261 lines
29 KiB
XML
261 lines
29 KiB
XML
<document id="7E12D419BB29D594A8C0F7575099DB38" ID-CLB-Dataset="29450" ID-DOI="10.11646/zootaxa.4472.2.3" ID-GBIF-Dataset="b9db02fb-a3bf-464f-a060-8da8ad880060" ID-ISSN="1175-5326" ID-Zenodo-Dep="1440175" ID-ZooBank="1468154D-628C-456B-A237-5E130AB3E9FF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538372054204" checkinUser="plazi" docAuthor="Gąsiorek, Piotr, Stec, Daniel, Zawierucha, Krzysztof, Kristensen, Reinhardt Møbjerg & Michalczyk, Łukasz" docDate="2018" docId="03EA879FB95A8818FF7391DEBE671D10" docLanguage="en" docName="zootaxa.4472.2.3.pdf" docOrigin="Zootaxa 4472 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Testechiniscus Kristensen 1987" docType="treatment" docVersion="8" lastPageNumber="266" masterDocId="FFD3FFE7B95E881DFFE4977BBC071E28" masterDocTitle="Revision of Testechiniscus Kristensen, 1987 (Heterotardigrada: Echiniscidae) refutes the polar-temperate distribution of the genus" masterLastPageNumber="297" masterPageNumber="261" pageNumber="265" updateTime="1698559050273" updateUser="ExternalLinkService">
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<mods:mods id="061A6E0C24D0D0F8FC97B5A4AC78954B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4323764E44F654AB5CAD980D625B679C">
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<mods:title id="A90E974A86693A5F12E4F052B1DC2C00">Revision of Testechiniscus Kristensen, 1987 (Heterotardigrada: Echiniscidae) refutes the polar-temperate distribution of the genus</mods:title>
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<mods:name id="7BB6DCCC32ED8690327D71B76181B2D7" type="personal">
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<mods:namePart id="B29E608589932BC69CE4E47137B8D222">Gąsiorek, Piotr</mods:namePart>
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<mods:roleTerm id="FE4290856ED964C445B8278CFDD34339">Author</mods:roleTerm>
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<mods:namePart id="B5D65B20A764527D878818FB7759BB88">Stec, Daniel</mods:namePart>
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<mods:name id="805FAF546E6B5443C5806847A11C16B8" type="personal">
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<mods:roleTerm id="A93593AF75FC9BC32C22B32E84C18C6F">Author</mods:roleTerm>
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<mods:namePart id="D353886B64361547A4383A504BBB3A8C">Zawierucha, Krzysztof</mods:namePart>
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</mods:name>
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<mods:roleTerm id="487A0E5F04663C7E7D43A31F3F3D7348">Author</mods:roleTerm>
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<mods:namePart id="02912C2F7EBB6465B04AE24505F23BD0">Kristensen, Reinhardt Møbjerg</mods:namePart>
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</mods:name>
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<mods:name id="160BFB5B5DA2781460AE0FA6990947B5" type="personal">
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<mods:namePart id="D811E891DB39FD4C13C7502C9208D312">Michalczyk, Łukasz</mods:namePart>
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<mods:typeOfResource id="F20274D044A01134509ED4E55AE6E014">text</mods:typeOfResource>
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<mods:title id="A88572F693C56F55CD3949D110D698FC">Zootaxa</mods:title>
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<mods:part id="48A74EE293715AE3313F0D3B0A28B6AC">
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<mods:date id="2611567302D43C389E5C40111D07C78A">2018</mods:date>
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<mods:number id="88F861E2AD7A61710A71A58166DB59F7">2018-09-10</mods:number>
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<mods:detail id="51F729A70409F626B64B6DA266B21853" type="volume">
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<mods:number id="5B5289C156022D449328747DF1378D7D">4472</mods:number>
