treatments-xml/data/03/F3/87/03F387B2FFBB5370CD2451AEFD71A641.xml
2024-06-21 12:22:17 +02:00

430 lines
52 KiB
XML
Raw Blame History

This file contains invisible Unicode characters

This file contains invisible Unicode characters that are indistinguishable to humans but may be processed differently by a computer. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="E717261F4C59000CF49CEA344949C55B" ID-CLB-Dataset="5458" ID-DOI="10.11646/zootaxa.353.1.1" ID-GBIF-Dataset="740af13a-ed5c-40d0-b0c1-8d3fd2b92f56" ID-ISSN="1175­5334" ID-Zenodo-Dep="5019783" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1624467278120" checkinUser="felipe" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="03F387B2FFBB5370CD2451AEFD71A641" docLanguage="en" docName="zt00353.pdf" docOrigin="Zootaxa 353 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Atheta (Tetropla) nigritula" docType="treatment" docVersion="10" lastPageNumber="61" masterDocId="FFCAFFCAFF80534CCC2C5352FFD8A23F" masterDocTitle="Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes" masterLastPageNumber="134" masterPageNumber="1" pageNumber="60" updateTime="1698972890617" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="4886C286DC85BDC7367762710FD7FA89" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B853174587729E3FA7F9A6D951974C22">
<mods:title id="E8DCBC353A350F3704F9E54886A456CD">Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes</mods:title>
</mods:titleInfo>
<mods:name id="91BF74E7AECC56E94B1F18FEB419841C" type="personal">
<mods:role id="8CAD44B7649AB27035917B1BC7390B20">
<mods:roleTerm id="13593F004DDF0CD75CBAC2F04144DA84">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="374F1D7C40F67D96958E9E5B16887613">Gusarov, Vladimir I.</mods:namePart>
</mods:name>
<mods:typeOfResource id="3BBA25087D7C05906F08EC76B8535DE6">text</mods:typeOfResource>
<mods:relatedItem id="B063E755BEB211EE17B16F57BECE69A6" type="host">
<mods:titleInfo id="0C7CFE4565BC24548876DBAD8CF39E98">
<mods:title id="0EF1F3F33F91A05F8D23676896206B5F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="595928B54BBA265F26323AC9A1F7E0B3">
<mods:date id="91C5AE84B23613B9CAE2717E8891726B">2003</mods:date>
<mods:detail id="D6FDBFA4C823B97345AFABB1662C089E" type="pubDate">
<mods:number id="79F4AC799BFE99BF40A1891B32E9ACC3">2003-11-17</mods:number>
</mods:detail>
<mods:detail id="BE82DADD039D85A31A1ECA910035EB09" type="volume">
<mods:number id="DC9EB76C8F32C3F7461940A5DAF01450">353</mods:number>
</mods:detail>
<mods:detail id="9AB4478A15FF836852DD20397682E97E" type="issue">
<mods:number id="A0DC612ECA87B93AA3D0D02A5251DD1C">1</mods:number>
</mods:detail>
<mods:extent id="13F26BAAE55E36EF17D9C576005F046D" unit="page">
<mods:start id="1EEAF2F6FEF5310E1C588D7D9107057B">1</mods:start>
<mods:end id="CFD8453E21EEA9B620DDB83FED0BCBBA">134</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="1583C8B778E331BDFAFAEEBC146F9145">
<mods:url id="B98BB6033BB43B6A8CE623D0CDECD6D8">https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1</mods:url>
</mods:location>
<mods:classification id="EA68699429A7CA36ECDD78394FD93909">journal article</mods:classification>
<mods:identifier id="47C98CE76F13A3015E571CA1D0D9AD5F" type="CLB-Dataset">5458</mods:identifier>
<mods:identifier id="6B2AEE814E5340AFA99936C9BBC0964A" type="DOI">10.11646/zootaxa.353.1.1</mods:identifier>
<mods:identifier id="959EFE0E4527B926C12E9148776D4928" type="GBIF-Dataset">740af13a-ed5c-40d0-b0c1-8d3fd2b92f56</mods:identifier>
<mods:identifier id="831D87FFB0E36F0D3049A76F508A15C0" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="62DB7A8B24F6E4DC802A713C18799AF5" type="Zenodo-Dep">5019783</mods:identifier>
<mods:identifier id="9B803401C0B546099E012D7B79DF3898" type="ZooBank">104E9C0B-B207-41E6-816C-E91FC1C7DBA0</mods:identifier>
</mods:mods>
<treatment id="03F387B2FFBB5370CD2451AEFD71A641" ID-DOI="http://doi.org/10.5281/zenodo.5103054" ID-GBIF-Taxon="183542856" ID-Zenodo-Dep="5103054" LSID="urn:lsid:plazi:treatment:03F387B2FFBB5370CD2451AEFD71A641" httpUri="http://treatment.plazi.org/id/03F387B2FFBB5370CD2451AEFD71A641" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
<subSubSection id="C340652FFFBB5377CD2451AEFC93A129" box="[264,843,764,790]" pageId="59" pageNumber="60" type="nomenclature">
<paragraph id="8BE536A4FFBB5377CD2451AEFC93A129" blockId="59.[264,1112,764,825]" box="[264,843,764,790]" pageId="59" pageNumber="60">
<heading id="D0AD81C8FFBB5377CD2451AEFC93A129" bold="true" box="[264,843,764,790]" fontSize="11" level="1" pageId="59" pageNumber="60" reason="1">
<taxonomicName id="4C5A4D27FFBB5377CD2451AEFC93A129" ID-CoL="JDYD" authority="(Gravenhorst, 1802)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[264,843,764,790]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="nigritula" subGenus="Tetropla">
<emphasis id="B92EEAB6FFBB5377CD2451AEFC93A129" bold="true" box="[264,843,764,790]" pageId="59" pageNumber="60">
<emphasis id="B92EEAB6FFBB5377CD2451AEFE8FA129" bold="true" box="[264,343,764,790]" italics="true" pageId="59" pageNumber="60">Atheta</emphasis>
(
<emphasis id="B92EEAB6FFBB5377CD4551AEFE15A129" bold="true" box="[361,461,764,790]" italics="true" pageId="59" pageNumber="60">Tetropla</emphasis>
)
<emphasis id="B92EEAB6FFBB5377CDF151AEFD9FA129" bold="true" box="[477,583,764,790]" italics="true" pageId="59" pageNumber="60">nigritula</emphasis>
(
<bibRefCitation id="EFCB4B55FFBB5377CE7B51AEFC9BA129" author="Gravenhorst, J. L. C." box="[599,835,764,790]" pageId="59" pageNumber="60" refId="ref77424" refString="Gravenhorst, J. L. C. (1802) Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera familias et species distribuit. C. Reichard, Brunswick, xxvi + 207 pp." type="book" year="1802">Gravenhorst, 1802</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C340652FFFBB5377CD24504DFB80A106" box="[264,1112,799,825]" pageId="59" pageNumber="60" type="description">
<paragraph id="8BE536A4FFBB5377CD24504DFB80A106" blockId="59.[264,1112,764,825]" box="[264,1112,799,825]" pageId="59" pageNumber="60">
(
<figureCitation id="13612A21FFBB5377CD3D504DFEAAA106" box="[273,370,799,825]" captionStart="FIGURES 116119" captionStartId="81.[264,383,1731,1755]" captionTargetBox="[346,1237,974,1699]" captionTargetId="figure-253@81.[341,1245,964,1705]" captionTargetPageId="81" captionText="FIGURES 116119. Abdominal segment 8 of Boreophilia angusticornis (Bernhauer) (male, lectotype of Atheta angusticornis (116117); and female, lectotype of Metaxya plutonica Casey (118119)). 116 — male tergum 8; 117 — male sternum 8; 118 — female tergum 8; 119 — female sternum 8. Scale bar 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5019864" httpUri="https://zenodo.org/record/5019864/files/figure.png" pageId="59" pageNumber="60">
Fig.
<quantity id="4CA29B41FFBB5377CD6B504DFEAAA106" box="[327,370,799,825]" metricMagnitude="0" metricUnit="m" metricValue="2.9718" pageId="59" pageNumber="60" unit="in" value="117.0">117</quantity>
</figureCitation>
in
<bibRefCitation id="EFCB4B55FFBB5377CDB4504DFD58A106" author="Strand, A. &amp; Vik, A." box="[408,640,799,825]" pageId="59" pageNumber="60" pagination="327 - 335" refId="ref81239" refString="Strand, A. &amp; Vik, A. (1964) Die Genitalorgane der nordischen Arten der Gattung Atheta Thoms. (Col., Staphylinidae). Norsk Entomologisk Tidsskrift, 12 (5 - 8), 327 - 335, Taf. I - XXI." type="journal article" year="1964">Strand &amp; Vik, 1964</bibRefCitation>
;
<figureCitation id="13612A21FFBB5377CEA0504DFD0AA106" box="[652,722,799,825]" captionStart="FIGURES 59" captionStartId="13.[264,382,1337,1361]" captionTargetBox="[298,1291,486,1318]" captionTargetId="figure-264@13.[288,1298,477,1322]" captionTargetPageId="13" captionText="FIGURES 59. Genitalia of Tinotus acerbus (Casey) (lectotype of Acrimea acerba (58) and lectotype of Acri. resecta (9)). 5 — median lobe, parameral view; 6 — median lobe with details of retracted internal sac, abparameral view; 7 — median lobe, lateral view; 8 — median lobe with details of retracted internal sac, lateral view; 9 — spermatheca. Scale bar 0.1 mm (9), 0.2 mm (5 8)." figureDoi="http://doi.org/10.5281/zenodo.5019795" httpUri="https://zenodo.org/record/5019795/files/figure.png" pageId="59" pageNumber="60">Fig. 7</figureCitation>
, p.
<quantity id="4CA29B41FFBB5377CED0504DFC9EA106" box="[764,838,799,825]" metricMagnitude="0" metricUnit="m" metricValue="3.9116" pageId="59" pageNumber="60" unit="in" value="154.0">
<figureCitation id="13612A21FFBB5377CED0504DFCF0A106" box="[764,808,799,825]" captionStart="FIGURES 147154" captionStartId="110.[264,382,1237,1261]" captionTargetBox="[283,1306,318,1200]" captionTargetId="figure-138@110.[273,1314,312,1212]" captionTargetPageId="110" captionText="FIGURES 147154. Genitalia of Strigota ambigua (Erichson) (male, paralectotype of St. gnava Casey (149154); and female, lectotype of St. oppidana Casey (147148)). 147148 — spermatheca (of the same specimen; different views); 149 — median lobe, parameral view; 150 — apex of median lobe, parameral view; 151 — median lobe, lateral view; 152 — apex of median lobe, lateral view; 153154 — apex of median lobe with retracted internal sac, lateral view. Scale bar 0.1 mm (150, 152), 0.2 mm (147149, 151, 153154)." figureDoi="http://doi.org/10.5281/zenodo.5019891" httpUri="https://zenodo.org/record/5019891/files/figure.png" pageId="59" pageNumber="60">154</figureCitation>
in
</quantity>
<bibRefCitation id="EFCB4B55FFBB5377CF61504DFB88A106" author="Benick, G. &amp; Lohse, G. A." box="[845,1104,799,825]" pageId="59" pageNumber="60" pagination="72 - 220" refId="ref75834" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse 1974</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C340652FFFBB5377CD24503DFD92A6B7" pageId="59" pageNumber="60" type="reference_group">
<paragraph id="8BE536A4FFBB5377CD24503DFD07A1BA" blockId="59.[264,1323,878,1094]" box="[264,735,878,902]" pageId="59" pageNumber="60">
<treatmentCitationGroup id="AB4A118AFFBB5377CD24503DFD07A1BA" box="[264,735,878,902]" pageId="59" pageNumber="60">
<treatmentCitation id="0AFB10B5FFBB5377CD24503DFD02A1BA" author="Gravenhorst, J. L. C." box="[264,730,878,902]" page="85" pageId="59" pageNumber="60" year="1802">
<taxonomicName id="4C5A4D27FFBB5377CD24503DFD02A1BA" authority="Gravenhorst, 1802: 85" authorityName="Gravenhorst" authorityPageNumber="85" authorityYear="1802" box="[264,730,878,902]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBB5377CD24503DFE03A1B9" box="[264,475,879,902]" italics="true" pageId="59" pageNumber="60">Aleochara nigritula</emphasis>
<bibRefCitation id="EFCB4B55FFBB5377CDCE503CFD02A1BA" author="Gravenhorst, J. L. C." box="[482,730,878,901]" pageId="59" pageNumber="60" refId="ref77424" refString="Gravenhorst, J. L. C. (1802) Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera familias et species distribuit. C. Reichard, Brunswick, xxvi + 207 pp." type="book" year="1802">Gravenhorst, 1802: 85</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE536A4FFBB5377CD2450DDFEA5A1FA" blockId="59.[264,1323,878,1094]" pageId="59" pageNumber="60">
<taxonomicName id="4C5A4D27FFBB5377CD2450DDFDAEA19A" ID-CoL="8Y4LW" authority="Erichson, 1839" authorityName="Erichson" authorityYear="1839" box="[264,630,910,934]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="socialis">
<emphasis id="B92EEAB6FFBB5377CD2450DDFE12A199" box="[264,458,911,934]" italics="true" pageId="59" pageNumber="60">Homalota socialis</emphasis>
<bibRefCitation id="EFCB4B55FFBB5377CDFE50DCFDAEA19A" author="Erichson, W. F." box="[466,630,910,933]" pageId="59" pageNumber="60" pagination="1 - 400" refId="ref76845" refString="Erichson, W. F. (1839) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, pp. i - viii, 1 - 400." type="journal article" year="1839">Erichson, 1839</bibRefCitation>
</taxonomicName>
, var. c: 103 (as valid name for
<taxonomicName id="4C5A4D27FFBB5377CFEA50DDFEA9A1FA" authority="Gravenhorst, 1802" authorityName="Gravenhorst" authorityYear="1802" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBB5377CFEA50DDFB41A199" box="[966,1177,911,934]" italics="true" pageId="59" pageNumber="60">Aleochara nigritula</emphasis>
<bibRefCitation id="EFCB4B55FFBB5377C88C50DCFEA9A1FA" author="Gravenhorst, J. L. C." pageId="59" pageNumber="60" refId="ref77424" refString="Gravenhorst, J. L. C. (1802) Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera familias et species distribuit. C. Reichard, Brunswick, xxvi + 207 pp." type="book" year="1802">Gravenhorst, 1802</bibRefCitation>
</taxonomicName>
).
</paragraph>
<paragraph id="8BE536A4FFBB5377CD24509DFB91A1DA" blockId="59.[264,1323,878,1094]" box="[264,1097,974,998]" pageId="59" pageNumber="60">
<treatmentCitationGroup id="AB4A118AFFBB5377CD24509DFB91A1DA" box="[264,1097,974,998]" pageId="59" pageNumber="60">
<treatmentCitation id="0AFB10B5FFBB5377CD24509DFC5BA1DA" author="Mulsant, M. E. &amp; Rey, C." box="[264,899,974,998]" page="529" pageId="59" pageNumber="60" year="1874">
<taxonomicName id="4C5A4D27FFBB5377CD24509DFC5BA1DA" authority=": Mulsant &amp; Rey, 1874 a: 529" authorityName="Mulsant &amp; Rey" authorityPageNumber="529" authorityYear="1874" box="[264,899,974,998]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="nigritula" subGenus="Tetropla">
<emphasis id="B92EEAB6FFBB5377CD24509DFEAAA1D9" box="[264,370,975,998]" italics="true" pageId="59" pageNumber="60">Homalota</emphasis>
(
<emphasis id="B92EEAB6FFBB5377CDAE509DFE02A1D9" box="[386,474,975,998]" italics="true" pageId="59" pageNumber="60">Tetropla</emphasis>
)
<emphasis id="B92EEAB6FFBB5377CDC6509DFD9FA1D9" box="[490,583,975,998]" italics="true" pageId="59" pageNumber="60">nigritula</emphasis>
:
<bibRefCitation id="EFCB4B55FFBB5377CE79509CFC5BA1DA" author="Mulsant, M. E. &amp; Rey, C." box="[597,899,974,997]" pageId="59" pageNumber="60" pagination="33 - 738" refId="ref79932" refString="Mulsant, M. E. &amp; Rey, C. (1874 a) Tribu des Brevipennes. Famille des Aleochariens. Septieme branche: Myrmedoniaires. Annales de la Societe d'Agriculture, Histoire Naturelle et Arts Utiles de Lyon, serie 4. 6 [1873], 33 - 738." type="journal article" year="1874">Mulsant &amp; Rey, 1874a: 529</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(as valid species).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE536A4FFBB5377CD2450BDFBB4A63A" blockId="59.[264,1323,878,1094]" box="[264,1132,1006,1030]" pageId="59" pageNumber="60">
<treatmentCitationGroup id="AB4A118AFFBB5377CD2450BDFBB4A63A" box="[264,1132,1006,1030]" pageId="59" pageNumber="60">
<treatmentCitation id="0AFB10B5FFBB5377CD2450BDFC7EA63A" author="Bernhauer, M. &amp; Scheerpeltz, O." box="[264,934,1006,1030]" page="646" pageId="59" pageNumber="60" year="1926">
<taxonomicName id="4C5A4D27FFBB5377CD2450BDFC7EA63A" ID-CoL="JDYD" authority=": Bernhauer &amp; Scheerpeltz, 1926: 646" authorityName="Bernhauer &amp; Scheerpeltz" authorityPageNumber="646" authorityYear="1926" box="[264,934,1006,1030]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBB5377CD2450BDFE96A639" box="[264,334,1007,1030]" italics="true" pageId="59" pageNumber="60">Atheta</emphasis>
(
<emphasis id="B92EEAB6FFBB5377CD7150BDFE48A639" box="[349,400,1007,1030]" italics="true" pageId="59" pageNumber="60">s. str</emphasis>
.)
<emphasis id="B92EEAB6FFBB5377CD8A50BDFDDBA639" box="[422,515,1007,1030]" italics="true" pageId="59" pageNumber="60">nigritula</emphasis>
:
<bibRefCitation id="EFCB4B55FFBB5377CE3450BCFC7EA63A" author="Bernhauer, M. &amp; Scheerpeltz, O." box="[536,934,1006,1029]" pageId="59" pageNumber="60" pagination="499 - 988" refId="ref76079" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988." type="book chapter" year="1926">Bernhauer &amp; Scheerpeltz, 1926: 646</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(as valid species).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE536A4FFBB5377CD24575DFC31A61A" blockId="59.[264,1323,878,1094]" box="[264,1001,1038,1062]" pageId="59" pageNumber="60">
<treatmentCitationGroup id="AB4A118AFFBB5377CD24575DFC31A61A" box="[264,1001,1038,1062]" pageId="59" pageNumber="60">
<treatmentCitation id="0AFB10B5FFBB5377CD24575DFCFCA61A" author="Strand, A. &amp; Vik, A." box="[264,804,1038,1062]" page="331" pageId="59" pageNumber="60" year="1964">
<taxonomicName id="4C5A4D27FFBB5377CD24575DFCFCA61A" ID-CoL="JDYD" authority=": Strand &amp; Vik, 1964: 331" authorityName="Strand &amp; Vik" authorityPageNumber="331" authorityYear="1964" box="[264,804,1038,1062]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBB5377CD24575DFE96A619" box="[264,334,1039,1062]" italics="true" pageId="59" pageNumber="60">Atheta</emphasis>
(
<emphasis id="B92EEAB6FFBB5377CD71575DFE4DA619" box="[349,405,1039,1062]" italics="true" pageId="59" pageNumber="60">s. str.</emphasis>
)
<emphasis id="B92EEAB6FFBB5377CD88575DFDD9A619" box="[420,513,1039,1062]" italics="true" pageId="59" pageNumber="60">nigritula</emphasis>
:
<bibRefCitation id="EFCB4B55FFBB5377CE3A575CFCFCA61A" author="Strand, A. &amp; Vik, A." box="[534,804,1038,1061]" pageId="59" pageNumber="60" pagination="327 - 335" refId="ref81239" refString="Strand, A. &amp; Vik, A. (1964) Die Genitalorgane der nordischen Arten der Gattung Atheta Thoms. (Col., Staphylinidae). Norsk Entomologisk Tidsskrift, 12 (5 - 8), 327 - 335, Taf. I - XXI." type="journal article" year="1964">Strand &amp; Vik, 1964: 331</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(as valid species).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE536A4FFBB5377CD24577DFBF3A67A" blockId="59.[264,1323,878,1094]" box="[264,1067,1070,1094]" pageId="59" pageNumber="60">
<treatmentCitationGroup id="AB4A118AFFBB5377CD24577DFBF3A67A" box="[264,1067,1070,1094]" pageId="59" pageNumber="60">
<treatmentCitation id="0AFB10B5FFBB5377CD24577DFCBDA67A" author="Benick, G. &amp; Lohse, G. A." box="[264,869,1070,1094]" page="154" pageId="59" pageNumber="60" year="1974">
<taxonomicName id="4C5A4D27FFBB5377CD24577DFCBDA67A" ID-CoL="JDYD" authority=": Benick &amp; Lohse, 1974: 154" authorityName="Benick &amp; Lohse" authorityPageNumber="154" authorityYear="1974" box="[264,869,1070,1094]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="nigritula" subGenus="Tetropla">
<emphasis id="B92EEAB6FFBB5377CD24577DFE96A679" box="[264,334,1071,1094]" italics="true" pageId="59" pageNumber="60">Atheta</emphasis>
(
<emphasis id="B92EEAB6FFBB5377CD71577DFE6DA679" box="[349,437,1071,1094]" italics="true" pageId="59" pageNumber="60">Tetropla</emphasis>
)
<emphasis id="B92EEAB6FFBB5377CDE9577DFDFAA679" box="[453,546,1071,1094]" italics="true" pageId="59" pageNumber="60">nigritula</emphasis>
:
<bibRefCitation id="EFCB4B55FFBB5377CE1B577CFCBDA67A" author="Benick, G. &amp; Lohse, G. A." box="[567,869,1070,1093]" pageId="59" pageNumber="60" pagination="72 - 220" refId="ref75834" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse, 1974: 154</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(as valid species).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE536A4FFBB5377CD245723FD92A6B7" blockId="59.[264,586,1137,1160]" box="[264,586,1137,1160]" pageId="59" pageNumber="60">(Other references are omitted)</paragraph>
</subSubSection>
<subSubSection id="C340652FFFBB5377CD2457EEFD4FA7A1" pageId="59" pageNumber="60" type="materials_examined">
<paragraph id="8BE536A4FFBB5377CD2457EEFB9AA6C1" blockId="59.[264,1323,1211,1878]" pageId="59" pageNumber="60">
<emphasis id="B92EEAB6FFBB5377CD2457EEFE67A6E9" bold="true" box="[264,447,1212,1238]" pageId="59" pageNumber="60">Type material.</emphasis>
<materialsCitation id="3B323CF9FFBB5377CDE057EEFB9AA6C1" ID-GBIF-Occurrence="3321919322" collectionCode="ZMHB" pageId="59" pageNumber="60" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<typeStatus id="54E18806FFBB5377CDE057EEFD99A6E9" box="[460,577,1212,1238]" pageId="59" pageNumber="60" type="lectotype">Lectotype</typeStatus>
of
<taxonomicName id="4C5A4D27FFBB5377CE4457EEFC97A6EA" authorityName="Gravenhorst" authorityYear="1802" box="[616,847,1212,1237]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBB5377CE4457EEFC97A6EA" box="[616,847,1212,1237]" italics="true" pageId="59" pageNumber="60">Aleochara nigritula</emphasis>
</taxonomicName>
(here designated):
<specimenCount id="9D5CFD2DFFBB5377C81B57E9FB93A6F0" box="[1079,1099,1211,1231]" pageId="59" pageNumber="60" type="female"></specimenCount>
(with the head and pronotum missing), “var. c.
<taxonomicName id="4C5A4D27FFBB5377CE7C57B6FCDFA6C1" authority="Gr." authorityName="Gravenhorst" authorityYear="1802" box="[592,775,1252,1278]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBB5377CE7C57B6FD00A6C2" box="[592,728,1252,1277]" italics="true" pageId="59" pageNumber="60">A. nigritula</emphasis>
Gr.
</taxonomicName>
”, “
<taxonomicName id="4C5A4D27FFBB5377CF0057B6FC18A6C1" authority="Gr." authorityName="Gravenhorst" authorityYear="1802" box="[812,960,1252,1278]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBB5377CF0057B6FC4BA6C2" box="[812,915,1252,1277]" italics="true" pageId="59" pageNumber="60">nigritula</emphasis>
Gr.
</taxonomicName>
” (
<collectionCode id="ED4BAE61FFBB5377CFF757B6FBEEA6C1" box="[987,1078,1252,1278]" pageId="59" pageNumber="60">ZMHB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BE536A4FFBB5377CD14565EFD4FA7A1" blockId="59.[264,1323,1211,1878]" pageId="59" pageNumber="60">
<emphasis id="B92EEAB6FFBB5377CD14565EFDE3A719" bold="true" box="[312,571,1292,1318]" pageId="59" pageNumber="60">Additional material.</emphasis>
<materialsCitation id="3B323CF9FFBB5377CE7A565EFEBCA749" ID-GBIF-Occurrence="3321919694" collectingDate="1990-05-27" collectionCode="SPSU" collectorName="V. I. Gusarov" country="Ukraine" elevation="400" location="Krasnoles'ye" municipality="Simferopol' distr." pageId="59" pageNumber="60" specimenCount="10" stateProvince="Crimea">
<emphasis id="B92EEAB6FFBB5377CE7A565EFCB4A719" bold="true" box="[598,876,1292,1318]" pageId="59" pageNumber="60">
<collectingCountry id="F34D7634FFBB5377CE7A565EFD3DA719" box="[598,741,1292,1318]" name="Ukraine" pageId="59" pageNumber="60">UKRAINE</collectingCountry>
:
<collectingRegion id="499EF846FFBB5377CF28565EFCBFA719" box="[772,871,1292,1318]" country="Ukraine" name="Crimea" pageId="59" pageNumber="60">Crimea</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6B81ACDEFFBB5377CFAB565EFBB8A719" box="[903,1120,1292,1318]" pageId="59" pageNumber="60">Simferopol distr.</collectingMunicipality>
:
<specimenCount id="9D5CFD2DFFBB5377C852565EFAFDA719" box="[1150,1317,1292,1318]" pageId="59" pageNumber="60" type="generic">10 specimens</specimenCount>
,
<location id="8E85607FFFBB5377CD245666FE7DA771" LSID="urn:lsid:plazi:treatment:03F387B2FFBB5370CD2451AEFD71A641:8E85607FFFBB5377CD245666FE7DA771" box="[264,421,1332,1358]" country="Ukraine" municipality="Simferopol' distr." name="Krasnoles'ye" pageId="59" pageNumber="60" stateProvince="Crimea">Krasnolesye</location>
,
<quantity id="4CA29B41FFBB5377CD965666FDC8A771" box="[442,528,1332,1358]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="59" pageNumber="60" unit="m" value="400.0">
<elevation id="0077D197FFBB5377CD965666FDC8A771" box="[442,528,1332,1358]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="59" pageNumber="60" unit="m" value="400.0">400 m</elevation>
</quantity>
, in rotting wood and on mushrooms,
<date id="FFE41064FFBB5377C82A5666FBA1A771" box="[1030,1145,1332,1358]" pageId="59" pageNumber="60" value="1990-05-27">
<collectingDate id="EFA0E98CFFBB5377C82A5666FBA1A771" box="[1030,1145,1332,1358]" pageId="59" pageNumber="60" value="1990-05-27">27.v.1990</collectingDate>
</date>
(
<collectorName id="26AF5372FFBB5377C8BE5666FAF9A771" box="[1170,1313,1332,1358]" pageId="59" pageNumber="60">V.I.Gusarov</collectorName>
) (
<collectionCode id="ED4BAE61FFBB5377CD3C560EFE84A749" box="[272,348,1372,1398]" pageId="59" pageNumber="60">SPSU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B323CF9FFBB5377CD5A560EFD4AA7A1" ID-GBIF-Occurrence="3321919754" collectingDate="1978-08-06" collectionCode="SPSU" collectorName="S. Mosyakin" country="Ukraine" location="Ivano-Frankovsk" pageId="59" pageNumber="60" specimenCount="7" stateProvince="Ivano-Frankivs'k">
<emphasis id="B92EEAB6FFBB5377CD5A560EFD44A749" bold="true" box="[374,668,1372,1398]" pageId="59" pageNumber="60">
<collectingRegion id="499EF846FFBB5377CD5A560EFD96A749" box="[374,590,1372,1398]" country="Ukraine" name="Ivano-Frankivs'k" pageId="59" pageNumber="60">Ivano­Frankovsk</collectingRegion>
Reg.:
</emphasis>
<specimenCount id="9D5CFD2DFFBB5377CE98560EFC93A749" box="[692,843,1372,1398]" pageId="59" pageNumber="60" type="generic">7 specimens</specimenCount>
, Nadvornaya, on polypore mushrooms,
<date id="FFE41064FFBB5377CD2456D6FE5FA7A1" box="[264,391,1412,1438]" pageId="59" pageNumber="60" value="1978-08-06">
<collectingDate id="EFA0E98CFFBB5377CD2456D6FE5FA7A1" box="[264,391,1412,1438]" pageId="59" pageNumber="60" value="1978-08-06">6.viii.1978</collectingDate>
</date>
(
<collectorName id="26AF5372FFBB5377CDBA56D6FDFEA7A1" box="[406,550,1412,1438]" pageId="59" pageNumber="60">S.Mosyakin</collectorName>
) (
<collectionCode id="ED4BAE61FFBB5377CE1256D6FD52A7A1" box="[574,650,1412,1438]" pageId="59" pageNumber="60">SPSU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C340652FFFBB5377CD1456FEFCCCA7F9" box="[312,788,1452,1478]" pageId="59" pageNumber="60" type="diagnosis">
<paragraph id="8BE536A4FFBB5377CD1456FEFCCCA7F9" blockId="59.[264,1323,1211,1878]" box="[312,788,1452,1478]" pageId="59" pageNumber="60">
<emphasis id="B92EEAB6FFBB5377CD1456FEFE6FA7F9" bold="true" box="[312,439,1452,1478]" pageId="59" pageNumber="60">Diagnosis.</emphasis>
See
<bibRefCitation id="EFCB4B55FFBB5377CDD456FEFCC9A7F9" author="Benick, G. &amp; Lohse, G. A." box="[504,785,1452,1478]" pageId="59" pageNumber="60" pagination="72 - 220" refId="ref75834" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse (1974)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C340652FFFBB5370CD145686FB10A189" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60" type="discussion">
<paragraph id="8BE536A4FFBB5370CD145686FEFFA159" blockId="59.[264,1323,1211,1878]" lastBlockId="60.[264,1323,284,1150]" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
<emphasis id="B92EEAB6FFBB5377CD145686FE1BA7D1" bold="true" box="[312,451,1492,1518]" pageId="59" pageNumber="60">Discussion.</emphasis>
The types of
<taxonomicName id="4C5A4D27FFBB5377CE445686FCDAA7D2" authorityName="Gravenhorst" authorityYear="1802" box="[616,770,1492,1517]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBB5377CE445686FCDAA7D2" box="[616,770,1492,1517]" italics="true" pageId="59" pageNumber="60">Ale. nigritula</emphasis>
</taxonomicName>
were examined to clarify the status of
<taxonomicName id="4C5A4D27FFBB5377C8EA5686FAF3A7D2" authorityName="Bernhauer" authorityYear="1909" box="[1222,1323,1492,1517]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="frosti">
<emphasis id="B92EEAB6FFBB5377C8EA5686FAF3A7D2" box="[1222,1323,1492,1517]" italics="true" pageId="59" pageNumber="60">At. frosti</emphasis>
</taxonomicName>
and
<emphasis id="B92EEAB6FFBB5377CD1756AEFE0AA42A" box="[315,466,1532,1557]" italics="true" pageId="59" pageNumber="60">
<taxonomicName id="4C5A4D27FFBB5377CD1756AEFE16A42A" authorityName="Gravenhorst" authorityYear="1802" box="[315,462,1532,1557]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="nigritula">At. nigritula</taxonomicName>
.
</emphasis>
The historic collection at ZMHB which contains the Gravenhorst types has several specimens arranged as
<taxonomicName id="4C5A4D27FFBB5377CE8D5576FC51A402" authorityName="Gravenhorst" authorityYear="1802" box="[673,905,1572,1597]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBB5377CE8D5576FC51A402" box="[673,905,1572,1597]" italics="true" pageId="59" pageNumber="60">Aleochara nigritula</emphasis>
</taxonomicName>
in the following order: the label at the bottom of the drawer “var. c.
<taxonomicName id="4C5A4D27FFBB5377CEA1551EFC9BA459" authority="Gr." authorityName="Gravenhorst" authorityYear="1802" box="[653,835,1612,1638]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBB5377CEA1551EFCCCA45A" box="[653,788,1612,1637]" italics="true" pageId="59" pageNumber="60">A. nigritula</emphasis>
Gr.
</taxonomicName>
” is followed by
<specimenCount id="9D5CFD2DFFBB5377C828551EFB7FA459" box="[1028,1191,1612,1638]" pageId="59" pageNumber="60" type="generic">11 specimens</specimenCount>
: 1)
<specimenCount id="9D5CFD2DFFBB5377C8FE5519FB3EA460" box="[1234,1254,1611,1631]" pageId="59" pageNumber="60" type="female"></specimenCount>
(with the head and pronotum missing), labeled “
<taxonomicName id="4C5A4D27FFBB5377CF335526FC64A4B1" authority="Gr." authorityName="Gravenhorst" authorityYear="1802" box="[799,956,1652,1678]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBB5377CF335526FC5EA4B2" box="[799,902,1652,1677]" italics="true" pageId="59" pageNumber="60">nigritula</emphasis>
Gr.
</taxonomicName>
”; 2)
<specimenCount id="9D5CFD2DFFBB5377C82C5521FBCCA4B8" box="[1024,1044,1651,1671]" pageId="59" pageNumber="60" type="female"></specimenCount>
, without labels; 3)
<specimenCount id="9D5CFD2DFFBB5377C93C5521FAFCA4B8" box="[1296,1316,1651,1671]" pageId="59" pageNumber="60" type="female"></specimenCount>
, without labels; 4)
<specimenCount id="9D5CFD2DFFBB5377CDF055C9FE28A490" box="[476,496,1691,1711]" pageId="59" pageNumber="60" type="male"></specimenCount>
, without labels; 5)
<specimenCount id="9D5CFD2DFFBB5377CEFE55C9FD3EA490" box="[722,742,1691,1711]" pageId="59" pageNumber="60" type="female"></specimenCount>
, without labels; 6) a specimen labeled “
<emphasis id="B92EEAB6FFBB5377C89055CEFAF2A48A" box="[1212,1322,1692,1717]" italics="true" pageId="59" pageNumber="60">fungicola</emphasis>
Kraatz”; 7) a specimen without labels; 8) a specimen labeled “var. d”; 9) a specimen without labels; 10) a specimen labeled “Sard. Géné”; 11) a specimen labeled “Ural Er.”. These
<specimenCount id="9D5CFD2DFFBB5377CD7B5446FE2EA511" box="[343,502,1812,1838]" pageId="59" pageNumber="60" type="generic">11 specimens</specimenCount>
complete the row. The next row starts with a label at the bottom of the drawer “var.?” which is followed by
<specimenCount id="9D5CFD2DFFBB5377CEC1546EFC5BA569" box="[749,899,1852,1878]" pageId="59" pageNumber="60" type="generic">7 specimens</specimenCount>
: 1215) four unlabeled specimens, 16) a specimen labeled “Aust. …[illegible]”; 1718)
<specimenCount id="9D5CFD2DFFBC5370CF56524EFBF0A309" box="[890,1064,284,310]" pageId="60" pageNumber="61" type="generic">two specimens</specimenCount>
mounted on the same card labeled “Pallau
<taxonomicName id="4C5A4D27FFBC5370CDD25216FCE5A361" authority="Gve." authorityName="Gve." box="[510,829,324,350]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="variety" species="socialis" variety="nigra">
<emphasis id="B92EEAB6FFBC5370CDD25216FDA4A362" box="[510,636,324,349]" italics="true" pageId="60" pageNumber="61">H. socialis</emphasis>
var.
<emphasis id="B92EEAB6FFBC5370CE945216FD2FA362" box="[696,759,324,349]" italics="true" pageId="60" pageNumber="61">nigra</emphasis>
Gve.
</taxonomicName>
”. Apparently, the specimens listed above had been arranged by Erichson, who described
<taxonomicName id="4C5A4D27FFBC5370CF7A523EFB2BA3B9" authority="Erichson, 1839" authorityName="Erichson" authorityYear="1839" box="[854,1267,364,390]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="socialis">
<emphasis id="B92EEAB6FFBC5370CF7A523EFBE9A3BA" box="[854,1073,364,389]" italics="true" pageId="60" pageNumber="61">Homalota socialis</emphasis>
<bibRefCitation id="EFCB4B55FFBC5370C811523EFB2BA3B9" author="Erichson, W. F." box="[1085,1267,364,390]" pageId="60" pageNumber="61" pagination="1 - 400" refId="ref76845" refString="Erichson, W. F. (1839) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, pp. i - viii, 1 - 400." type="journal article" year="1839">Erichson, 1839</bibRefCitation>
</taxonomicName>
and listed
<taxonomicName id="4C5A4D27FFBC5370CD7852C6FD07A391" authority="Gravenhorst" authorityName="Gravenhorst" authorityYear="1802" box="[340,735,404,430]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBC5370CD7852C6FD9AA392" box="[340,578,404,429]" italics="true" pageId="60" pageNumber="61">Aleochara nigritula</emphasis>
Gravenhorst
</taxonomicName>
as “Var. c” of
<taxonomicName id="4C5A4D27FFBC5370CF8F52C6FBFDA392" authorityName="Erichson" authorityYear="1839" box="[931,1061,404,429]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="socialis">
<emphasis id="B92EEAB6FFBC5370CF8F52C6FBFDA392" box="[931,1061,404,429]" italics="true" pageId="60" pageNumber="61">H. socialis</emphasis>
</taxonomicName>
. Therefore only the specimens (Nos. 17) which follow the label “var. c” qualify as potential
<typeStatus id="54E18806FFBC5370C88C52EEFADEA3E9" box="[1184,1286,444,470]" pageId="60" pageNumber="61" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C5A4D27FFBC5370CD2452B6FE2BA3C2" authorityName="Gravenhorst" authorityYear="1802" box="[264,499,484,509]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBC5370CD2452B6FE2BA3C2" box="[264,499,484,509]" italics="true" pageId="60" pageNumber="61">Aleochara nigritula</emphasis>
</taxonomicName>
. My examination of these
<specimenCount id="9D5CFD2DFFBC5370CF6A52B6FBD6A3C1" box="[838,1038,484,510]" pageId="60" pageNumber="61" type="generic">seven specimens</specimenCount>
demonstrated that only specimen No. 1 belongs to
<taxonomicName id="4C5A4D27FFBC5370CE62515EFD06A01A" authorityName="Gravenhorst" authorityYear="1802" box="[590,734,524,549]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBC5370CE62515EFD06A01A" box="[590,734,524,549]" italics="true" pageId="60" pageNumber="61">At. nigritula</emphasis>
</taxonomicName>
in the commonly used sense (
<emphasis id="B92EEAB6FFBC5370C865515EFBA4A01A" box="[1097,1148,524,549]" italics="true" pageId="60" pageNumber="61">e. g.</emphasis>
,
<bibRefCitation id="EFCB4B55FFBC5370C8A0515EFE9EA071" author="Strand, A. &amp; Vik, A." pageId="60" pageNumber="61" pagination="327 - 335" refId="ref81239" refString="Strand, A. &amp; Vik, A. (1964) Die Genitalorgane der nordischen Arten der Gattung Atheta Thoms. (Col., Staphylinidae). Norsk Entomologisk Tidsskrift, 12 (5 - 8), 327 - 335, Taf. I - XXI." type="journal article" year="1964">Strand &amp; Vik 1964</bibRefCitation>
;
<bibRefCitation id="EFCB4B55FFBC5370CD7B5166FDB4A071" author="Benick, G. &amp; Lohse, G. A." box="[343,620,564,590]" pageId="60" pageNumber="61" pagination="72 - 220" refId="ref75834" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse 1974</bibRefCitation>
); specimens No. 2 and No. 5 belong to
<taxonomicName id="4C5A4D27FFBC5370C84B5166FE01A049" authority="(Thomson, 1856)" baseAuthorityName="Thomson" baseAuthorityYear="1856" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="pallidicornis">
<emphasis id="B92EEAB6FFBC5370C84B5166FAF2A072" box="[1127,1322,564,589]" italics="true" pageId="60" pageNumber="61">At. pallidicornis</emphasis>
(
<bibRefCitation id="EFCB4B55FFBC5370CD3C510EFE08A049" author="Thomson, C. G." box="[272,464,604,630]" pageId="60" pageNumber="61" pagination="91 - 107" refId="ref81328" refString="Thomson, C. G. (1856) Nagra nya arter af Insekt-slagtet Homalota. Ofversigt af Kongl. Vetenskaps- Akademiens Forhandlingar, 13 (4), 91 - 107." type="journal article" year="1856">Thomson, 1856</bibRefCitation>
)
</taxonomicName>
; specimen No. 3 belongs to
<taxonomicName id="4C5A4D27FFBC5370CF1D510EFB75A049" authority="(Kraatz, 1856)" authorityName="Gusarov" authorityYear="2003" baseAuthorityName="Kraatz" baseAuthorityYear="1856" box="[817,1197,604,630]" class="Insecta" family="Staphylinidae" genus="Dalotia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="coriaria">
<emphasis id="B92EEAB6FFBC5370CF1D510EFC2CA04A" box="[817,1012,604,629]" italics="true" pageId="60" pageNumber="61">Dalotia coriaria</emphasis>
(
<bibRefCitation id="EFCB4B55FFBC5370C829510EFB7CA049" author="Kraatz, G." box="[1029,1188,604,630]" pageId="60" pageNumber="61" pagination="1 - 376" refId="ref78849" refString="Kraatz, G. (1856) Naturgeschichte der Insecten Deutschlands. Erste Abteilung. Coleoptera. Bd. 2. Lief. 1 - 2. Verlag der Nicolaischen Buchhandlung, Berlin, pp. 1 - 376." type="book chapter" year="1856">Kraatz, 1856</bibRefCitation>
)
</taxonomicName>
; specimen No. 4 belongs to
<taxonomicName id="4C5A4D27FFBC5370CDF651D6FCC6A0A1" authority="(Baudi, 1848)" baseAuthorityName="Baudi" baseAuthorityYear="1848" box="[474,798,644,670]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="gagatina">
<emphasis id="B92EEAB6FFBC5370CDF651D6FDB4A0A2" box="[474,620,644,669]" italics="true" pageId="60" pageNumber="61">At. gagatina</emphasis>
(Baudi, 1848)
</taxonomicName>
; specimen No. 6 is not conspecific with
<taxonomicName id="4C5A4D27FFBC5370C92551D6FEB6A0FA" authorityName="Gravenhorst" authorityYear="1802" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBC5370C92551D6FEB6A0FA" italics="true" pageId="60" pageNumber="61">At. nigritula</emphasis>
</taxonomicName>
and has the pronotal pubescence of type I; specimen No. 7 is not conspecific with
<taxonomicName id="4C5A4D27FFBC5370CD245186FE40A0D2" authorityName="Gravenhorst" authorityYear="1802" box="[264,408,724,749]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBC5370CD245186FE40A0D2" box="[264,408,724,749]" italics="true" pageId="60" pageNumber="61">At. nigritula</emphasis>
</taxonomicName>
and has the pronotal pubescence of type II. Specimen No. 1 was selected as the
<typeStatus id="54E18806FFBC5370CD1551AEFE7CA129" box="[313,420,764,790]" pageId="60" pageNumber="61" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C5A4D27FFBC5370CDFB51AEFD1DA12A" authorityName="Gravenhorst" authorityYear="1802" box="[471,709,764,789]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBC5370CDFB51AEFD1DA12A" box="[471,709,764,789]" italics="true" pageId="60" pageNumber="61">Aleochara nigritula</emphasis>
</taxonomicName>
. The spermatheca of that specimen matches the illustrations by
<bibRefCitation id="EFCB4B55FFBC5370CD915076FD64A101" author="Strand, A. &amp; Vik, A." box="[445,700,804,830]" pageId="60" pageNumber="61" pagination="327 - 335" refId="ref81239" refString="Strand, A. &amp; Vik, A. (1964) Die Genitalorgane der nordischen Arten der Gattung Atheta Thoms. (Col., Staphylinidae). Norsk Entomologisk Tidsskrift, 12 (5 - 8), 327 - 335, Taf. I - XXI." type="journal article" year="1964">Strand and Vik (1964</bibRefCitation>
:
<figureCitation id="13612A21FFBC5370CEE25076FCF7A101" box="[718,815,804,830]" captionStart="FIGURES 116119" captionStartId="81.[264,383,1731,1755]" captionTargetBox="[346,1237,974,1699]" captionTargetId="figure-253@81.[341,1245,964,1705]" captionTargetPageId="81" captionText="FIGURES 116119. Abdominal segment 8 of Boreophilia angusticornis (Bernhauer) (male, lectotype of Atheta angusticornis (116117); and female, lectotype of Metaxya plutonica Casey (118119)). 116 — male tergum 8; 117 — male sternum 8; 118 — female tergum 8; 119 — female sternum 8. Scale bar 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5019864" httpUri="https://zenodo.org/record/5019864/files/figure.png" pageId="60" pageNumber="61">Fig. 117</figureCitation>
) and
<bibRefCitation id="EFCB4B55FFBC5370CF5C5076FB49A101" author="Benick, G. &amp; Lohse, G. A." box="[880,1169,804,830]" pageId="60" pageNumber="61" pagination="72 - 220" refId="ref75834" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick and Lohse (1974</bibRefCitation>
, p. 154:
<figureCitation id="13612A21FFBC5370C8D05076FEC1A159" captionStart="FIGURES 59" captionStartId="13.[264,382,1337,1361]" captionTargetBox="[298,1291,486,1318]" captionTargetId="figure-264@13.[288,1298,477,1322]" captionTargetPageId="13" captionText="FIGURES 59. Genitalia of Tinotus acerbus (Casey) (lectotype of Acrimea acerba (58) and lectotype of Acri. resecta (9)). 5 — median lobe, parameral view; 6 — median lobe with details of retracted internal sac, abparameral view; 7 — median lobe, lateral view; 8 — median lobe with details of retracted internal sac, lateral view; 9 — spermatheca. Scale bar 0.1 mm (9), 0.2 mm (5 8)." figureDoi="http://doi.org/10.5281/zenodo.5019795" httpUri="https://zenodo.org/record/5019795/files/figure.png" pageId="60" pageNumber="61">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8BE536A4FFBC5370CD145026FB10A189" blockId="60.[264,1323,284,1150]" pageId="60" pageNumber="61">
<taxonomicName id="4C5A4D27FFBC5370CD145026FD2DA1B1" authority="(Gravenhorst, 1802)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[312,757,884,910]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBC5370CD145026FE2EA1B2" box="[312,502,884,909]" italics="true" pageId="60" pageNumber="61">Atheta nigritula</emphasis>
(
<bibRefCitation id="EFCB4B55FFBC5370CE275026FD35A1B1" author="Gravenhorst, J. L. C." box="[523,749,884,910]" pageId="60" pageNumber="61" refId="ref77424" refString="Gravenhorst, J. L. C. (1802) Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera familias et species distribuit. C. Reichard, Brunswick, xxvi + 207 pp." type="book" year="1802">Gravenhorst, 1802</bibRefCitation>
)
</taxonomicName>
is the
<typeStatus id="54E18806FFBC5370CF7C5026FC55A1B1" box="[848,909,884,910]" pageId="60" pageNumber="61">types</typeStatus>
species of the subgenus
<taxonomicName id="4C5A4D27FFBC5370C8E55026FDC9A189" authority="Mulsant &amp; Rey, 1874 a" authorityName="Mulsant &amp; Rey" authorityYear="1874" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="subGenus" subGenus="Tetropla">
<emphasis id="B92EEAB6FFBC5370C8E55026FAF2A1B2" box="[1225,1322,884,909]" italics="true" pageId="60" pageNumber="61">Tetropla</emphasis>
<bibRefCitation id="EFCB4B55FFBC5370CD2450CEFDC9A189" author="Mulsant, M. E. &amp; Rey, C." box="[264,529,924,950]" pageId="60" pageNumber="61" pagination="33 - 738" refId="ref79932" refString="Mulsant, M. E. &amp; Rey, C. (1874 a) Tribu des Brevipennes. Famille des Aleochariens. Septieme branche: Myrmedoniaires. Annales de la Societe d'Agriculture, Histoire Naturelle et Arts Utiles de Lyon, serie 4. 6 [1873], 33 - 738." type="journal article" year="1874">Mulsant &amp; Rey, 1874a</bibRefCitation>
</taxonomicName>
(by subsequent designation, fixed by
<bibRefCitation id="EFCB4B55FFBC5370CFE250CEFB1CA189" author="Blackwelder, R. E." box="[974,1220,924,950]" pageId="60" pageNumber="61" pagination="1 - 483" refId="ref76128" refString="Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae, with an essay on genotypy. U. S. National Museum Bulletin, 200, 1 - 483." type="journal article" year="1952">Blackwelder (1952))</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C340652FFFBC5370CD145096FD71A641" pageId="60" pageNumber="61" type="distribution">
<paragraph id="8BE536A4FFBC5370CD145096FD71A641" blockId="60.[264,1323,284,1150]" pageId="60" pageNumber="61">
<emphasis id="B92EEAB6FFBC5370CD145096FE00A1E1" bold="true" box="[312,472,964,990]" pageId="60" pageNumber="61">Distribution.</emphasis>
<taxonomicName id="4C5A4D27FFBC5370CDCA5096FD78A1E2" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[486,672,964,989]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBC5370CDCA5096FD78A1E2" box="[486,672,964,989]" italics="true" pageId="60" pageNumber="61">Atheta nigritula</emphasis>
</taxonomicName>
is a common European species associated with fungi. It has been reported from North America (
<bibRefCitation id="EFCB4B55FFBC5370CECF50BEFC7DA639" author="Bernhauer, M." box="[739,933,1004,1030]" pageId="60" pageNumber="61" pagination="381 - 405" refId="ref76008" refString="Bernhauer, M. (1907) Neue Aleocharini aus Nordamerika. (Col.) (3. Stuck.). Deutsche Entomologische Zeitschrift, 1907 (4), 381 - 405." type="journal article" year="1907">Bernhauer 1907</bibRefCitation>
;
<bibRefCitation id="EFCB4B55FFBC5370CF9E50BEFAF8A639" author="Bernhauer, M. &amp; Scheerpeltz, O." box="[946,1312,1004,1030]" pageId="60" pageNumber="61" pagination="499 - 988" refId="ref76079" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988." type="book chapter" year="1926">Bernhauer &amp; Scheerpeltz 1926</bibRefCitation>
) but, to the best of my knowledge, does not occur there. The examined North American specimens formerly identified as
<taxonomicName id="4C5A4D27FFBC5370CEA1576EFCC4A66A" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[653,796,1084,1109]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="nigritula">
<emphasis id="B92EEAB6FFBC5370CEA1576EFCC4A66A" box="[653,796,1084,1109]" italics="true" pageId="60" pageNumber="61">At. nigritula</emphasis>
</taxonomicName>
turned out to be
<taxonomicName id="4C5A4D27FFBC5370CFCD576EFAC9A669" authority="Bernhauer, 1909" authorityName="Bernhauer" authorityYear="1909" box="[993,1297,1084,1110]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="frosti">
<emphasis id="B92EEAB6FFBC5370CFCD576EFB9FA66A" box="[993,1095,1084,1109]" italics="true" pageId="60" pageNumber="61">At. frosti</emphasis>
<bibRefCitation id="EFCB4B55FFBC5370C861576EFAC9A669" author="Bernhauer, M." box="[1101,1297,1084,1110]" pageId="60" pageNumber="61" pagination="515 - 528" refId="ref76044" refString="Bernhauer, M. (1909) Neue Aleocharini aus Nordamerika. (Col.) (4. Stuck.) Deutsche Entomologische Zeitschrift, 1909 (4), 515 - 528." type="journal article" year="1909">Bernhauer, 1909</bibRefCitation>
</taxonomicName>
, a closely related but separate species.
</paragraph>
</subSubSection>
</treatment>
</document>