021</date>
; 1 ♂,
<date id="FF831078CA17FFB2FC26FBC8FBCEFBF3" box="[982,1100,1060,1086]" pageId="16" pageNumber="17" value="2022-05-28">28.V.2022</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276533CA17FFB2FF37FBA4FEE2FB67" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8B8236B8CA17FFB2FF37FBA4FEE2FB67" blockId="16.[151,1437,988,1914]" pageId="16" pageNumber="17">
<emphasis id="B949EAAACA17FFB2FF37FBA4FEE3FBAF" bold="true" box="[199,353,1096,1122]" pageId="16" pageNumber="17">Distribution.</emphasis>
<collectingCountry id="F32A7628CA17FFB2FE98FBA4FE31FBAF" box="[360,435,1096,1122]" name="Russia" pageId="16" pageNumber="17">Russia</collectingCountry>
(S RFE: S
<collectingRegion id="49F9F85ACA17FFB2FDC6FBA4FD19FBAF" box="[566,667,1096,1122]" country="Russia" name="Sakhalin" pageId="16" pageNumber="17">Sakhalin</collectingRegion>
, S Kurils—Kunashir (
<bibRefCitation id="EFAC4B49CA17FFB2FC66FBA4FB00FBAF" author="Rybalkin, S. A. &amp; Benedek, B. &amp; Dubatolov, V. V." box="[918,1154,1096,1122]" pageId="16" pageNumber="17" pagination="13 - 32" refId="ref28740" refString="Rybalkin, S. A., Benedek, B. &amp; Dubatolov, V. V. (2022) New for the fauna of Kunashir Island moths and butterflies (Lepidoptera: Carposinidae, Zygaenidae, Tortricidae, Geometridae, Notodontidae, Erebidae, Nolidae, Noctuidae, Lycaenidae). Far Eastern Entomologist, 457, 13 - 32. https: // doi. org / 10.25221 / fee. 457.3" type="journal article" year="2022">
Rybalkin
<emphasis id="B949EAAACA17FFB2FBF5FBA5FBB5FBAF" box="[1029,1079,1096,1122]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
. 2022
</bibRefCitation>
), S Khabarovskii Krai, S Amurskaya Oblast, Primorskii Krai),
<collectingCountry id="F32A7628CA17FFB2FDCDFB80FDFFFB4B" box="[573,637,1132,1158]" name="Japan" pageId="16" pageNumber="17">Japan</collectingCountry>
(
<collectingRegion id="49F9F85ACA17FFB2FD7BFB80FD7CFB4B" box="[651,766,1132,1158]" country="Japan" name="Hokkaido" pageId="16" pageNumber="17">Hokkaido</collectingRegion>
, Honshu, Shikoku, Kyushu, Tsushima),
<collectingCountry id="F32A7628CA17FFB2FB30FB80FAD2FB4B" box="[1216,1360,1132,1158]" name="North Korea" pageId="16" pageNumber="17">North Korea</collectingCountry>
,
<collectingCountry id="F32A7628CA17FFB2FAABFB80FF5CFB67" name="South Korea" pageId="16" pageNumber="17">South Korea</collectingCountry>
, NE
<collectingCountry id="F32A7628CA17FFB2FEE6FB7CFEDFFB67" box="[278,349,1168,1194]" name="China" pageId="16" pageNumber="17">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276533CA17FFB2FF37FB58FD0DF8B7" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8B8236B8CA17FFB2FF37FB58FBD9FADB" blockId="16.[151,1437,988,1914]" pageId="16" pageNumber="17">
<emphasis id="B949EAAACA17FFB2FF37FB58FEB9FB03" bold="true" box="[199,315,1204,1230]" pageId="16" pageNumber="17">Remarks.</emphasis>
The finding of
<taxonomicName id="4C3D4D3BCA17FFB2FE01FB59FDE5FB03" baseAuthorityName="Butler" baseAuthorityYear="1878" box="[497,615,1204,1230]" class="Insecta" family="Geometridae" genus="Trichopteryx" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hemana">
<emphasis id="B949EAAACA17FFB2FE01FB59FDE5FB03" box="[497,615,1204,1230]" italics="true" pageId="16" pageNumber="17">T. hemana</emphasis>
</taxonomicName>
in Sakhalin significantly expands its distribution range to the northeast. The larvae are polyphagous on broadleaved trees, in
<collectingCountry id="F32A7628CA17FFB2FD1FFB34FCADFB3F" box="[751,815,1240,1266]" name="Japan" pageId="16" pageNumber="17">Japan</collectingCountry>
noted on
<taxonomicName id="4C3D4D3BCA17FFB2FC51FB34FC7DFB3F" box="[929,1023,1240,1266]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="genus">
<emphasis id="B949EAAACA17FFB2FC51FB34FC7DFB3F" box="[929,1023,1240,1266]" italics="true" pageId="16" pageNumber="17">Quercus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C3D4D3BCA17FFB2FBFEFB34FBFEFB3F" authorityName="Dumortier" authorityYear="1829" box="[1038,1148,1240,1266]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
),
<taxonomicName id="4C3D4D3BCA17FFB2FB60FB34FB47FB3F" box="[1168,1221,1240,1266]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="genus">
<emphasis id="B949EAAACA17FFB2FB60FB34FB47FB3F" box="[1168,1221,1240,1266]" italics="true" pageId="16" pageNumber="17">Acer</emphasis>
</taxonomicName>
(
<taxonomicName id="4C3D4D3BCA17FFB2FB25FB34FAE7FB3F" box="[1237,1381,1240,1266]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="family">Sapindaceae</taxonomicName>
) and
<taxonomicName id="4C3D4D3BCA17FFB2FF67FB10FE95FADB" box="[151,279,1276,1302]" class="Magnoliopsida" family="Hamamelidaceae" genus="Hamamelis" kingdom="Plantae" order="Saxifragales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="genus">
<emphasis id="B949EAAACA17FFB2FF67FB10FE95FADB" box="[151,279,1276,1302]" italics="true" pageId="16" pageNumber="17">Hamamelis</emphasis>
</taxonomicName>
(
<taxonomicName id="4C3D4D3BCA17FFB2FED5FB10FE6FFADB" box="[293,493,1276,1302]" class="Magnoliopsida" family="Hamamelidaceae" kingdom="Plantae" order="Saxifragales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="family">Hamamelidaceae</taxonomicName>
). In Sakhalin the moths fly from mid-May to mid-June.
</paragraph>
<paragraph id="8B8236B8CA17FFB2FF37FACCFD0DF8B7" blockId="16.[151,1437,988,1914]" pageId="16" pageNumber="17">
The report of
<taxonomicName id="4C3D4D3BCA17FFB2FE99FACDFE54FAF7" authorityName="Yazaki" authorityYear="1978" box="[361,470,1313,1338]" class="Insecta" family="Geometridae" genus="Trichopteryx" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="incerta">
<emphasis id="B949EAAACA17FFB2FE99FACDFE54FAF7" box="[361,470,1313,1338]" italics="true" pageId="16" pageNumber="17">T. incerta</emphasis>
</taxonomicName>
from Sakhalin by
<bibRefCitation id="EFAC4B49CA17FFB2FD41FACCFC5CFAF7" author="Beljaev, E. A. &amp; Mironov, V. G." box="[689,990,1312,1338]" pageId="16" pageNumber="17" pagination="235 - 281" refId="ref24431" refString="Beljaev, E. A. &amp; Mironov, V. G. (2019) Geometridae. In: Sinev, S. Yu. (Ed.), Catalogue of the Lepidoptera of Russia. 2 nd Edition. Zoological Institute RAS, St. Petersburg, pp. 235 - 281. [in Russian]" type="book chapter" year="2019">Beljaev &amp; Mironov (2019</bibRefCitation>
, 2021) is erroneous. It was based on a shabby male with the “unusual” male genitalia, which is marked with an asterisk in the Material examined. Unlike the other examined specimens of
<taxonomicName id="4C3D4D3BCA17FFB2FDE6FA85FD0EFA4F" baseAuthorityName="Butler" baseAuthorityYear="1878" box="[534,652,1384,1410]" class="Insecta" family="Geometridae" genus="Trichopteryx" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hemana">
<emphasis id="B949EAAACA17FFB2FDE6FA85FD0EFA4F" box="[534,652,1384,1410]" italics="true" pageId="16" pageNumber="17">T. hemana</emphasis>
</taxonomicName>
from Russian Far East, which have the distal projection of a valvula almost digital and the excision between this projection and the costa of valva deep (form A,
<figureCitation id="13062A3DCA17FFB2FB62FA60FB72FA6B" box="[1170,1264,1420,1446]" captionStart="FIGURES 5862" captionStartId="27.[151,264,1809,1833]" captionTargetBox="[161,1417,194,1777]" captionTargetId="figure-17@27.[157,1430,181,1785]" captionTargetPageId="27" captionText="FIGURES 5862. Genitalia of Geometridae from Sakhalin. 58—Abraxas niphonibia, 5960—Acasis exviretata, 61—Trichopteryx hemana, form A, 62—T. hemana, form B. 58, 60— female genitalia, 59, 61, 62—male genitalia (a—genital segment, b—aedeagus). Scale—1 mm." figureDoi="http://doi.org/10.5281/zenodo.10147417" httpUri="https://zenodo.org/record/10147417/files/figure.png" pageId="16" pageNumber="17">Fig. 61a</figureCitation>
), this specimen from Sakhalin has wide triangular the distal projection of a valvula, and shallow the excision between this projection and the costa of valva, and somewhat shorter distal process of sacculus (form B,
<figureCitation id="13062A3DCA17FFB2FBFFFA38FBECFA23" box="[1039,1134,1492,1518]" captionStart="FIGURES 5862" captionStartId="27.[151,264,1809,1833]" captionTargetBox="[161,1417,194,1777]" captionTargetId="figure-17@27.[157,1430,181,1785]" captionTargetPageId="27" captionText="FIGURES 5862. Genitalia of Geometridae from Sakhalin. 58—Abraxas niphonibia, 5960—Acasis exviretata, 61—Trichopteryx hemana, form A, 62—T. hemana, form B. 58, 60— female genitalia, 59, 61, 62—male genitalia (a—genital segment, b—aedeagus). Scale—1 mm." figureDoi="http://doi.org/10.5281/zenodo.10147417" httpUri="https://zenodo.org/record/10147417/files/figure.png" pageId="16" pageNumber="17">Fig. 62a</figureCitation>
). This makes general shape of the genitalia similar to those of
<taxonomicName id="4C3D4D3BCA17FFB2FDEFFA15FD09F9DF" authorityName="Yazaki" authorityYear="1978" box="[543,651,1529,1554]" class="Insecta" family="Geometridae" genus="Trichopteryx" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="incerta">
<emphasis id="B949EAAACA17FFB2FDEFFA15FD09F9DF" box="[543,651,1529,1554]" italics="true" pageId="16" pageNumber="17">T. incerta</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFAC4B49CA17FFB2FD37FA14FCD5F9DF" author="Yazaki, K." box="[711,855,1528,1554]" pageId="16" pageNumber="17" pagination="111 - 113" refId="ref30843" refString="Yazaki, K. (1978) A new species of Trichopteryx from Japan (Lepidoptera: Geometridae). Transactions of the lepidopterological Society of Japan, 29, 111 - 113." type="journal article" year="1978">Yazaki 1978</bibRefCitation>
; figs 3, 4;
<bibRefCitation id="EFAC4B49CA17FFB2FC20FA14FBDAF9DF" author="Choi, S. - W." box="[976,1112,1528,1554]" pageId="16" pageNumber="17" pagination="46 - 53" refId="ref25053" refString="Choi, S. - W. (2007 a) Taxonomic study of the genus Trichopteryx Hubner (Lepidoptera: Geometridae) in Korea. Entomological Research, 37 (1), 46 - 53. https: // doi. org / 10.1111 / j. 1748 - 5967.2007.00052. x" type="journal article" year="2007">Choi 2007a</bibRefCitation>
: figs 13, 22;
<bibRefCitation id="EFAC4B49CA17FFB2FB1CFA14FAE4F9DF" author="Choi, S. - W." box="[1260,1382,1528,1554]" pageId="16" pageNumber="17" pagination="1 - 130" refId="ref25161" refString="Choi, S. - W. (2012) Arthropoda: Insecta: Lepidoptera: Geometridae. Geometrids. In: Insect fauna of Korea. Vol. 16. No. 5. National Institute of Biological Resources, Ministry of Environment, Incheon, pp. 1 - 130." type="book chapter" year="2012">Choi 2012</bibRefCitation>
: figs 154, 233;
<bibRefCitation id="EFAC4B49CA17FFB2FEFCF9F0FE4EF9FB" author="Hashimoto S." box="[268,460,1564,1590]" pageId="16" pageNumber="17" pagination="1 - 146" refId="ref25743" refString="Hashimoto S. (2021) Taxonomic study of the Japanese Trichopterygini (Lepidoptera: Geometridae: Larentiinae), with a proposal of a new tribe Heterophlebini. Japanese Journal of Systematic Entomology, Monographic Series, 6, 1 - 146." type="journal article" year="2021">Hashimoto 2021</bibRefCitation>
: fig. 69 [uncus looks to be broken off distally]). But both these forms possess almost identical aedeagus which is strictly characteristic for
<taxonomicName id="4C3D4D3BCA17FFB2FD1AF9ADFCDDF997" baseAuthorityName="Butler" baseAuthorityYear="1878" box="[746,863,1600,1626]" class="Insecta" family="Geometridae" genus="Trichopteryx" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hemana">
<emphasis id="B949EAAACA17FFB2FD1AF9ADFCDDF997" box="[746,863,1600,1626]" italics="true" pageId="16" pageNumber="17">T. hemana</emphasis>
</taxonomicName>
(
<figureCitation id="13062A3DCA17FFB2FC9DF9ACFB8EF997" box="[877,1036,1600,1626]" captionStart="FIGURES 5862" captionStartId="27.[151,264,1809,1833]" captionTargetBox="[161,1417,194,1777]" captionTargetId="figure-17@27.[157,1430,181,1785]" captionTargetPageId="27" captionText="FIGURES 5862. Genitalia of Geometridae from Sakhalin. 58—Abraxas niphonibia, 5960—Acasis exviretata, 61—Trichopteryx hemana, form A, 62—T. hemana, form B. 58, 60— female genitalia, 59, 61, 62—male genitalia (a—genital segment, b—aedeagus). Scale—1 mm." figureDoi="http://doi.org/10.5281/zenodo.10147417" httpUri="https://zenodo.org/record/10147417/files/figure.png" pageId="16" pageNumber="17">Figs 61b, 62b</figureCitation>
).
<bibRefCitation id="EFAC4B49CA17FFB2FBD0F9ACFB6EF997" author="Hashimoto S." box="[1056,1260,1600,1626]" pageId="16" pageNumber="17" pagination="1 - 146" refId="ref25743" refString="Hashimoto S. (2021) Taxonomic study of the Japanese Trichopterygini (Lepidoptera: Geometridae: Larentiinae), with a proposal of a new tribe Heterophlebini. Japanese Journal of Systematic Entomology, Monographic Series, 6, 1 - 146." type="journal article" year="2021">Hashimoto (2021)</bibRefCitation>
illustrated both these shapes of valva in the moths from
<collectingCountry id="F32A7628CA17FFB2FDACF988FD1EF9B3" box="[604,668,1636,1662]" name="Japan" pageId="16" pageNumber="17">Japan</collectingCountry>
(
<emphasis id="B949EAAACA17FFB2FD5CF988FC82F9B3" box="[684,768,1636,1662]" italics="true" pageId="16" pageNumber="17">loc. cit.</emphasis>
: fig. 14: A, E—form B; fig. 14: B—form A), whereas from the continental
<collectingCountry id="F32A7628CA17FFB2FEB3F964FE29F96F" box="[323,427,1672,1698]" name="China" pageId="16" pageNumber="17">East Asia</collectingCountry>
only form A is known (for
<collectingCountry id="F32A7628CA17FFB2FD2CF964FCA2F96F" box="[732,800,1672,1698]" name="South Korea" pageId="16" pageNumber="17">Korea</collectingCountry>
see
<bibRefCitation id="EFAC4B49CA17FFB2FCA1F964FC57F96F" author="Choi, S. - W." box="[849,981,1672,1698]" pageId="16" pageNumber="17" pagination="46 - 53" refId="ref25053" refString="Choi, S. - W. (2007 a) Taxonomic study of the genus Trichopteryx Hubner (Lepidoptera: Geometridae) in Korea. Entomological Research, 37 (1), 46 - 53. https: // doi. org / 10.1111 / j. 1748 - 5967.2007.00052. x" type="journal article" year="2007">Choi 2007a</bibRefCitation>
, fig. 11, and
<bibRefCitation id="EFAC4B49CA17FFB2FB95F964FB5FF96F" author="Choi, S. - W." box="[1125,1245,1672,1698]" pageId="16" pageNumber="17" pagination="1 - 130" refId="ref25161" refString="Choi, S. - W. (2012) Arthropoda: Insecta: Lepidoptera: Geometridae. Geometrids. In: Insect fauna of Korea. Vol. 16. No. 5. National Institute of Biological Resources, Ministry of Environment, Incheon, pp. 1 - 130." type="book chapter" year="2012">Choi 2012</bibRefCitation>
, fig. 150; and for
<collectingCountry id="F32A7628CA17FFB2FF67F940FF58F90B" box="[151,218,1708,1734]" name="China" pageId="16" pageNumber="17">China</collectingCountry>
see
<bibRefCitation id="EFAC4B49CA17FFB2FEFAF940FE36F90B" author="Xue, D. - Y. &amp; Zhu, H. - F." box="[266,436,1708,1734]" pageId="16" pageNumber="17" refId="ref30790" refString="Xue, D. - Y. &amp; Zhu, H. - F. (1999) Lepidoptera, Geometridae, Larentiinae. In: Fauna Sinica. Insecta. Vol. 15. Science Press, Beijing, xxxii + 1090 pp., 1197 figs., 25 pls." type="book" year="1999">Xue, Zhu 1999</bibRefCitation>
, fig. 135). Possibly,
<taxonomicName id="4C3D4D3BCA17FFB2FD64F941FC8AF90B" baseAuthorityName="Butler" baseAuthorityYear="1878" box="[660,776,1708,1734]" class="Insecta" family="Geometridae" genus="Trichopteryx" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hemana">
<emphasis id="B949EAAACA17FFB2FD64F941FC8AF90B" box="[660,776,1708,1734]" italics="true" pageId="16" pageNumber="17">T. hemana</emphasis>
</taxonomicName>
in the current understanding contains two “cryptic” species with the similar moths without distinct differences in the wing pattern. Of these, one is widespread in
<collectingCountry id="F32A7628CA17FFB2FB0EF93CFAEBF927" box="[1278,1385,1744,1770]" name="China" pageId="16" pageNumber="17">East Asia</collectingCountry>
, and the other one limited to
<collectingCountry id="F32A7628CA17FFB2FE5CF918FE6EF8C3" box="[428,492,1780,1806]" name="Japan" pageId="16" pageNumber="17">Japan</collectingCountry>
and the adjacent islands. However, polymorphism in the male genitalia is also not excluded, because we have not yet been able to find
<taxonomicName id="4C3D4D3BCA17FFB2FD17F8F5FCDEF8FF" baseAuthorityName="Butler" baseAuthorityYear="1878" box="[743,860,1816,1842]" class="Insecta" family="Geometridae" genus="Trichopteryx" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hemana">
<emphasis id="B949EAAACA17FFB2FD17F8F5FCDEF8FF" box="[743,860,1816,1842]" italics="true" pageId="16" pageNumber="17">T. hemana</emphasis>
</taxonomicName>
specimens that differ either in appearance or in the female genitalia, which could be associated with this male. Therefore, we retain identification of the form B as
<taxonomicName id="4C3D4D3BCA17FFB2FA78F8D1FF72F8B7" baseAuthorityName="Butler" baseAuthorityYear="1878" class="Insecta" family="Geometridae" genus="Trichopteryx" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hemana">
<emphasis id="B949EAAACA17FFB2FA78F8D1FF72F8B7" italics="true" pageId="16" pageNumber="17">T. hemana</emphasis>
</taxonomicName>
. This issue requires a separate study.
</paragraph>
</subSubSection>
</treatment>
</document>