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<mods:number id="ABFA93DAF2B398D8EE56FD07401C95FD">2</mods:number>
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<mods:start id="B98707DBC9ACC7F1DF98EF9C08296C67">261</mods:start>
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<mods:identifier id="70A545E4492DE4A3627A14CE6F4A9D75" type="DOI">10.11646/zootaxa.4472.2.3</mods:identifier>
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<treatment id="03EA879FB95A8818FF7391DEBE671D10" ID-DOI="http://doi.org/10.5281/zenodo.5975776" ID-GBIF-Taxon="148421337" ID-Zenodo-Dep="5975776" LSID="urn:lsid:plazi:treatment:03EA879FB95A8818FF7391DEBE671D10" httpUri="http://treatment.plazi.org/id/03EA879FB95A8818FF7391DEBE671D10" lastPageId="5" lastPageNumber="266" pageId="4" pageNumber="265">
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<subSubSection id="C3596502B95A8819FF7391DEBE781897" box="[151,639,1701,1727]" pageId="4" pageNumber="265" type="nomenclature">
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<paragraph id="8BFC3689B95A8819FF7391DEBE781897" blockId="4.[151,639,1701,1727]" box="[151,639,1701,1727]" pageId="4" pageNumber="265">
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<heading id="D0B481E5B95A8819FF7391DEBE781897" bold="true" box="[151,639,1701,1727]" fontSize="11" level="1" pageId="4" pageNumber="265" reason="1">
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<emphasis id="B937EA9BB95A8819FF7391DEBE781897" bold="true" box="[151,639,1701,1727]" pageId="4" pageNumber="265">
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Genus:
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<taxonomicName id="4C434D0AB95A8819FF1C91DEBE781897" ID-CoL="646Z4" authority="Kristensen, 1987" authorityName="Kristensen" authorityYear="1987" box="[248,639,1701,1727]" class="Heterotardigrada" family="Echiniscidae" genus="Testechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="265" phylum="Tardigrada" rank="genus">
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<emphasis id="B937EA9BB95A8819FF1C91DEBDA01897" bold="true" box="[248,423,1701,1727]" italics="true" pageId="4" pageNumber="265">Testechiniscus</emphasis>
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<bibRefCitation id="EFD24B78B95A8819FE4A91DEBE781897" box="[430,639,1701,1727]" pageId="4" pageNumber="265" refId="ref12358" refString="Kristensen, R. M. (1987) Generic revision of the Echiniscidae (Heterotardigrada), with a discussion of the origin of the family. In: Bertolani, R. (Ed.), Biology of Tardigrada. Selected Symposia and Monographs. U. Z. I. Mucci, Modena, pp. 261 - 335." type="book chapter">Kristensen, 1987</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3596502B95A8818FF739197BD581C48" lastPageId="5" lastPageNumber="266" pageId="4" pageNumber="265" type="diagnosis">
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<paragraph id="8BFC3689B95A8819FF739197BE7219BD" blockId="4.[151,1436,1772,2013]" pageId="4" pageNumber="265">
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<emphasis id="B937EA9BB95A8819FF739197BDDF192D" bold="true" box="[151,472,1772,1797]" pageId="4" pageNumber="265">Amended genus diagnosis:</emphasis>
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Medium-sized echiniscids with black, crystalline eyes. Rigid buccal tube with large cuticular stylet supports. Appendaged,
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<emphasis id="B937EA9BB95A8819FDBE906ABE7B1900" box="[602,636,1809,1832]" italics="true" pageId="4" pageNumber="265">i.e.</emphasis>
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having both cephalic and trunk cirri. Two pairs of segmental plates, unpaired scapular and caudal plates. Three median plates. Cuticular sculpture composed of large true round or polygonal pores that gradually become reticulum. Incisions (notches) on caudal (terminal) plate. Eight rows of ventral plates. No pseudosegmental plates.
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</paragraph>
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<paragraph id="8BFC3689B95A8818FF2390E4B9631F17" blockId="4.[151,1436,1772,2013]" lastBlockId="5.[151,1437,151,824]" lastPageId="5" lastPageNumber="266" pageId="4" pageNumber="265">
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<emphasis id="B937EA9BB95A8819FF2390E4BE651990" bold="true" box="[199,610,1951,1976]" pageId="4" pageNumber="265">Differential diagnosis. Phenotypic.</emphasis>
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<taxonomicName id="4C434D0AB95A8819FD8D90E4BF0D1990" authorityName="Kristensen" authorityYear="1987" box="[617,778,1951,1976]" class="Heterotardigrada" family="Echiniscidae" genus="Testechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="265" phylum="Tardigrada" rank="genus">
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<emphasis id="B937EA9BB95A8819FD8D90E4BF0D1990" box="[617,778,1951,1976]" italics="true" pageId="4" pageNumber="265">Testechiniscus</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName id="4C434D0AB95A8819FC7A90E4B82A1990" authorityName="Thulin" authorityYear="1928" box="[926,1069,1951,1976]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="265" phylum="Tardigrada" rank="genus">
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<emphasis id="B937EA9BB95A8819FC7A90E4B82A1990" box="[926,1069,1951,1976]" italics="true" pageId="4" pageNumber="265">Bryodelphax</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C434D0AB95A8819FBD890E4B96B1990" authority="Marcus, 1936" authorityName="Marcus" authorityYear="1936" box="[1084,1388,1951,1976]" class="Heterotardigrada" family="Echiniscidae" genus="Bryochoerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="265" phylum="Tardigrada" rank="genus">
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<emphasis id="B937EA9BB95A8819FBD890E4B8CC1990" box="[1084,1227,1951,1976]" italics="true" pageId="4" pageNumber="265">Bryochoerus</emphasis>
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<bibRefCitation id="EFD24B78B95A8819FB3690DBB96B1990" box="[1234,1388,1951,1976]" pageId="4" pageNumber="265" refId="ref12558" refString="Marcus, E. (1936) Tardigrada. Das Tierreich, 66, 1 - 340." type="journal article">Marcus, 1936</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4C434D0AB95A8819FF7390BFBD1719F5" box="[151,272,1988,2013]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="265" phylum="Tardigrada" rank="genus">
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<emphasis id="B937EA9BB95A8819FF7390BFBD1719F5" box="[151,272,1988,2013]" italics="true" pageId="4" pageNumber="265">Echiniscus</emphasis>
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</taxonomicName>
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in having black eyes. Moreover,
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<taxonomicName id="4C434D0AB95A8819FD6790BFBF2319F5" authorityName="Kristensen" authorityYear="1987" box="[643,804,1988,2013]" class="Heterotardigrada" family="Echiniscidae" genus="Testechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="265" phylum="Tardigrada" rank="genus">
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<emphasis id="B937EA9BB95A8819FD6790BFBF2319F5" box="[643,804,1988,2013]" italics="true" pageId="4" pageNumber="265">Testechiniscus</emphasis>
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</taxonomicName>
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can be separated from
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<taxonomicName id="4C434D0AB95A8819FBC290BFB8B219F5" authorityName="Thulin" authorityYear="1928" box="[1062,1205,1988,2013]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="265" phylum="Tardigrada" rank="genus">
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<emphasis id="B937EA9BB95A8819FBC290BFB8B219F5" box="[1062,1205,1988,2013]" italics="true" pageId="4" pageNumber="265">Bryodelphax</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C434D0AB95A8819FB0E90BFB97E19F5" authorityName="Marcus" authorityYear="1936" box="[1258,1401,1988,2013]" class="Heterotardigrada" family="Echiniscidae" genus="Bryochoerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="265" phylum="Tardigrada" rank="genus">
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<emphasis id="B937EA9BB95A8819FB0E90BFB97E19F5" box="[1258,1401,1988,2013]" italics="true" pageId="4" pageNumber="265">Bryochoerus</emphasis>
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</taxonomicName>
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by the presence of incisions on the caudal plate and cirri in positions other than
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<emphasis id="B937EA9BB95B8818FBD797E3B8431E87" box="[1075,1092,152,175]" italics="true" pageId="5" pageNumber="266">A</emphasis>
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, and from
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<taxonomicName id="4C434D0AB95B8818FB2997ECB9411E98" box="[1229,1350,151,176]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="genus">
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<emphasis id="B937EA9BB95B8818FB2997ECB9411E98" box="[1229,1350,151,176]" italics="true" pageId="5" pageNumber="266">Echiniscus</emphasis>
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</taxonomicName>
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by the presence of ventral armature covering the entire venter. Furthermore,
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<taxonomicName id="4C434D0AB95B8818FC4697C7B8501EFD" box="[930,1111,188,213]" class="Heterotardigrada" family="Echiniscidae" genus="Diploechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="genus">
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<emphasis id="B937EA9BB95B8818FC4697C7B8501EFD" box="[930,1111,188,213]" italics="true" pageId="5" pageNumber="266">Diploechiniscus</emphasis>
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</taxonomicName>
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has only two rows of ventral plates (subcephalic and genital) and double cuticular sculpture absent in
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<taxonomicName id="4C434D0AB95B8818FC0397A4B88F1ED0" authorityName="Kristensen" authorityYear="1987" box="[999,1160,223,248]" class="Heterotardigrada" family="Echiniscidae" genus="Testechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="genus">
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<emphasis id="B937EA9BB95B8818FC0397A4B88F1ED0" box="[999,1160,223,248]" italics="true" pageId="5" pageNumber="266">Testechiniscus</emphasis>
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</taxonomicName>
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. Finally,
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<taxonomicName id="4C434D0AB95B8818FB1D97A4BD2F1F34" authority="Thulin, 1928" authorityName="Thulin" authorityYear="1928" class="Heterotardigrada" family="Echiniscidae" genus="Hypechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="genus">
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<emphasis id="B937EA9BB95B8818FB1D97A4B99B1ED0" box="[1273,1436,223,248]" italics="true" pageId="5" pageNumber="266">Hypechiniscus</emphasis>
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<bibRefCitation id="EFD24B78B95B8818FF73967FBD2F1F34" box="[151,296,260,285]" pageId="5" pageNumber="266" refId="ref14646" refString="Thulin, G. (1928) Uber die Phylogenie und das System der Tardigraden. Hereditas, 11, 207 - 266. https: // doi. org / 10.1111 / j. 1601 - 5223.1928. tb 02488. x" type="journal article">Thulin, 1928</bibRefCitation>
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</taxonomicName>
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lacks trunk cirri (sometimes only an autapomorphic mediodorsal cirrus is present) and ventral plates; dorsal armature is very weakly sclerotised compared to large and strongly sclerotised plates in
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<taxonomicName id="4C434D0AB95B8818FB58965CB95A1F68" authorityName="Kristensen" authorityYear="1987" box="[1212,1373,295,320]" class="Heterotardigrada" family="Echiniscidae" genus="Testechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="genus">
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<emphasis id="B937EA9BB95B8818FB58965CB95A1F68" box="[1212,1373,295,320]" italics="true" pageId="5" pageNumber="266">Testechiniscus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BFC3689B95B8818FF239637BD581C48" blockId="5.[151,1437,151,824]" pageId="5" pageNumber="266">
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<emphasis id="B937EA9BB95B8818FF239637BD4E1F4D" bold="true" box="[199,329,332,357]" pageId="5" pageNumber="266">Genotypic.</emphasis>
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Based on the currently available sequences, the 18S rRNA p-distances within the genus
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<taxonomicName id="4C434D0AB95B8818FF739614BD3F1FA0" authorityName="Kristensen" authorityYear="1987" box="[151,312,367,392]" class="Heterotardigrada" family="Echiniscidae" genus="Testechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="genus">
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<emphasis id="B937EA9BB95B8818FF739614BD3F1FA0" box="[151,312,367,392]" italics="true" pageId="5" pageNumber="266">Testechiniscus</emphasis>
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</taxonomicName>
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vary from 0.2% to 0.8%, and between
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<taxonomicName id="4C434D0AB95B8818FD149614BF961FA0" authorityName="Kristensen" authorityYear="1987" box="[752,913,367,392]" class="Heterotardigrada" family="Echiniscidae" genus="Testechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="genus">
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<emphasis id="B937EA9BB95B8818FD149614BF961FA0" box="[752,913,367,392]" italics="true" pageId="5" pageNumber="266">Testechiniscus</emphasis>
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</taxonomicName>
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spp. and other currently recognised echiniscid genera they vary from 4.6% (
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<taxonomicName id="4C434D0AB95B8818FE0096EEBF331F84" authority="(Doyere, 1840)" baseAuthorityName="Doyere" baseAuthorityYear="1840" box="[484,820,404,429]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="species" species="granulatus">
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<emphasis id="B937EA9BB95B8818FE0096EEBE7A1F85" box="[484,637,404,429]" italics="true" pageId="5" pageNumber="266">E. granulatus</emphasis>
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(
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<bibRefCitation id="EFD24B78B95B8818FD6996EFBF2B1F84" box="[653,812,404,428]" pageId="5" pageNumber="266" refId="ref11163" refString="Doyere, M. L. (1840) Memorie sur les Tardigrades. Annales des Sciences Naturelles. Zoologie, 14, 269 - 362." type="journal article">Doyère, 1840</bibRefCitation>
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)
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</taxonomicName>
|
|
,
|
|
<accessionNumber id="9410AB6AB95B8818FCA496EFBFC41F85" box="[832,963,404,429]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MG016454" pageId="5" pageNumber="266">MG016454</accessionNumber>
|
|
) to 15.7% (
|
|
<taxonomicName id="4C434D0AB95B8818FBAE96EFBCD51FF8" authority="Thulin, 1928" authorityName="Thulin" authorityYear="1928" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="species" species="parvulus">
|
|
<emphasis id="B937EA9BB95B8818FBAE96EFB9451F85" box="[1098,1346,404,429]" italics="true" pageId="5" pageNumber="266">Bryodelphax parvulus</emphasis>
|
|
<bibRefCitation id="EFD24B78B95B8818FAAE96EFBCD51FF8" pageId="5" pageNumber="266" refId="ref14646" refString="Thulin, G. (1928) Uber die Phylogenie und das System der Tardigraden. Hereditas, 11, 207 - 266. https: // doi. org / 10.1111 / j. 1601 - 5223.1928. tb 02488. x" type="journal article">Thulin, 1928</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<accessionNumber id="9410AB6AB95B8818FF3B96C3BD641FF8" box="[223,355,440,464]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HM193371" pageId="5" pageNumber="266">HM193371</accessionNumber>
|
|
, and
|
|
<taxonomicName id="4C434D0AB95B8818FE4796C2BF311FF8" authority="Gasiorek et al., 2017" authorityName="Gasiorek et al." authorityYear="2017" box="[419,822,439,464]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="species" species="maculatus">
|
|
<emphasis id="B937EA9BB95B8818FE4796C2BE301FF8" box="[419,567,439,464]" italics="true" pageId="5" pageNumber="266">B. maculatus</emphasis>
|
|
Gąsiorek
|
|
<emphasis id="B937EA9BB95B8818FD5496C2BEE41FF8" box="[688,739,439,464]" italics="true" pageId="5" pageNumber="266">et al</emphasis>
|
|
., 2017
|
|
</taxonomicName>
|
|
,
|
|
<accessionNumber id="9410AB6AB95B8818FCA096CCBFC21FF8" box="[836,965,439,464]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY609137" pageId="5" pageNumber="266">KY609137</accessionNumber>
|
|
). The corresponding comparisons for the 28S rRNA fragment yield higher minimal but lower maximal p-values between genera; respectively from 6.3% (
|
|
<taxonomicName id="4C434D0AB95B8818FA6096A6BDB51C30" authority="Cuenot, 1932" authorityName="Cuenot" authorityYear="1932" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="species" species="trisetosus">
|
|
<emphasis id="B937EA9BB95B8818FA6096A6BD041C30" italics="true" pageId="5" pageNumber="266">E. trisetosus</emphasis>
|
|
<bibRefCitation id="EFD24B78B95B8818FEF6957BBDB51C30" box="[274,434,512,536]" pageId="5" pageNumber="266" refId="ref10828" refString="Cuenot, L. (1932) Tardigrades. In: Lechevalier, P. (Ed.), Faune de France, 24, pp. 1 - 96." type="journal article">Cuénot, 1932</bibRefCitation>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C434D0AB95B8818FE1F957ABF521C30" authority="Murray, 1910" authorityName="Murray" authorityYear="1910" box="[507,853,511,536]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="species" species="canadensis">
|
|
<emphasis id="B937EA9BB95B8818FE1F957ABEA61C30" box="[507,673,511,536]" italics="true" pageId="5" pageNumber="266">E. canadensis</emphasis>
|
|
<bibRefCitation id="EFD24B78B95B8818FD54957BBF521C30" box="[688,853,512,536]" pageId="5" pageNumber="266" refId="ref13216" refString="Murray, J. (1910) Tardigrada. British Antarctic Expedition 1907 - 1909. Reports on the Scientific Investigations, 1, Biology (Part V), 83 - 187." type="book chapter">Murray, 1910</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<accessionNumber id="9410AB6AB95B8818FC8D9684BFD01C30" box="[873,983,511,536]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ435781" pageId="5" pageNumber="266">FJ435781</accessionNumber>
|
|
and
|
|
<accessionNumber id="9410AB6AB95B8818FBC49684B8951C30" box="[1056,1170,511,536]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ435784" pageId="5" pageNumber="266">FJ435784</accessionNumber>
|
|
,
|
|
<accessionNumber id="9410AB6AB95B8818FB429684B9101C30" box="[1190,1303,511,536]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ435786" pageId="5" pageNumber="266">FJ435786</accessionNumber>
|
|
) to 13.9% (
|
|
<taxonomicName id="4C434D0AB95B8818FF44955FBE731C15" authority="Petersen, 1951" authorityName="Petersen" authorityYear="1951" box="[160,628,548,573]" class="Heterotardigrada" family="Echiniscidae" genus="Pseudechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="species" species="facettalis">
|
|
<emphasis id="B937EA9BB95B8818FF44955FBDC01C15" box="[160,455,548,573]" italics="true" pageId="5" pageNumber="266">Pseudechiniscus facettalis</emphasis>
|
|
<bibRefCitation id="EFD24B78B95B8818FE29955EBE731C15" box="[461,628,548,573]" pageId="5" pageNumber="266" refId="ref13434" refString="Petersen, B. (1951) The Tardigrade Fauna of Greenland. Meddelelser om GrOnland, 150 (5), 1 - 94." type="journal article">Petersen, 1951</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<accessionNumber id="9410AB6AB95B8818FD64955FBEF31C15" box="[640,756,548,573]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ435788" pageId="5" pageNumber="266">FJ435788</accessionNumber>
|
|
). Full p-distance matrices are provided in the Supplementary Materials (SM.2).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C3596502B95B8818FF239517BEEC1CD8" pageId="5" pageNumber="266" type="description">
|
|
<paragraph id="8BFC3689B95B8818FF239517BEEC1CD8" blockId="5.[151,1437,151,824]" pageId="5" pageNumber="266">
|
|
<emphasis id="B937EA9BB95B8818FF239517BD6E1CAD" bold="true" box="[199,361,620,645]" pageId="5" pageNumber="266">Composition:</emphasis>
|
|
<emphasis id="B937EA9BB95B8818FE979517BEAA1CAC" box="[371,685,620,645]" italics="true" pageId="5" pageNumber="266">
|
|
<taxonomicName id="4C434D0AB95B8818FE979517BE5C1CAC" box="[371,603,620,645]" class="Heterotardigrada" family="Echiniscidae" genus="Testechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="subSpecies" subSpecies="sensu">
|
|
Testechiniscus
|
|
<taxonomicNameLabel id="A20457E0B95B8818FDF99516BE5C1CAC" box="[541,603,621,644]" pageId="5" pageNumber="266" sensu="stricto">sensu</taxonomicNameLabel>
|
|
</taxonomicName>
|
|
stricto
|
|
</emphasis>
|
|
:
|
|
<taxonomicName id="4C434D0AB95B8818FD5B9516B8231CAD" authority="spitsbergensis" authorityName="spitsbergensis (Scourfield" authorityYear="1897" box="[703,1060,620,645]" class="Heterotardigrada" family="Echiniscidae" genus="Testechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="subSpecies" species="spitsbergensis" subSpecies="spitsbergensis">
|
|
<emphasis id="B937EA9BB95B8818FD5B9516B8231CAD" box="[703,1060,620,645]" italics="true" pageId="5" pageNumber="266">T. spitsbergensis spitsbergensis</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C434D0AB95B8818FBD09516B9621CAD" authority="Gąsiorek & Stec & Zawierucha & Kristensen & Michalczyk, 2018" authorityName="Gąsiorek & Stec & Zawierucha & Kristensen & Michalczyk" authorityYear="2018" box="[1076,1381,620,645]" class="Heterotardigrada" family="Echiniscidae" genus="Testechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="subSpecies" species="spitsbergensis" status="subsp. nov." subSpecies="tropicalis">
|
|
<emphasis id="B937EA9BB95B8818FBD09516B9621CAD" box="[1076,1381,620,645]" italics="true" pageId="5" pageNumber="266">T. spitsbergensis tropicalis</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A20457E0B95B8818FA8B9517BCCF1C80" pageId="5" pageNumber="266" rank="subSpecies">
|
|
<emphasis id="B937EA9BB95B8818FA8B9517BCCF1C80" bold="true" pageId="5" pageNumber="266">
|
|
<taxonomicName id="4C434D0AB95B8818FA8B9517BCC31C80" pageId="5" pageNumber="266" rank="subSpecies" subSpecies="nov">ssp. nov</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
</taxonomicNameLabel>
|
|
and
|
|
<taxonomicName id="4C434D0AB95B8818FEE495EABD8C1C80" authority="Gąsiorek & Stec & Zawierucha & Kristensen & Michalczyk, 2018" authorityName="Gąsiorek & Stec & Zawierucha & Kristensen & Michalczyk" authorityYear="2018" box="[256,395,655,680]" class="Heterotardigrada" family="Echiniscidae" genus="Testechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="species" sensu="lato" species="laterculus">
|
|
<emphasis id="B937EA9BB95B8818FEE495EABD8C1C80" box="[256,395,655,680]" italics="true" pageId="5" pageNumber="266">T. laterculus</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
<emphasis id="B937EA9BB95B8818FE7E95F4BEB31C80" box="[410,692,655,680]" italics="true" pageId="5" pageNumber="266">
|
|
<taxonomicName id="4C434D0AB95B8818FE7E95F4BE871C80" box="[410,640,655,680]" class="Heterotardigrada" family="Echiniscidae" genus="Testechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="subSpecies" subSpecies="sensu">
|
|
Testechiniscus
|
|
<taxonomicNameLabel id="A20457E0B95B8818FDA695EABE871C80" box="[578,640,657,680]" pageId="5" pageNumber="266" sensu="lato">sensu</taxonomicNameLabel>
|
|
</taxonomicName>
|
|
lato
|
|
</emphasis>
|
|
includes
|
|
<taxonomicName id="4C434D0AB95B8818FCC495EABFAE1C80" baseAuthorityName="Richters" baseAuthorityYear="1907" box="[800,937,657,680]" class="Heterotardigrada" family="Echiniscidae" genus="Testechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="species" species="macronyx">
|
|
<emphasis id="B937EA9BB95B8818FCC495EABFAE1C80" box="[800,937,657,680]" italics="true" pageId="5" pageNumber="266">T. macronyx</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C434D0AB95B8818FC0695EAB88C1C80" baseAuthorityName="Murray" baseAuthorityYear="1906" box="[994,1163,655,680]" class="Heterotardigrada" family="Echiniscidae" genus="Testechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="266" phylum="Tardigrada" rank="species" species="meridionalis">
|
|
<emphasis id="B937EA9BB95B8818FC0695EAB88C1C80" box="[994,1163,655,680]" italics="true" pageId="5" pageNumber="266">T. meridionalis</emphasis>
|
|
</taxonomicName>
|
|
(our analyses described below show that the two last species do not belong to the genus. However, their taxonomic status remains unchanged, pending formal taxonomic designations).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C3596502B95B8818FF239587BE671D10" pageId="5" pageNumber="266" type="etymology">
|
|
<paragraph id="8BFC3689B95B8818FF239587BE671D10" blockId="5.[151,1437,151,824]" pageId="5" pageNumber="266">
|
|
<emphasis id="B937EA9BB95B8818FF239587BD561D3D" bold="true" box="[199,337,764,789]" pageId="5" pageNumber="266">Etymology:</emphasis>
|
|
From Latin
|
|
<emphasis id="B937EA9BB95B8818FE3A9586BE141D3C" box="[478,531,765,788]" italics="true" pageId="5" pageNumber="266">testa</emphasis>
|
|
= shell, armour. The name refers to well-developed plates covering both dorsum and venter of animals (
|
|
<bibRefCitation id="EFD24B78B95B8818FE739464BE531D10" box="[407,596,799,824]" pageId="5" pageNumber="266" refId="ref12358" refString="Kristensen, R. M. (1987) Generic revision of the Echiniscidae (Heterotardigrada), with a discussion of the origin of the family. In: Bertolani, R. (Ed.), Biology of Tardigrada. Selected Symposia and Monographs. U. Z. I. Mucci, Modena, pp. 261 - 335." type="book chapter">Kristensen 1987</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |