270 lines
37 KiB
XML
270 lines
37 KiB
XML
<document id="7C0CFBD7596CE8B53CA3990D96F33BC7" ID-CLB-Dataset="20492" ID-DOI="10.5169/seals-787049" ID-GBIF-Dataset="ce0756e2-5184-4b74-8472-ab69fed7951c" ID-ISSN="2624-9170" ID-Zenodo-Dep="6285683" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637935326602" checkinUser="carolina" docAuthor="Huber, Charles & Schmidt, Joachim" docDate="2017" docId="03948799FFEDFFFAFF41FB58FE5BFD43" docLanguage="en" docName="ContribNatHist.36.1-85.pdf.imf" docOrigin="Contributions to Natural History 36" docStyle="DocumentStyle:D319E9290ED630111D2A11900DAFC11F.4:ContribNatHist.2011-.journal_article" docStyleId="D319E9290ED630111D2A11900DAFC11F" docStyleName="ContribNatHist.2011-.journal_article" docStyleVersion="4" docTitle="Nebria (Epinebriola) retingensis Huber & Schmidt, 2017, sp. nov." docType="treatment" docVersion="13" lastPageNumber="59" masterDocId="FFADFFE1FFDBFFC0FF9CFFE9FFB0FFA7" masterDocTitle="Notes on the Nebria subgenus Epinebriola K. DANIEL, 1904 with the description of Barbonebriola subgen. nov. and 13 new species from the Himalaya-Tibet orogen (Coleoptera, Carabidae, Nebriini)" masterLastPageNumber="85" masterPageNumber="1" pageNumber="55" updateTime="1699470525451" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="02435395EBBD92DA1DBC797BF3C8CEC5">Notes on the Nebria subgenus Epinebriola K. DANIEL, 1904 with the description of Barbonebriola subgen. nov. and 13 new species from the Himalaya-Tibet orogen (Coleoptera, Carabidae, Nebriini)</mods:title>
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<mods:namePart id="9ED7718EAFBE0ED18C911F66E98D1C6A">Huber, Charles</mods:namePart>
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<mods:namePart id="D0273D039D45DB5E19FCF953AF918674">Schmidt, Joachim</mods:namePart>
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<mods:typeOfResource id="5A0A7EA829258CE2486AF37EC28C8DDB">text</mods:typeOfResource>
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<mods:title id="58AE6AA268ED9E4EB69D249DA44CC44C">Contributions to Natural History</mods:title>
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<mods:part id="F366A488BC965BB733E8FD5A51D27DAC">
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<mods:date id="901CAE672439F9F98213B30F8F4AB2B9">2017</mods:date>
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<mods:number id="4E1C118992EC242A97DF61D1BD51F97B">2017-07-17</mods:number>
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<mods:detail id="16BCB9EFA6AD6DE395156576CA3669D2" type="volume">
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<mods:number id="19984CA0C2325018F00064AD4D68B647">36</mods:number>
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<mods:classification id="D66E76FC823E93199D50DDF8C5526EC6">journal article</mods:classification>
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<treatment id="03948799FFEDFFFAFF41FB58FE5BFD43" ID-DOI="http://doi.org/10.5281/zenodo.6315675" ID-GBIF-Taxon="193820244" ID-Zenodo-Dep="6315675" LSID="urn:lsid:plazi:treatment:03948799FFEDFFFAFF41FB58FE5BFD43" httpUri="http://treatment.plazi.org/id/03948799FFEDFFFAFF41FB58FE5BFD43" lastPageId="58" lastPageNumber="59" pageId="54" pageNumber="55">
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<subSubSection id="C3276504FFEDFFF6FF41FB58FCBDFB6E" box="[221,781,1201,1226]" pageId="54" pageNumber="55" type="nomenclature">
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<paragraph id="8B82368FFFEDFFF6FF41FB58FCBDFB6E" blockId="54.[221,781,1201,1226]" box="[221,781,1201,1226]" pageId="54" pageNumber="55">
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<heading id="D0CA81E3FFEDFFF6FF41FB58FCBDFB6E" box="[221,781,1201,1226]" centered="true" fontSize="12" level="2" pageId="54" pageNumber="55" reason="2">
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<emphasis id="B949EA9DFFEDFFF6FF41FB58FD19FB6D" bold="true" box="[221,681,1201,1226]" pageId="54" pageNumber="55">
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<taxonomicName id="4C3D4D0CFFEDFFF6FF41FB58FDFBFB6D" ID-CoL="8MHV8" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[221,587,1201,1226]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="retingensis" status="sp. nov." subGenus="Epinebriola">
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<emphasis id="B949EA9DFFEDFFF6FF41FB58FE99FB6D" bold="true" box="[221,297,1201,1226]" italics="true" pageId="54" pageNumber="55">Nebria</emphasis>
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(
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<emphasis id="B949EA9DFFEDFFF6FEA5FB58FE0BFB6D" bold="true" box="[313,443,1201,1226]" italics="true" pageId="54" pageNumber="55">Epinebriola</emphasis>
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)
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<emphasis id="B949EA9DFFEDFFF6FE57FB58FDFBFB6D" bold="true" box="[459,587,1201,1226]" italics="true" pageId="54" pageNumber="55">retingensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27A57E6FFEDFFF6FDCDFB58FD19FB6D" box="[593,681,1201,1226]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="13062A0AFFEDFFF6FD2AFB58FCB3FB6E" box="[694,771,1201,1225]" captionStart="Fig" captionStartId="56.[223,251,160,178]" captionTargetBox="[463,1106,153,1322]" captionTargetId="figure-104@56.[463,1106,150,1322]" captionTargetPageId="56" captionText="Fig. 28: Nebria (Epinebriola) retingensis sp. nov. Holotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6285804" httpUri="https://zenodo.org/record/6285804/files/figure.png" pageId="54" pageNumber="55">Fig. 28</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3276504FFEDFFF6FF41FB15FC03FAF9" pageId="54" pageNumber="55" type="materials_examined">
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<paragraph id="8B82368FFFEDFFF6FF41FB15FDE9FA9E" blockId="54.[221,1107,1273,1598]" pageId="54" pageNumber="55">
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<materialsCitation id="3B553CD2FFEDFFF6FF41FB15FDE9FA9E" ID-GBIF-Occurrence="3497686316" collectingDate="2015-08-01" collectionCode="cSCHM" collectorName="J. Schmidt" elevation="4650" latitude="30.265833" location="side valley S Reting" longLatPrecision="20" longitude="91.53445" pageId="54" pageNumber="55" specimenCount="1" specimenCount-male="1" stateProvince="Tibet" typeStatus="holotype">
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<typeStatus id="5486882DFFEDFFF6FF41FB15FEF1FAB3" box="[221,321,1276,1300]" pageId="54" pageNumber="55" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D3BFD06FFEDFFF6FEDBFB10FEEBFAB3" box="[327,347,1273,1300]" pageId="54" pageNumber="55" type="male">♂</specimenCount>
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:
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<collectingRegion id="49F9F86DFFEDFFF6FEF5FB15FE15FAB3" box="[361,421,1276,1300]" country="China" name="Xizang" pageId="54" pageNumber="55">Tibet</collectingRegion>
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,
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<location id="8EE26054FFEDFFF6FE31FB15FD34FAB3" LSID="urn:lsid:plazi:treatment:03948799FFEDFFFAFF41FB58FE5BFD43:8EE26054FFEDFFF6FE31FB15FD34FAB3" box="[429,644,1276,1300]" latitude="30.265833" longLatPrecision="20" longitude="91.53445" name="side valley S Reting" pageId="54" pageNumber="55" stateProvince="Tibet">side valley S Reting</location>
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,
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<geoCoordinate id="EE095048FFEDFFF6FD11FB15FCA6FAB3" box="[653,790,1276,1300]" degrees="30" direction="north" minutes="15" orientation="latitude" pageId="54" pageNumber="55" precision="15" seconds="57" value="30.265833">30°15'57" N</geoCoordinate>
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,
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<geoCoordinate id="EE095048FFEDFFF6FCBDFB15FC17FAB3" box="[801,935,1276,1300]" degrees="91" direction="east" minutes="32" orientation="longitude" pageId="54" pageNumber="55" precision="15" seconds="04" value="91.53445">91°32'04" E</geoCoordinate>
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,
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<quantity id="4CC59B6AFFEDFFF6FC2DFB15FBFEFAB3" box="[945,1102,1276,1300]" metricMagnitude="3" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="54" pageNumber="55" unit="m" value="4650.0" valueMax="4800.0" valueMin="4500.0">
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<elevation id="0010D1BCFFEDFFF6FC2DFB15FBFEFAB3" box="[945,1102,1276,1300]" metricMagnitude="3" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="54" pageNumber="55" unit="m" value="4650.0" valueMax="4800.0" valueMin="4500.0">4500–4800 m</elevation>
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</quantity>
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,
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<date id="FF83104FFFEDFFF6FF41FAC8FE8FFA9E" box="[221,319,1313,1337]" pageId="54" pageNumber="55" value="2015-08-01">
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<collectingDate id="EFC7E9A7FFEDFFF6FF41FAC8FE8FFA9E" box="[221,319,1313,1337]" pageId="54" pageNumber="55" value="2015-08-01">1.8.2015</collectingDate>
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</date>
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, leg.
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<collectorName id="26C85359FFEDFFF6FEE5FAC8FE58FA9E" box="[377,488,1313,1337]" pageId="54" pageNumber="55">J. Schmidt</collectorName>
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(
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<collectionCode id="ED2CAE4AFFEDFFF6FE68FAC8FDFFFA9E" box="[500,591,1313,1337]" pageId="54" pageNumber="55">cSCHM</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8B82368FFFEDFFF6FE9FFAAFFC03FAF9" blockId="54.[221,1107,1273,1598]" box="[259,947,1347,1374]" pageId="54" pageNumber="55">
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<materialsCitation id="3B553CD2FFEDFFF6FE9FFAAFFC03FAF9" ID-GBIF-Occurrence="3497686353" box="[259,947,1347,1374]" collectingDate="2015-08-01" collectionCode="NMBE, cSCHM" collectorName="S Reting & J. Schmidt" elevation="4650" latitude="30.265833" location="Tibet" longLatPrecision="20" longitude="91.53445" pageId="54" pageNumber="55" specimenCount="13" specimenCount-female="12" specimenCount-male="1" stateProvince="Xizang" typeStatus="paratype">
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<typeStatus id="5486882DFFEDFFF6FE9FFAAFFECCFAF9" box="[259,380,1350,1374]" pageId="54" pageNumber="55" type="paratype">Paratypes</typeStatus>
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||
:
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<specimenCount id="9D3BFD06FFEDFFF6FE18FAAFFE1DFAF9" box="[388,429,1347,1374]" pageId="54" pageNumber="55" type="male">1 ♂</specimenCount>
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||
<specimenCount id="9D3BFD06FFEDFFF6FE2FFAAFFE59FAF9" box="[435,489,1347,1374]" pageId="54" pageNumber="55" type="female">12 ♀</specimenCount>
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, same data as holotype (
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<collectionCode id="ED2CAE4AFFEDFFF6FC9EFAAFFCFFFAF9" box="[770,847,1350,1374]" collectionName="Switzerland, Bern, Naturhistorische Museums" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="54" pageNumber="55">NMBE</collectionCode>
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,
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<collectionCode id="ED2CAE4AFFEDFFF6FCC4FAAFFC1DFAF9" box="[856,941,1350,1374]" pageId="54" pageNumber="55">cSCHM</collectionCode>
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)
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3276504FFEDFFF9FE9FFA85FECBF999" lastPageId="57" lastPageNumber="58" pageId="54" pageNumber="55" type="description">
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<paragraph id="8B82368FFFEDFFF6FE9FFA85FD54FA0E" blockId="54.[221,1107,1273,1598]" pageId="54" pageNumber="55">
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Body length
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<quantity id="4CC59B6AFFEDFFF6FE13FA85FE4AFA23" box="[399,506,1388,1412]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="54" pageNumber="55" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9–11 mm</quantity>
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. Colour dark brown to black. Appendages of the head, antennae, tibiae and tarsi brownish lightened.
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</paragraph>
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<paragraph id="8B82368FFFEDFFF7FE9FFA5FFEFCFE79" blockId="54.[221,1107,1273,1598]" lastBlockId="55.[184,1069,156,1636]" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
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Head: Labrum finely punctate with anterior margin straight, bearing six setae. Clypeus with anterior margin straight. Frons transversally wrinkled. Eyes large, protruding, temples very long. One supraorbital seta. Head with a faint collar constriction. Vertex impunctate. Antennae long and slender extending at least to the middle of the elytra. Antennal scape thickened, cylindrical, shorter than the eye's diameter (or as long as the diameter), basally narrowed, with one dorsal seta. 2
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<superScript id="7C489BC7FFECFFF7FDA5FF0AFDE0FF51" attach="left" box="[569,592,227,246]" fontSize="8" pageId="55" pageNumber="56">nd</superScript>
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antennomere with one seta ventroapically, rarely with an additional dorsoapical seta (asymmetrically in the
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<typeStatus id="5486882DFFECFFF7FC5BFEE5FB9CFE83" box="[967,1068,268,292]" pageId="55" pageNumber="56" type="holotype">holotype</typeStatus>
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male and in
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<specimenCount id="9D3BFD06FFECFFF7FED5FED8FDF2FEEE" box="[329,578,305,329]" pageId="55" pageNumber="56" type="female" typeStatus="paratype">one paratype female</specimenCount>
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). Maxillar palpomeres short. Maxillary stipes laterally straight, with 5–6 setae. Penultimate labial palpomere trisetose. Ligula spiny, with two lateral setae. Mentum with bifid medial tooth. Submentum with a row of 6–10 setae. Vertex impunctate. Microreticulation of the head isodiametric.
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</paragraph>
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<paragraph id="8B82368FFFECFFF7FF41FE05FE14FB03" blockId="55.[184,1069,156,1636]" pageId="55" pageNumber="56">
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Pronotum slender, ratio width/length of the pronotum = 1.23. Lateral margin faintly convex, slightly narrowed basally (ratio apical width/basal width = 1.12), faintly rounded to the anterior and hardly rounded to the posterior angles, straight in front of the posterior angles (
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<figureCitation id="13062A0AFFECFFF7FD47FDB5FC8FFDD3" box="[731,831,604,628]" captionStart="Fig" captionStartId="61.[184,213,1317,1335]" captionTargetBox="[185,1068,153,1276]" captionTargetId="figure-106@61.[551,733,151,716]" captionTargetPageId="61" captionText="Fig. 31: Left part of the pronotum of Epinebriola species. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) tuberculata sp. nov.; F: N. (E.) numburica sp. nov.; G: N. (E.) retingensis sp. nov.; H: N. (E.) delicata sp. nov.; I: N. (E.) incarinata sp. nov.; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6285816" httpUri="https://zenodo.org/record/6285816/files/figure.png" pageId="55" pageNumber="56">Fig. 31G</figureCitation>
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). Lateral groove very narrow, also apically, broadened basally. Anterior angles faintly acuminate, slightly protruding, margined only at apex. Posterior angles acute, slightly protruding backwards. Basal margin medially straight, in front of the posterior angles weakly sinuate. Pronotal disc weakly convex. Basal fovea deep, anterior transverse impression weak, posterior transverse impression distinct, median line shallow. Lateral groove and anterior transverse impression impunctate; basal fovea and posterior transverse impression sparsely punctate. Apical margination of the pronotum restricted to lateral one-fourth, merging with the apex of the anterior angle. Base of the pronotum not margined. Basolateral seta present. One midlateral seta in the apical half of the pronotum, inserted within the narrow lateral groove; outline of the lateral margin at the insertion site of the seta weakly disturbed. Pronotum impunctate on disc, microreticulation isodiametric. Proepisternum craniad coarsely and shallowly punctate. Prosternal process laterally distinctly margined, apex margined or bulging.
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</paragraph>
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<paragraph id="8B82368FFFECFFF7FF41FB58FC95F9BE" blockId="55.[184,1069,156,1636]" pageId="55" pageNumber="56">Elytral silhouette slender, long-ovoid, with the lateral margin almost straight in the middle, widest at apical third behind the middle. Ratio length/width of the elytra = 1.69. Distinctly wider than the pronotum, ratio elytral width/pronotal width = 1.71. Elytral apex acutely rounded, apical carina faint or even absent. Basal margination straight, at right angles with the elytral suture, obliquely merged with the lateral margination. Humeral carina present. Striae distinct, deep, reaching the apex, striae coarsely punctate; lateral striae obliterate towards the apex. Intervals flat or slightly convex. Interval 3 with 2–4 setae adjoining stria 3, intervals 5 and 7 asetose. Scutellar seta absent. Microreticulation isodiametric. Mesepisterna sparsely punctate.</paragraph>
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<paragraph id="8B82368FFFECFFF8FF41F9CFFBB5FA69" blockId="55.[184,1069,156,1636]" lastBlockId="56.[221,1107,1388,1636]" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">Sternum 2 laterally coarsely punctate. Sternum 3 medially unisetose or bisetose. Sterna 4–6 posteriorly each with 3–6 paramedial setae; sterna laterally with shallow impressions. Anal sternum with two paramedial setae in the male, with 2–4 setae in the female. Metepisterna twice as long as wide, sparsely punctate. Metacoxa basally with 2–5 setae, apically unisetose.</paragraph>
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<caption id="DF426607FFE3FFF8FF43FF49FEC9FEB5" ID-DOI="http://doi.org/10.5281/zenodo.6285804" ID-Zenodo-Dep="6285804" httpUri="https://zenodo.org/record/6285804/files/figure.png" pageId="56" pageNumber="57" startId="56.[223,251,160,178]" targetBox="[463,1106,153,1322]" targetPageId="56">
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<paragraph id="8B82368FFFE3FFF8FF43FF49FEC9FEB5" blockId="56.[221,390,160,274]" pageId="56" pageNumber="57">
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Fig. 28:
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<taxonomicName id="4C3D4D0CFFE3FFF8FEBFFF49FEFEFF6D" authority="(Epinebriola)" authorityName="LATREILLE" authorityYear="1802" baseAuthorityName="Epinebriola" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
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Nebria
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<emphasis id="B949EA9DFFE3FFF8FF41FF51FEFEFF6D" bold="true" box="[221,334,184,202]" pageId="56" pageNumber="57">
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||
(
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<emphasis id="B949EA9DFFE3FFF8FF78FF51FEF7FF6D" bold="true" box="[228,327,184,202]" italics="true" pageId="56" pageNumber="57">Epinebriola</emphasis>
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)
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</emphasis>
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</taxonomicName>
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<emphasis id="B949EA9DFFE3FFF8FF41FF39FEC9FEB5" bold="true" pageId="56" pageNumber="57">
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<taxonomicName id="4C3D4D0CFFE3FFF8FF41FF39FE8EFF45" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[221,318,208,226]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="retingensis" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFE3FFF8FF41FF39FE8EFF45" bold="true" box="[221,318,208,226]" italics="true" pageId="56" pageNumber="57">retingensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE3FFF8FEDFFF39FE36FF45" box="[323,390,208,226]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<typeStatus id="5486882DFFE3FFF8FF41FF01FE9DFF5D" box="[221,301,232,250]" pageId="56" pageNumber="57" type="holotype">Holotype</typeStatus>
|
||
. Scale bar =
|
||
<quantity id="4CC59B6AFFE3FFF8FED8FEE9FEC5FEB5" box="[324,373,256,274]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="56" pageNumber="57" unit="mm" value="5.0">5 mm</quantity>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B82368FFFE3FFF9FE9FFA35FD27FBA9" blockId="56.[221,1107,1388,1636]" lastBlockId="57.[184,1069,977,1636]" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">Legs long and slender. All tarsomeres dorsally glabrous. Protarsus of male stoutly built with tarsomeres 1–3 distinctly broader than in the female, and ventrally with pads of adhesive setae. Ventroapical tooth of the metatarsomere 4 markedly projected. Meso- and metatarsomeres 2–4 ventroapically with long setae. All tarsomeres 5 ventrally with two rows of 3–4 short setae. Metatarsomere 5 as long as the metatarsomeres 3+4.</paragraph>
|
||
<caption id="DF426607FFE2FFF9FF24FCA8FD84FC2F" ID-DOI="http://doi.org/10.5281/zenodo.6285808" ID-Zenodo-Dep="6285808" httpUri="https://zenodo.org/record/6285808/files/figure.png" pageId="57" pageNumber="58" startId="57.[184,213,833,851]" targetBox="[184,1068,153,808]" targetPageId="57">
|
||
<paragraph id="8B82368FFFE2FFF9FF24FCA8FD84FC2F" blockId="57.[184,1068,833,904]" pageId="57" pageNumber="58">
|
||
Fig. 29: Habitat of
|
||
<taxonomicName id="4C3D4D0CFFE2FFF9FEFEFCA8FD30FCF4" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[354,640,833,851]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="retingensis" status="sp. nov." subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE2FFF9FEFEFCA8FE2CFCF4" bold="true" box="[354,412,833,851]" italics="true" pageId="57" pageNumber="58">Nebria</emphasis>
|
||
(
|
||
<emphasis id="B949EA9DFFE2FFF9FE37FCA8FDBFFCF4" bold="true" box="[427,527,833,851]" italics="true" pageId="57" pageNumber="58">Epinebriola</emphasis>
|
||
)
|
||
<emphasis id="B949EA9DFFE2FFF9FD81FCA8FD30FCF4" bold="true" box="[541,640,833,851]" italics="true" pageId="57" pageNumber="58">retingensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE2FFF9FD1BFCA8FD7AFCF4" box="[647,714,833,851]" pageId="57" pageNumber="58" rank="species">sp.nov.</taxonomicNameLabel>
|
||
: The specimens were collected on melt water rivulets in stone fields in the lower alpine zone (4800 m a.s.l.) of the Gangdise Shan near the Reting Monastery. Photo: Joachim Schmidt.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B82368FFFE2FFF9FF41FBF5FC02FB6E" blockId="57.[184,1069,977,1636]" pageId="57" pageNumber="58">
|
||
Male genitalia: Edeagus (
|
||
<figureCitation id="13062A0AFFE2FFF9FE6EFBF5FDE0FB93" box="[498,592,1052,1076]" captionStart="Fig" captionStartId="38.[221,250,1317,1335]" captionTargetBox="[221,1106,153,1296]" captionTargetId="figure-70@38.[221,704,627,867]" captionTargetPageId="38" captionText="Fig. 20: Male edeagi of new Nebria species of the subgenus Epinebriola K. DANIEL, 1904. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) numburica sp. nov., with parameres; F: N. (E.) retingensis sp. nov.; G: N. (E.) incarinata sp. nov.; H: N. (E.) delicata sp. nov.; I: N. (E.) incarinata sp. nov., apex in dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6285764" httpUri="https://zenodo.org/record/6285764/files/figure.png" pageId="57" pageNumber="58">Fig. 20F</figureCitation>
|
||
): Base of the median lobe small, with small basolateral lobes; the base decreasing abruptly to the thin mid-shaft. Mid-shaft strongly curved at base on inner side as well as on outer side; moderately curved to the apex. Apex straight, faintly deflected to the left, in front of the short tip on the ventral side faintly convex. Mid-shaft of the endophallus asetose.
|
||
</paragraph>
|
||
<paragraph id="8B82368FFFE2FFF9FF41FB3FFC68FA0E" blockId="57.[184,1069,977,1636]" pageId="57" pageNumber="58">
|
||
Female genitalia: Gonocoxa (
|
||
<figureCitation id="13062A0AFFE2FFF9FDBDFB3FFDCAFB49" box="[545,634,1238,1262]" captionStart="Fig" captionStartId="20.[221,250,1430,1448]" captionTargetBox="[228,1106,153,1365]" captionTargetId="figure-16@20.[574,757,751,1260]" captionTargetPageId="20" captionText="Fig. 10: Right gonocoxae of Epinebriola species, in ventral view. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) tuberculata sp. nov.; F: N. (E.) numburica sp. nov.; G: N. (E.) numburica sp. nov. in lateral view with the distale gonocoxite 2 dorsally grooved (arrow); H: N. (E.) christinae HUBER & SCHMIDT, 2007; I: N. (E.) pseudorestias sp. nov.; J: N. (E.) retingensis sp. nov.; Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6285726" httpUri="https://zenodo.org/record/6285726/files/figure.png" pageId="57" pageNumber="58">Fig. 10J</figureCitation>
|
||
,
|
||
<figureCitation id="13062A0AFFE2FFF9FD18FB3FFD07FB49" box="[644,695,1238,1262]" captionStart="Fig" captionStartId="64.[221,251,1018,1036]" captionTargetBox="[221,992,153,958]" captionTargetId="figure-110@64.[423,633,150,988]" captionTargetPageId="64" captionText="Fig. 34: Right gonocoxa, in ventral view. A: Nebria (Epinebriola) retingensis sp. nov. (= Fig. 10J); B: N. (Eonebria) kurentzovi LAFER, 1989; C: N. (Eonebria) gemina LEDOUx, ROUx & WRASE, 1996; D: N. (Eonebria) amabilis LEDOUx, ROUx & SAWADA, 1991. Scale bar = 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.6285830" httpUri="https://zenodo.org/record/6285830/files/figure.png" pageId="57" pageNumber="58">34A</figureCitation>
|
||
): Gonocoxite 2 very long, as long or longer than gonocoxite 1, slender, curved dagger-like, broad at base, distinctly narrowing near base, parallel sided in median part. Apex acuminate, slightly arcuate dorsally, apex dorsally not grooved. Ventral preapical insertion furrow small, short-oval, with two nematiform setae. Gonocoxites ventrally unjointed and continuously sclerotized, dorsally separated by a membranous area.
|
||
</paragraph>
|
||
<paragraph id="8B82368FFFE2FFF9FF41FA5FFECBF999" blockId="57.[184,1069,977,1636]" pageId="57" pageNumber="58">Body ratios: hea.w/fro.w = 1.50±0.012 (1.48–1.51); prm.w/hea.w = 1.15 (1.13–1.17); prm.w/prp.w = 1.52 (1.51–1.53); pra.w/prp.w = 1.12 (1.11–1.13); ely.w/prm.w = 1.71 (1.66–1.74); prm.w/prm.l = 1.23 (1.22–1.25(; ely.l/ely.w = 1.69 (1.66–1.73).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFE2FFF9FF41F9A5FCDAF9C3" box="[221,874,1612,1636]" pageId="57" pageNumber="58" type="etymology">
|
||
<paragraph id="8B82368FFFE2FFF9FF41F9A5FCDAF9C3" blockId="57.[184,1069,977,1636]" box="[221,874,1612,1636]" pageId="57" pageNumber="58">
|
||
Etymology: The specific epithet refers to the
|
||
<typeStatus id="5486882DFFE2FFF9FD4BF9A5FCBAF9C3" box="[727,778,1612,1636]" pageId="57" pageNumber="58">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFE1FFFAFE9FFF75FB8BFE79" pageId="58" pageNumber="59" type="diagnosis">
|
||
<paragraph id="8B82368FFFE1FFFAFE9FFF75FB8BFE79" blockId="58.[221,1108,156,740]" pageId="58" pageNumber="59">
|
||
Diagnosis:
|
||
<taxonomicName id="4C3D4D0CFFE1FFFAFE1FFF75FD92FF13" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[387,546,156,180]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="retingensis" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFE1FFFAFE1FFF75FD92FF13" box="[387,546,156,180]" italics="true" pageId="58" pageNumber="59">N. retingensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE1FFFAFDB6FF75FD34FF13" box="[554,644,156,180]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is closely related to
|
||
<taxonomicName id="4C3D4D0CFFE1FFFAFCEAFF75FC40FF13" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[886,1008,156,180]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="delicata" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFE1FFFAFCEAFF75FC40FF13" box="[886,1008,156,180]" italics="true" pageId="58" pageNumber="59">N. delicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE1FFFAFC65FF75FBE4FF13" box="[1017,1108,156,180]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
due to the resemblance in the shape of the median lobes (
|
||
<figureCitation id="13062A0AFFE1FFFAFCF9FF28FC55FF7E" box="[869,997,193,217]" captionStart="Fig" captionStartId="29.[184,213,1019,1037]" captionTargetBox="[184,1068,153,981]" captionTargetId="figure-89@29.[184,1068,153,981]" captionTargetPageId="29" captionText="Fig. 16: Locations with occurrences of Nebria (Barbonebriola sp. nov.) species. 1: Lac de Gangabal, Kashmir Himalaya of India (N. barbata ANDREWES, 1929). 2: Lac de Gadsar, Kashmir Himalaya of India (N. barbata ANDREWES, 1929). 3: Sonamarg, Kashmir Himalaya of India (N. barbata ANDREWES, 1929). 4: Babussar Pass, Kashmir Himalaya of Pakistan (N. barbata ANDREWES, 1929). 5: Rama Lake, Kashmir Himalaya of Pakistan (N. barbata ANDREWES, 1929). 6: Rohtang Pass, Himachal Pradesh, India (N. ganeshi LEDOUx, 1984). 7: Badrinath, Uttarakhand, India (N. restricta LEDOUx & ROUx, 2005). 8: South slope of Api Himal, Ghusa, Nepal (N. tenuisulcata sp. nov.). 9: East slope of Saipal Himal, Sankha La, Nepal (N. rubostipes sp. nov.). 10: South slope of Kanjiroba Himal, Kagmara La, Nepal (N. kagmara sp. nov.)." figureDoi="http://doi.org/10.5281/zenodo.6285752" httpUri="https://zenodo.org/record/6285752/files/figure.png" pageId="58" pageNumber="59">Figs 16F, H</figureCitation>
|
||
).
|
||
<taxonomicName id="4C3D4D0CFFE1FFFAFC6BFF28FE96FF59" authorityName="C.Huber & J.Schmidt" authorityYear="2017" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="retingensis" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFE1FFFAFC6BFF28FE96FF59" italics="true" pageId="58" pageNumber="59">N. retingensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE1FFFAFEB2FF0FFE37FF59" box="[302,391,230,254]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from the similar
|
||
<taxonomicName id="4C3D4D0CFFE1FFFAFD06FF0FFCA3FF59" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[666,787,230,254]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="delicata" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFE1FFFAFD06FF0FFCA3FF59" box="[666,787,230,254]" italics="true" pageId="58" pageNumber="59">N. delicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE1FFFAFC87FF0FFCC4FF59" box="[795,884,230,254]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the labrum with six setae (eight setae in
|
||
<taxonomicName id="4C3D4D0CFFE1FFFAFE63FEE5FDCBFE83" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[511,635,268,292]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="delicata" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFE1FFFAFE63FEE5FDCBFE83" box="[511,635,268,292]" italics="true" pageId="58" pageNumber="59">N. delicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE1FFFAFD18FEE5FD55FE83" box="[644,741,268,292]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
), by the finely punctate labrum (impunctate in
|
||
<taxonomicName id="4C3D4D0CFFE1FFFAFE0CFED8FDBCFEEE" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[400,524,305,329]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="delicata" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFE1FFFAFE0CFED8FDBCFEEE" box="[400,524,305,329]" italics="true" pageId="58" pageNumber="59">N. delicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE1FFFAFD89FED8FDC7FEEE" box="[533,631,305,329]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
), by large and protruding eyes with long temples (small eyes, not protruding, temples shorter in
|
||
<taxonomicName id="4C3D4D0CFFE1FFFAFCF8FEBFFC6FFEC9" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[868,991,342,366]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="delicata" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFE1FFFAFCF8FEBFFC6FFEC9" box="[868,991,342,366]" italics="true" pageId="58" pageNumber="59">N. delicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE1FFFAFC7BFEBFFBF8FEC9" box="[999,1096,342,366]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
), by the lateral margin of the pronotum which is straight towards the posterior angles (with a faint concave sinuation in front of the posterior angles in
|
||
<taxonomicName id="4C3D4D0CFFE1FFFAFB9DFE48FEBCFE79" authorityName="C.Huber & J.Schmidt" authorityYear="2017" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="delicata" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFE1FFFAFB9DFE48FEBCFE79" italics="true" pageId="58" pageNumber="59">N. delicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE1FFFAFE8EFE2FFEDFFE79" box="[274,367,454,478]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
), and by hanging shoulders (accentuated in
|
||
<taxonomicName id="4C3D4D0CFFE1FFFAFCCBFE2FFC7EFE79" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[855,974,454,478]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="delicata" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFE1FFFAFCCBFE2FFC7EFE79" box="[855,974,454,478]" italics="true" pageId="58" pageNumber="59">N. delicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE1FFFAFC4FFE2FFB80FE79" box="[979,1072,454,478]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFE1FFFAFE9FFE05FE04FD3E" pageId="58" pageNumber="59" type="distribution">
|
||
<paragraph id="8B82368FFFE1FFFAFE9FFE05FE04FD3E" blockId="58.[221,1108,156,740]" pageId="58" pageNumber="59">
|
||
Distribution (
|
||
<figureCitation id="13062A0AFFE1FFFAFE04FE05FE56FDA3" box="[408,486,492,516]" captionStart="Fig" captionStartId="76.[221,250,160,178]" captionTargetId="figure-8@75.[184,1068,152,1645]" captionTargetPageId="75" captionText="Fig. 41: Locations with occurrences of Nebria (Epinebriola) species from the Central and Eastern Himalaya and from South Tibet." figureDoi="http://doi.org/10.5281/zenodo.6285892" httpUri="https://zenodo.org/record/6285892/files/figure.png" pageId="58" pageNumber="59">Fig. 41</figureCitation>
|
||
): Known only from the
|
||
<typeStatus id="5486882DFFE1FFFAFD6AFE05FC99FDA3" box="[758,809,492,516]" pageId="58" pageNumber="59">type</typeStatus>
|
||
locality in the Transhimalaya (Gangdise Shan) near the famous Reting Monastery immediately south of the river Reting Zhangbo.
|
||
<taxonomicName id="4C3D4D0CFFE1FFFAFD99FDDFFD13FDE9" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[517,675,566,590]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="retingensis" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFE1FFFAFD99FDDFFD13FDE9" box="[517,675,566,590]" italics="true" pageId="58" pageNumber="59">N. retingensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE1FFFAFD36FDDFFCB3FDE9" box="[682,771,566,590]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is one of the two
|
||
<taxonomicName id="4C3D4D0CFFE1FFFAFC4CFDDFFBE2FDE9" authorityName="K. DANIEL" authorityYear="1904" box="[976,1106,566,590]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="79" phylum="Arthropoda" rank="subGenus" subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE1FFFAFC4CFDDFFBE2FDE9" box="[976,1106,566,590]" italics="true" pageId="58" pageNumber="59">Epinebriola</emphasis>
|
||
</taxonomicName>
|
||
species (in addition to
|
||
<taxonomicName id="4C3D4D0CFFE1FFFAFE42FDB5FDE6FDD3" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[478,598,604,628]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="delicata" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFE1FFFAFE42FDB5FDE6FDD3" box="[478,598,604,628]" italics="true" pageId="58" pageNumber="59">N. delicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE1FFFAFDC0FDB5FD07FDD3" box="[604,695,604,628]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) known to inhabit the central part of the Transhimalaya.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFE1FFFAFE9FFD4FFE5BFD43" pageId="58" pageNumber="59" type="biology_ecology">
|
||
<paragraph id="8B82368FFFE1FFFAFE9FFD4FFE5BFD43" blockId="58.[221,1108,156,740]" pageId="58" pageNumber="59">
|
||
Habitat (
|
||
<figureCitation id="13062A0AFFE1FFFAFEFCFD4FFE1BFD19" box="[352,427,678,702]" captionStart="Fig" captionStartId="57.[184,213,833,851]" captionTargetBox="[184,1068,153,808]" captionTargetId="figure-253@57.[184,1068,153,811]" captionTargetPageId="57" captionText="Fig. 29: Habitat of Nebria (Epinebriola) retingensis sp.nov.: The specimens were collected on melt water rivulets in stone fields in the lower alpine zone (4800 m a.s.l.) of the Gangdise Shan near the Reting Monastery. Photo: Joachim Schmidt." figureDoi="http://doi.org/10.5281/zenodo.6285808" httpUri="https://zenodo.org/record/6285808/files/figure.png" pageId="58" pageNumber="59">Fig. 29</figureCitation>
|
||
): All specimens were collected alongside a melt water stream in the lower alpine zone.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |