584 lines
80 KiB
XML
584 lines
80 KiB
XML
<document id="2280010854ED56196C513BF9A3C28D53" ID-DOI="10.11646/megataxa.11.1.1" ID-ISSN="2703-3090" ID-Zenodo-Dep="10993768" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1713461317092" checkinUser="felipe" docAuthor="Kerr, Isaac A. R., Camens, Aaron B., Van Zoelen, Jacob D., Worthy, Trevor H. & Prideaux, Gavin J." docDate="2024" docId="03E587FDFF50D58BFCA2728CFA47FA09" docLanguage="en" docName="Megataxa.11.1.001-261.pdf" docOrigin="Megataxa 11 (1)" docStyle="DocumentStyle:50B6B96D96A55C21793B52C23E95B310.5:Megataxa.2020-.journal_article" docStyleId="50B6B96D96A55C21793B52C23E95B310" docStyleName="Megataxa.2020-.journal_article" docStyleVersion="5" docTitle="Protemnodon Owen 1874" docType="treatment" docVersion="4" lastPageNumber="191" masterDocId="FFDCFF85FFECD535FF887446FFD0FFEC" masterDocTitle="Systematics and palaeobiology of kangaroos of the late Cenozoic genus Protemnodon (Marsupialia, Macropodidae)" masterLastPageNumber="261" masterPageNumber="1" pageNumber="189" updateTime="1713463467402" updateUser="ExternalLinkService">
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<mods:title id="E8626F6C726C2C505408B0064D7F2687">Systematics and palaeobiology of kangaroos of the late Cenozoic genus Protemnodon (Marsupialia, Macropodidae)</mods:title>
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<mods:namePart id="2094867A6132B27D2BD6B43EA109AD0E">Kerr, Isaac A. R.</mods:namePart>
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<mods:affiliation id="41374CAFC91F892006F438699D23E964">College of Science and Engineering, Flinders University, Bedford Park, South Australia, Australia, 5042</mods:affiliation>
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<mods:roleTerm id="D6A7631B0DB22CB1592D4F59878BCCD6">Author</mods:roleTerm>
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<mods:namePart id="56349D4A63F1DCE358FCF83638F3D857">Camens, Aaron B.</mods:namePart>
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<mods:affiliation id="0381937F37D42B0222645A6B9EA06848">College of Science and Engineering, Flinders University, Bedford Park, South Australia, Australia, 5042</mods:affiliation>
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<mods:namePart id="343FB0C381BF0B3DD580E0DA616DCFE7">Van Zoelen, Jacob D.</mods:namePart>
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<mods:affiliation id="2A05595087D2F29922193BAB5A311750">College of Science and Engineering, Flinders University, Bedford Park, South Australia, Australia, 5042</mods:affiliation>
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<mods:namePart id="3BFEDA4C63E079C0C3D5A1C61757E7CD">Worthy, Trevor H.</mods:namePart>
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<mods:nameIdentifier id="19CEDA8369FFCAD40C9EE3867A82C49D" type="ORCID">0000-0001-7047-4680</mods:nameIdentifier>
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<mods:affiliation id="0DA38D5485F1D8D8A2913D4175EC840A">College of Science and Engineering, Flinders University, Bedford Park, South Australia, Australia, 5042</mods:affiliation>
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<mods:namePart id="1068E7D9B8246D2ADBE37C57F15CC2CF">Prideaux, Gavin J.</mods:namePart>
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<mods:affiliation id="68EBE1AF443FCB84DBABAA0D0C33EE6E">College of Science and Engineering, Flinders University, Bedford Park, South Australia, Australia, 5042</mods:affiliation>
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<mods:title id="4793FA1B502CF8FFB9017A78157EB897">Megataxa</mods:title>
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<mods:part id="C09DFFFB67DD7F4344815FD4927141A1">
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<mods:date id="1CF716A6488C3ED22CD8723AD949541F">2024</mods:date>
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<mods:number id="A4C7A360CAA4E80D0A0FDEA65ACC74C1">2024-04-15</mods:number>
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<mods:number id="EF5F8EA56F8C94DF4B7DE09DE7C66833">11</mods:number>
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<mods:number id="028196BF6AD87B11B5C2D2F9B3136511">1</mods:number>
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<mods:start id="AAB06459CDEBB0DCCCFA084CCDC9D248">1</mods:start>
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<mods:identifier id="5D2831B56ADBE37319F16E4E5D7EF271" type="DOI">10.11646/megataxa.11.1.1</mods:identifier>
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<mods:identifier id="524A92D466ACED6F368709697DF11873" type="ISSN">2703-3090</mods:identifier>
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<treatment id="03E587FDFF50D58BFCA2728CFA47FA09" ID-DOI="http://doi.org/10.5281/zenodo.10993702" ID-Zenodo-Dep="10993702" LSID="urn:lsid:plazi:treatment:03E587FDFF50D58BFCA2728CFA47FA09" httpUri="http://treatment.plazi.org/id/03E587FDFF50D58BFCA2728CFA47FA09" lastPageId="190" lastPageNumber="191" pageId="188" pageNumber="189">
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<subSubSection id="C3566560FF50D589FCA2728CFC1FF8E9" pageId="188" pageNumber="189" type="nomenclature">
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<paragraph id="8BF336EBFF50D589FCA2728CFC1FF8E9" blockId="188.[810,1451,1738,1797]" pageId="188" pageNumber="189">
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<emphasis id="B938EAF9FF50D589FCA2728CFC1FF8E9" bold="true" pageId="188" pageNumber="189">
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Status of other species previously referred to
|
||
<taxonomicName id="4C4C4D68FF50D589FCA272ACFC1FF8E9" authorityName="Owen" authorityYear="1874" box="[810,975,1770,1797]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="188" pageNumber="189" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF50D589FCA272ACFC1FF8E9" bold="true" box="[810,975,1770,1797]" italics="true" pageId="188" pageNumber="189">Protemnodon</emphasis>
|
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</taxonomicName>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3566560FF50D589FCA2736CFB47F8A9" box="[810,1175,1834,1861]" pageId="188" pageNumber="189" type="multiple">
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<paragraph id="8BF336EBFF50D589FCA2736CFB47F8A9" blockId="188.[810,1175,1834,1861]" box="[810,1175,1834,1861]" pageId="188" pageNumber="189">
|
||
<emphasis id="B938EAF9FF50D589FCA2736CFB47F8A9" bold="true" box="[810,1175,1834,1861]" italics="true" pageId="188" pageNumber="189">Australian Pleistocene species</emphasis>
|
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3566560FF50D588FCA27328FD3AF968" lastPageId="189" lastPageNumber="190" pageId="188" pageNumber="189" type="multiple">
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<paragraph id="8BF336EBFF50D589FCA27328FBFFF848" blockId="188.[810,1452,1902,2084]" pageId="188" pageNumber="189">
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<taxonomicName id="4C4C4D68FF50D589FCA27328FB54F869" authority="(Owen, 1874)" baseAuthorityName="Owen" baseAuthorityYear="1874" box="[810,1156,1902,1925]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="188" pageNumber="189" phylum="Chordata" rank="species" species="brehus">
|
||
<emphasis id="B938EAF9FF50D589FCA27328FC23F86F" box="[810,1011,1902,1923]" italics="true" pageId="188" pageNumber="189">Protemnodon brehus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDD4B1AFF50D589FB897329FBADF869" author="Owen, R." box="[1025,1149,1903,1925]" pageId="188" pageNumber="189" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen, 1874</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="B938EAF9FF50D589FB067328FB66F86F" box="[1166,1206,1902,1923]" italics="true" pageId="188" pageNumber="189">Phil</emphasis>
|
||
.
|
||
<emphasis id="B938EAF9FF50D589FB4B7329FA8BF868" box="[1219,1371,1903,1924]" italics="true" pageId="188" pageNumber="189">Trans. Roy. Soc</emphasis>
|
||
. 164, p. 272—
|
||
<emphasis id="B938EAF9FF50D589FC1373C9FBF9F84F" box="[923,1065,1934,1956]" italics="true" pageId="188" pageNumber="189">nomen dubium</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF50D589FCA273E8FBFFF808" blockId="188.[810,1452,1902,2084]" pageId="188" pageNumber="189">
|
||
<taxonomicName id="4C4C4D68FF50D589FCA273E8FBA9F828" authority="Owen, 1874" authorityName="Owen" authorityYear="1874" box="[810,1145,1966,1988]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="188" pageNumber="189" phylum="Chordata" rank="species" species="mimas">
|
||
<emphasis id="B938EAF9FF50D589FCA273E8FC24F828" box="[810,1012,1966,1988]" italics="true" pageId="188" pageNumber="189">Protemnodon mimas</emphasis>
|
||
<bibRefCitation id="EFDD4B1AFF50D589FC7473E9FBA9F828" author="Owen, R." box="[1020,1145,1967,1988]" pageId="188" pageNumber="189" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen, 1874</bibRefCitation>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="B938EAF9FF50D589FB0D73E8FA88F828" box="[1157,1368,1966,1988]" italics="true" pageId="188" pageNumber="189">Phil. Trans. Roy. Soc</emphasis>
|
||
. 164, p. 278—
|
||
<emphasis id="B938EAF9FF50D589FC137389FBF9F80F" box="[923,1065,1998,2020]" italics="true" pageId="188" pageNumber="189">nomen dubium</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF50D589FCA273A8FBFFF7C8" blockId="188.[810,1452,1902,2084]" pageId="188" pageNumber="189">
|
||
<taxonomicName id="4C4C4D68FF50D589FCA273A8FB51F7E8" authority="Owen, 1874" authorityName="Owen" authorityYear="1874" box="[810,1153,2030,2052]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="188" pageNumber="189" phylum="Chordata" rank="species" species="roechus">
|
||
<emphasis id="B938EAF9FF50D589FCA273A8FC2EF7EF" box="[810,1022,2030,2051]" italics="true" pageId="188" pageNumber="189">Protemnodon roechus</emphasis>
|
||
<bibRefCitation id="EFDD4B1AFF50D589FB8E73A9FB51F7E8" author="Owen, R." box="[1030,1153,2031,2052]" pageId="188" pageNumber="189" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen, 1874</bibRefCitation>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="B938EAF9FF50D589FB0373A8FAB1F7E8" box="[1163,1377,2030,2052]" italics="true" pageId="188" pageNumber="189">Phil. Trans. Roy. Soc.</emphasis>
|
||
164, p. 281—
|
||
<emphasis id="B938EAF9FF50D589FC137C49FBF9F7CF" box="[923,1065,2062,2084]" italics="true" pageId="188" pageNumber="189">nomen dubium</emphasis>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF336663FF51D588FF00710AFD95FA29" ID-DOI="http://doi.org/10.5281/zenodo.10994012" ID-Zenodo-Dep="10994012" httpUri="https://zenodo.org/record/10994012/files/figure.png" pageId="189" pageNumber="190" startId="189.[136,235,1356,1380]" targetBox="[162,1425,183,1327]" targetPageId="189" targetType="figure">
|
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<paragraph id="8BF336EBFF51D588FF00710AFD95FA29" blockId="189.[136,1452,1356,1477]" pageId="189" pageNumber="190">
|
||
<emphasis id="B938EAF9FF51D588FF00710AFECDFA89" bold="true" box="[136,285,1356,1381]" pageId="189" pageNumber="190">FIGURE 108.</emphasis>
|
||
<bibRefCitation id="EFDD4B1AFF51D588FEAD710BFE66FA89" author="Owen, R." box="[293,438,1357,1381]" pageId="189" pageNumber="190" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen’s (1874</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDD4B1AFF51D588FE49710BFE25FA89" author="Owen, R." box="[449,501,1357,1381]" pageId="189" pageNumber="190" refId="ref152945" refString="Owen, R. (1877) Researches on the fossil remains of the extinct mammals of Australia; with a notice of the extinct marsupials of England. 1. J. Erxleben, London, 522 pp. https: // doi. org / 10.5962 / bhl. title. 77375" type="book" year="1877">1877</bibRefCitation>
|
||
) type specimens for species of
|
||
<taxonomicName id="4C4C4D68FF51D588FCBE710BFC6FFA89" authorityName="Owen" authorityYear="1874" box="[822,959,1357,1381]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="189" pageNumber="190" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF51D588FCBE710BFC6FFA89" box="[822,959,1357,1381]" italics="true" pageId="189" pageNumber="190">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
: (a–d, f) all partial left dentaries in buccal view, the holotypes of (a)
|
||
<taxonomicName id="4C4C4D68FF51D588FEDD712BFD4DFA68" authority=", NHMUK PVM 2258" authorityName=", NHMUK PVM" authorityYear="2258" box="[341,669,1388,1412]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="189" pageNumber="190" phylum="Chordata" rank="species" species="antaeus">
|
||
<emphasis id="B938EAF9FF51D588FEDD712BFE6DFA68" box="[341,445,1389,1412]" italics="true" pageId="189" pageNumber="190">P. antaeus</emphasis>
|
||
, NHMUK PVM2258
|
||
</taxonomicName>
|
||
; (b)
|
||
<taxonomicName id="4C4C4D68FF51D588FD43712BFC25FA68" authority=", NHMUK PVM 1895" authorityName=", NHMUK PVM" authorityYear="1895" box="[715,1013,1388,1413]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="189" pageNumber="190" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF51D588FD43712BFCC5FA69" box="[715,789,1389,1413]" italics="true" pageId="189" pageNumber="190">P. anak</emphasis>
|
||
, NHMUK PVM1895
|
||
</taxonomicName>
|
||
; (c)
|
||
<taxonomicName id="4C4C4D68FF51D588FBAA712BFA93FA68" authority=", NHMUK PVOR" authorityName="NHMUK PVOR" box="[1058,1347,1388,1413]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="189" pageNumber="190" phylum="Chordata" rank="species" species="roechus">
|
||
<emphasis id="B938EAF9FF51D588FBAA712BFB5AFA69" box="[1058,1162,1389,1413]" italics="true" pageId="189" pageNumber="190">P. roechus</emphasis>
|
||
, NHMUK PVOR
|
||
</taxonomicName>
|
||
35968; (d)
|
||
<emphasis id="B938EAF9FF51D588FF0071CBFF6CFA48" box="[136,188,1421,1444]" italics="true" pageId="189" pageNumber="190">P. og</emphasis>
|
||
, NHMUK PVOR35963; (f)
|
||
<taxonomicName id="4C4C4D68FF51D588FE6D71CBFD2EFA48" authority=", NHMUK PVOR" authorityName="NHMUK PVOR" box="[485,766,1420,1444]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="189" pageNumber="190" phylum="Chordata" rank="species" species="mimas">
|
||
<emphasis id="B938EAF9FF51D588FE6D71CBFD91FA48" box="[485,577,1421,1444]" italics="true" pageId="189" pageNumber="190">P. mimas</emphasis>
|
||
, NHMUK PVOR
|
||
</taxonomicName>
|
||
43351; and (e) the lectotype of
|
||
<taxonomicName id="4C4C4D68FF51D588FBCA71CBFA8DFA49" authority=", NHMUK PVOR" authorityName="NHMUK PVOR" box="[1090,1373,1421,1445]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="189" pageNumber="190" phylum="Chordata" rank="species" species="brehus">
|
||
<emphasis id="B938EAF9FF51D588FBCA71CBFB71FA49" box="[1090,1185,1421,1445]" italics="true" pageId="189" pageNumber="190">P. brehus</emphasis>
|
||
, NHMUK PVOR
|
||
</taxonomicName>
|
||
43303a, partial maxillae and palate in occlusal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336EBFF51D588FF007268FD3AF968" blockId="189.[136,777,1582,1668]" pageId="189" pageNumber="190">
|
||
<taxonomicName id="4C4C4D68FF51D588FF007268FE03F9A8" authority="Owen, 1877" authorityName="Owen" authorityYear="1877" box="[136,467,1582,1604]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="189" pageNumber="190" phylum="Chordata" rank="species" species="antaeus">
|
||
<emphasis id="B938EAF9FF51D588FF007268FE89F9A8" box="[136,345,1582,1604]" italics="true" pageId="189" pageNumber="190">Protemnodon antaeus</emphasis>
|
||
<bibRefCitation id="EFDD4B1AFF51D588FED47269FE03F9A8" author="Owen, R." box="[348,467,1583,1604]" pageId="189" pageNumber="190" refId="ref152945" refString="Owen, R. (1877) Researches on the fossil remains of the extinct mammals of Australia; with a notice of the extinct marsupials of England. 1. J. Erxleben, London, 522 pp. https: // doi. org / 10.5962 / bhl. title. 77375" type="book" year="1877">Owen, 1877</bibRefCitation>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="B938EAF9FF51D588FE527268FE0EF96F" italics="true" pageId="189" pageNumber="190">
|
||
Researches on the fossil remains of the extinct mammals of
|
||
<collectingCountry id="F35B767BFF51D588FE5A7208FDFDF98F" box="[466,557,1614,1635]" name="Australia" pageId="189" pageNumber="190">Australia</collectingCountry>
|
||
; with a notice of the extinct marsupials of
|
||
<collectingCountry id="F35B767BFF51D588FE057228FE0EF96F" box="[397,478,1646,1667]" name="United Kingdom" pageId="189" pageNumber="190">England</collectingCountry>
|
||
</emphasis>
|
||
. 1, p. 448—
|
||
<emphasis id="B938EAF9FF51D588FDDE7229FD34F96F" box="[598,740,1646,1668]" italics="true" pageId="189" pageNumber="190">nomen dubium</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3566560FF51D58BFF0072EDFA47FA09" lastPageId="190" lastPageNumber="191" pageId="189" pageNumber="190" type="discussion">
|
||
<paragraph id="8BF336EBFF51D588FF0072EDFB66F949" blockId="189.[136,778,1707,2086]" lastBlockId="189.[810,1452,1579,2085]" pageId="189" pageNumber="190">
|
||
This section discusses the taxonomic history and status of the remaining species of
|
||
<taxonomicName id="4C4C4D68FF51D588FE11728DFDE0F909" authorityName="Owen" authorityYear="1874" box="[409,560,1739,1765]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="189" pageNumber="190" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF51D588FE11728DFDE0F909" box="[409,560,1739,1765]" italics="true" pageId="189" pageNumber="190">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
described by
|
||
<bibRefCitation id="EFDD4B1AFF51D588FD4F728DFF1CF8E9" author="Owen, R." pageId="189" pageNumber="190" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen (1874</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDD4B1AFF51D588FF5372ADFEC7F8E9" author="Owen, R." box="[219,279,1771,1797]" pageId="189" pageNumber="190" refId="ref152945" refString="Owen, R. (1877) Researches on the fossil remains of the extinct mammals of Australia; with a notice of the extinct marsupials of England. 1. J. Erxleben, London, 522 pp. https: // doi. org / 10.5962 / bhl. title. 77375" type="book" year="1877">1877</bibRefCitation>
|
||
), including justification for the designation of
|
||
<taxonomicName id="4C4C4D68FF51D588FF20734DFE71F8C9" box="[168,417,1803,1829]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="189" pageNumber="190" phylum="Chordata" rank="species" species="roechus">
|
||
<emphasis id="B938EAF9FF51D588FF20734DFE71F8C9" box="[168,417,1803,1829]" italics="true" pageId="189" pageNumber="190">Protemnodon roechus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF51D588FE53734AFD93F8C9" box="[475,579,1803,1829]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="189" pageNumber="190" phylum="Chordata" rank="species" species="brehus">
|
||
<emphasis id="B938EAF9FF51D588FE53734AFD93F8C9" box="[475,579,1803,1829]" italics="true" pageId="189" pageNumber="190">P. brehus</emphasis>
|
||
</taxonomicName>
|
||
as
|
||
<emphasis id="B938EAF9FF51D588FDE5734AFCDAF8C9" box="[621,778,1803,1829]" italics="true" pageId="189" pageNumber="190">nomina dubia</emphasis>
|
||
and for the erection of two
|
||
<emphasis id="B938EAF9FF51D588FE66736AFD42F8A9" box="[494,658,1836,1861]" italics="true" pageId="189" pageNumber="190">species novae</emphasis>
|
||
from the Australian Pleistocene. For reference, all of the
|
||
<typeStatus id="54F78849FF51D588FD15730DFCDAF889" box="[669,778,1867,1893]" pageId="189" pageNumber="190" type="holotype">holotypes</typeStatus>
|
||
and
|
||
<typeStatus id="54F78849FF51D588FF34732DFEFEF869" box="[188,302,1899,1925]" pageId="189" pageNumber="190" type="lectotype">lectotypes</typeStatus>
|
||
for
|
||
<bibRefCitation id="EFDD4B1AFF51D588FEEE732DFDDCF869" author="Owen, R." box="[358,524,1899,1925]" pageId="189" pageNumber="190" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen’s (1874</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDD4B1AFF51D588FD94732DFD85F869" author="Owen, R." box="[540,597,1899,1925]" pageId="189" pageNumber="190" refId="ref152945" refString="Owen, R. (1877) Researches on the fossil remains of the extinct mammals of Australia; with a notice of the extinct marsupials of England. 1. J. Erxleben, London, 522 pp. https: // doi. org / 10.5962 / bhl. title. 77375" type="book" year="1877">1877</bibRefCitation>
|
||
) six species of
|
||
<taxonomicName id="4C4C4D68FF51D588FF0073CDFECFF849" authorityName="Owen" authorityYear="1874" box="[136,287,1931,1957]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="189" pageNumber="190" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF51D588FF0073CDFECFF849" box="[136,287,1931,1957]" italics="true" pageId="189" pageNumber="190">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
are figured here (
|
||
<figureCitation id="13772A6EFF51D588FE7F73CDFD8EF849" box="[503,606,1931,1957]" captionStart="FIGURE 108" captionStartId="189.[136,235,1356,1380]" captionTargetBox="[162,1425,183,1327]" captionTargetId="figure-18@189.[151,1436,172,1334]" captionTargetPageId="189" captionText="FIGURE 108. Owen’s (1874, 1877) type specimens for species of Protemnodon: (a–d, f) all partial left dentaries in buccal view, the holotypes of (a) P. antaeus, NHMUK PVM2258; (b) P. anak, NHMUK PVM1895; (c) P. roechus, NHMUK PVOR35968; (d) P. og, NHMUK PVOR35963; (f) P. mimas, NHMUK PVOR43351; and (e) the lectotype of P. brehus, NHMUK PVOR43303a, partial maxillae and palate in occlusal view." figureDoi="http://doi.org/10.5281/zenodo.10994012" httpUri="https://zenodo.org/record/10994012/files/figure.png" pageId="189" pageNumber="190">Fig. 108</figureCitation>
|
||
). Note that, as per ICZN Article 61, the
|
||
<typeStatus id="54F78849FF51D588FE4E73EDFDF8F829" box="[454,552,1963,1989]" pageId="189" pageNumber="190" type="holotype">holotype</typeStatus>
|
||
specimen (
|
||
<emphasis id="B938EAF9FF51D588FD3F73EAFD0AF829" box="[695,730,1964,1989]" italics="true" pageId="189" pageNumber="190">i.e.</emphasis>
|
||
the name-bearing type) is the sole specimen that provides the diagnostic features of the
|
||
<emphasis id="B938EAF9FF51D588FE2773AAFE2AF7E9" box="[431,506,2028,2053]" italics="true" pageId="189" pageNumber="190">nomen</emphasis>
|
||
—the type (not the type series,
|
||
<emphasis id="B938EAF9FF51D588FF5D7C4AFF27F7C9" box="[213,247,2060,2085]" italics="true" pageId="189" pageNumber="190">i.e.</emphasis>
|
||
<typeStatus id="54F78849FF51D588FF747C4DFEBAF7C9" box="[252,362,2059,2085]" pageId="189" pageNumber="190" type="paratype">paratypes</typeStatus>
|
||
, nor referred specimens) provides the objective standard of reference for the application of the name it bears (ICZN 1999).A loss of the utility of a
|
||
<emphasis id="B938EAF9FF51D588FAE9720AFA7CF989" box="[1377,1452,1612,1637]" italics="true" pageId="189" pageNumber="190">nomen</emphasis>
|
||
is the criterion for status as
|
||
<emphasis id="B938EAF9FF51D588FBE0722AFADFF969" box="[1128,1295,1643,1669]" italics="true" pageId="189" pageNumber="190">nomen dubium</emphasis>
|
||
, although the
|
||
<emphasis id="B938EAF9FF51D588FCA272CAFCA5F949" box="[810,885,1676,1701]" italics="true" pageId="189" pageNumber="190">nomen</emphasis>
|
||
remains valid and available.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF51D58BFCD272EDFF0FFE89" blockId="189.[810,1452,1579,2085]" lastBlockId="190.[136,780,171,2085]" lastPageId="190" lastPageNumber="191" pageId="189" pageNumber="190">
|
||
The following is an extract from the redescription of
|
||
<taxonomicName id="4C4C4D68FF51D588FCC0728AFC6CF909" box="[840,956,1739,1765]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="189" pageNumber="190" phylum="Chordata" rank="species" species="roechus">
|
||
<emphasis id="B938EAF9FF51D588FCC0728AFC6CF909" box="[840,956,1739,1765]" italics="true" pageId="189" pageNumber="190">P. roechus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<bibRefCitation id="EFDD4B1AFF51D588FB8A728DFB35F909" author="Bartholomai, A." box="[1026,1253,1739,1765]" pageId="189" pageNumber="190" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai (1973)</bibRefCitation>
|
||
, where the dental morphology of the species was contrasted with that of
|
||
<taxonomicName id="4C4C4D68FF51D588FCC7734AFC78F8C9" authorityName=", NHMUK PVM" authorityYear="1895" box="[847,936,1803,1829]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="189" pageNumber="190" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF51D588FCC7734AFC78F8C9" box="[847,936,1803,1829]" italics="true" pageId="189" pageNumber="190">P. anak</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF51D588FC64734AFB8AF8C9" box="[1004,1114,1803,1829]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="189" pageNumber="190" phylum="Chordata" rank="species" species="brehus">
|
||
<emphasis id="B938EAF9FF51D588FC64734AFB8AF8C9" box="[1004,1114,1803,1829]" italics="true" pageId="189" pageNumber="190">P. brehus</emphasis>
|
||
</taxonomicName>
|
||
: ‘[
|
||
<taxonomicName id="4C4C4D68FF51D588FB0B734DFACCF8C9" authorityName="Owen" authorityYear="1874" box="[1155,1308,1803,1829]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="189" pageNumber="190" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF51D588FB0B734DFACCF8C9" box="[1155,1308,1803,1829]" italics="true" pageId="189" pageNumber="190">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
]
|
||
<taxonomicName id="4C4C4D68FF51D588FABA734DFA5AF8C9" box="[1330,1418,1803,1829]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="189" pageNumber="190" phylum="Chordata" rank="species" species="roechus">
|
||
<emphasis id="B938EAF9FF51D588FABA734DFA5AF8C9" box="[1330,1418,1803,1829]" italics="true" pageId="189" pageNumber="190">roechus</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from other known species of
|
||
<taxonomicName id="4C4C4D68FF51D588FA9A736DFA7BF8A9" authorityName="Owen" authorityYear="1874" box="[1298,1451,1835,1861]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="189" pageNumber="190" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF51D588FA9A736DFA7BF8A9" box="[1298,1451,1835,1861]" italics="true" pageId="189" pageNumber="190">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
by both its extreme size and generally distinct morphology. In particular, the relatively less ornamented nature of its permanent lower premolar, the broadly rounded, swollen condition of the posterior crown bases of the lower molars, the non-vertical transecting ridges of the permanent upper premolars, the crescentic shape of this tooth in occlusal view and the generally marked tuberculation of the lingual extremity of the median valley in the upper molars are all useful in the morphological separation of the species. Overlap in size does occur with
|
||
<taxonomicName id="4C4C4D68FF52D58BFD1074AAFCD3FEE9" box="[664,771,235,261]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="brehus">
|
||
<emphasis id="B938EAF9FF52D58BFD1074AAFCD3FEE9" box="[664,771,235,261]" italics="true" pageId="190" pageNumber="191">P. brehus</emphasis>
|
||
</taxonomicName>
|
||
, but by applying a combination of the characters available separation of the species is achieved’ (
|
||
<bibRefCitation id="EFDD4B1AFF52D58BFDBC756DFCD5FEA9" author="Bartholomai, A." box="[564,773,299,325]" pageId="190" pageNumber="191" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai 1973</bibRefCitation>
|
||
, p. 346).
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF52D58BFF30752DFD88FC29" blockId="190.[136,780,171,2085]" pageId="190" pageNumber="191">
|
||
Nevertheless, the distinction as made by
|
||
<bibRefCitation id="EFDD4B1AFF52D58BFDF6752DFF02FE49" author="Bartholomai, A." pageId="190" pageNumber="191" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai (1973)</bibRefCitation>
|
||
is in fact not so clear. For a variety of reasons,
|
||
<taxonomicName id="4C4C4D68FF52D58BFD7D75CAFF03FE29" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="brehus">
|
||
<emphasis id="B938EAF9FF52D58BFD7D75CAFF03FE29" italics="true" pageId="190" pageNumber="191">P. brehus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF52D58BFE8375EAFEAFFE29" box="[267,383,427,453]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="roechus">
|
||
<emphasis id="B938EAF9FF52D58BFE8375EAFEAFFE29" box="[267,383,427,453]" italics="true" pageId="190" pageNumber="191">P. roechus</emphasis>
|
||
</taxonomicName>
|
||
have never been clearly delimited. The
|
||
<typeStatus id="54F78849FF52D58BFF44758DFEE3FE09" box="[204,307,459,485]" pageId="190" pageNumber="191" type="lectotype">lectotype</typeStatus>
|
||
specimen for
|
||
<taxonomicName id="4C4C4D68FF52D58BFD80758AFF05FDE9" authority=", NHMUK PVOR" authorityName="NHMUK PVOR" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="brehus">
|
||
<emphasis id="B938EAF9FF52D58BFD80758AFD50FE09" box="[520,640,459,485]" italics="true" pageId="190" pageNumber="191">P. brehus</emphasis>
|
||
, NHMUK PVOR
|
||
</taxonomicName>
|
||
43303a, is a partial palate containing left M3–M4 and right M2–M4. However, the
|
||
<typeStatus id="54F78849FF52D58BFDA5764DFD5FFDC9" box="[557,655,523,549]" pageId="190" pageNumber="191" type="holotype">holotype</typeStatus>
|
||
specimen for
|
||
<taxonomicName id="4C4C4D68FF52D58BFF3B766AFE2DFDAA" authority=", NHMUK PVOR" authorityName="NHMUK PVOR" box="[179,509,555,582]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="roechus">
|
||
<emphasis id="B938EAF9FF52D58BFF3B766AFEFAFDA9" box="[179,298,555,581]" italics="true" pageId="190" pageNumber="191">P. roechus</emphasis>
|
||
, NHMUK PVOR
|
||
</taxonomicName>
|
||
35968, is a left dentary fragment containing heavily worn p3–m3, and as such the
|
||
<typeStatus id="54F78849FF52D58BFF00762DFF25FD69" box="[136,245,619,645]" pageId="190" pageNumber="191" type="holotype">holotypes</typeStatus>
|
||
have no shared features for direct comparison. Three of the five characteristics stated by
|
||
<bibRefCitation id="EFDD4B1AFF52D58BFDF676CDFF02FD29" author="Bartholomai, A." pageId="190" pageNumber="191" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai (1973)</bibRefCitation>
|
||
as distinguishing
|
||
<taxonomicName id="4C4C4D68FF52D58BFE2576EAFDF5FD29" box="[429,549,683,709]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="roechus">
|
||
<emphasis id="B938EAF9FF52D58BFE2576EAFDF5FD29" box="[429,549,683,709]" italics="true" pageId="190" pageNumber="191">P. roechus</emphasis>
|
||
</taxonomicName>
|
||
from other species of
|
||
<taxonomicName id="4C4C4D68FF52D58BFF26768DFE95FD09" authorityName="Owen" authorityYear="1874" box="[174,325,715,741]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF52D58BFF26768DFE95FD09" box="[174,325,715,741]" italics="true" pageId="190" pageNumber="191">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
are in the upper cheek teeth, which are not preserved in the
|
||
<typeStatus id="54F78849FF52D58BFE2B76ADFDD5FCE9" box="[419,517,747,773]" pageId="190" pageNumber="191" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4C4D68FF52D58BFDA776AAFD75FCE9" box="[559,677,747,773]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="roechus">
|
||
<emphasis id="B938EAF9FF52D58BFDA776AAFD75FCE9" box="[559,677,747,773]" italics="true" pageId="190" pageNumber="191">P. roechus</emphasis>
|
||
</taxonomicName>
|
||
, and the
|
||
<typeStatus id="54F78849FF52D58BFF00774DFF3AFCC9" box="[136,234,779,805]" pageId="190" pageNumber="191" type="holotype">holotype</typeStatus>
|
||
of neither species preserves a P3. The results of our analyses and the morphological comparisons of the material of
|
||
<taxonomicName id="4C4C4D68FF52D58BFE87770DFE76FC89" authorityName="Owen" authorityYear="1874" box="[271,422,843,869]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF52D58BFE87770DFE76FC89" box="[271,422,843,869]" italics="true" pageId="190" pageNumber="191">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
from across
|
||
<collectingCountry id="F35B767BFF52D58BFDC8770DFD77FC89" box="[576,679,843,869]" name="Australia" pageId="190" pageNumber="191">Australia</collectingCountry>
|
||
indicate that the characteristics described above do not distinguish two distinct morphotypes, but instead describe features that vary continuously within the sample.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF52D58BFF30778DFD04F8E9" blockId="190.[136,780,171,2085]" pageId="190" pageNumber="191">
|
||
The statement by
|
||
<bibRefCitation id="EFDD4B1AFF52D58BFE19778DFDBDFC09" author="Bartholomai, A." box="[401,621,971,997]" pageId="190" pageNumber="191" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai (1973</bibRefCitation>
|
||
, p. 346) that specimens of
|
||
<taxonomicName id="4C4C4D68FF52D58BFEA377AAFE73FBE9" box="[299,419,1003,1029]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="roechus">
|
||
<emphasis id="B938EAF9FF52D58BFEA377AAFE73FBE9" box="[299,419,1003,1029]" italics="true" pageId="190" pageNumber="191">P. roechus</emphasis>
|
||
</taxonomicName>
|
||
are characterised by “extreme size” that differentiates them from those of
|
||
<taxonomicName id="4C4C4D68FF52D58BFD0C704AFD3DFBC9" box="[644,749,1035,1061]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="brehus">
|
||
<emphasis id="B938EAF9FF52D58BFD0C704AFD3DFBC9" box="[644,749,1035,1061]" italics="true" pageId="190" pageNumber="191">P. brehus</emphasis>
|
||
</taxonomicName>
|
||
is not well-supported by the data presented in that paper (
|
||
<bibRefCitation id="EFDD4B1AFF52D58BFF07700DFEB1FB89" author="Bartholomai, A." box="[143,353,1099,1125]" pageId="190" pageNumber="191" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai 1973</bibRefCitation>
|
||
, fig. 9), which show almost identical mean dimensions for the lower dentition, nor by the data presented here (See
|
||
<figureCitation id="13772A6EFF52D58BFEEE70CDFDD2FB4A" box="[358,514,1163,1190]" captionStart-0="FIGURE 116" captionStart-1="FIGURE 117" captionStart-2="FIGURE 118" captionStart-3="FIGURE 119" captionStart-4="FIGURE 120" captionStart-5="FIGURE 121" captionStart-6="FIGURE 122" captionStartId-0="198.[136,235,1068,1092]" captionStartId-1="198.[136,235,2028,2052]" captionStartId-2="199.[136,235,1068,1092]" captionStartId-3="199.[136,235,2028,2052]" captionStartId-4="200.[136,235,1068,1092]" captionStartId-5="200.[136,235,2028,2052]" captionStartId-6="201.[136,235,1068,1092]" captionTargetBox-0="[158,1429,201,1042]" captionTargetBox-1="[158,1429,1156,2002]" captionTargetBox-2="[158,1429,201,1042]" captionTargetBox-3="[158,1429,1161,2002]" captionTargetBox-4="[158,1429,201,1042]" captionTargetBox-5="[158,1429,1160,2002]" captionTargetBox-6="[158,1429,193,1041]" captionTargetId-0="figure-27@198.[151,1436,194,1049]" captionTargetId-1="figure-65@198.[151,1436,1149,2009]" captionTargetId-2="figure-18@199.[151,1436,194,1049]" captionTargetId-3="figure-56@199.[151,1436,1154,2009]" captionTargetId-4="figure-29@200.[151,1436,194,1049]" captionTargetId-5="figure-67@200.[151,1436,1153,2009]" captionTargetId-6="figure-19@201.[151,1436,186,1048]" captionTargetPageId-0="198" captionTargetPageId-1="198" captionTargetPageId-2="199" captionTargetPageId-3="199" captionTargetPageId-4="200" captionTargetPageId-5="200" captionTargetPageId-6="201" captionText-0="FIGURE 116. scatterplot of P3 anterior width versus length in specimens of P. mamkurra sp. nov., P. viator sp. nov., and specimens allocated to P. mamkurra sp. nov. or viator sp. nov." captionText-1="FIGURE 117. scatterplot of P3 posterior width versus length in specimens of P. mamkurra sp. nov., P. viator sp. nov., and specimens allocated to P. mamkurra sp. nov. or viator sp. nov." captionText-2="FIGURE 118. scatterplot of M1 anterior width versus length in specimens of P. mamkurra sp. nov., P. viator sp. nov., and specimens allocated to P. mamkurra sp. nov. or viator sp. nov." captionText-3="FIGURE 119. scatterplot of M2 anterior width versus length in specimens of P. mamkurra sp. nov., P. viator sp. nov., and specimens allocated to P. mamkurra sp. nov. or viator sp. nov." captionText-4="FIGURE 120. scatterplot of M3 anterior width versus length in specimens of P. mamkurra sp. nov., P. viator sp. nov., and specimens allocated to P. mamkurra sp. nov. or viator sp. nov." captionText-5="FIGURE 121. boxplot of p3 length divided by m3 length in specimens of P. anak, P. mamkurra sp. nov., P. viator sp. nov., P. dawsonae sp. nov., P. snewini, P. otibandus, P. tumbuna, and four extrageneric taxa." captionText-6="FIGURE 122. scatterplot of m2 length versus anterior width in specimens of P. anak, P. mamkurra sp. nov., P. viator sp. nov., P. dawsonae sp. nov., P. snewini, P. otibandus, P. tumbuna, and five extrageneric taxa." figureDoi-0="http://doi.org/10.5281/zenodo.10994028" figureDoi-1="http://doi.org/10.5281/zenodo.10994030" figureDoi-2="http://doi.org/10.5281/zenodo.10994032" figureDoi-3="http://doi.org/10.5281/zenodo.10994034" figureDoi-4="http://doi.org/10.5281/zenodo.10994036" figureDoi-5="http://doi.org/10.5281/zenodo.10994038" figureDoi-6="http://doi.org/10.5281/zenodo.10994040" httpUri-0="https://zenodo.org/record/10994028/files/figure.png" httpUri-1="https://zenodo.org/record/10994030/files/figure.png" httpUri-2="https://zenodo.org/record/10994032/files/figure.png" httpUri-3="https://zenodo.org/record/10994034/files/figure.png" httpUri-4="https://zenodo.org/record/10994036/files/figure.png" httpUri-5="https://zenodo.org/record/10994038/files/figure.png" httpUri-6="https://zenodo.org/record/10994040/files/figure.png" pageId="190" pageNumber="191">Figs 116–122</figureCitation>
|
||
). The ‘crescentic shape’ of the P3 varies greatly within the sample of Pleistocene P3s not attributable to
|
||
<taxonomicName id="4C4C4D68FF52D58BFE0D708AFE06FB09" authorityName=", NHMUK PVM" authorityYear="1895" box="[389,470,1227,1253]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF52D58BFE0D708AFE06FB09" box="[389,470,1227,1253]" italics="true" pageId="190" pageNumber="191">P. anak</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13772A6EFF52D58BFE6E708DFD98FB09" box="[486,584,1227,1253]" captionStart="FIGURE 109" captionStartId="191.[136,235,1932,1956]" captionTargetBox="[330,1258,177,1902]" captionTargetId="figure-17@191.[321,1266,172,1910]" captionTargetPageId="191" captionText="FIGURE 109. occlusal view of 16 unworn to lightly worn right (a) upper and (b) lower third premolars of SAMA P24098 (single specimen number for multiple individuals) of Protemnodon mamkurra sp. nov. or P. viator sp. nov. from Goulden’s Hole, southeastern South Australia, showing the morphological and size variation visible in permanent premolars within a sample of one or possibly two species from a single locality." figureDoi="http://doi.org/10.5281/zenodo.10994014" httpUri="https://zenodo.org/record/10994014/files/figure.png" pageId="190" pageNumber="191">Fig. 109</figureCitation>
|
||
). The description of the ‘relatively less-ornamented nature of its [p3]’ most probably refers to the narrow, roughly dorsoventrally orientated ridgelets on the lingual and buccal faces of the p3. These ridgelets, when numerous and thin but distinct, are diagnostic of
|
||
<taxonomicName id="4C4C4D68FF52D58BFEC4712AFE4EFA69" authorityName=", NHMUK PVM" authorityYear="1895" box="[332,414,1387,1413]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF52D58BFEC4712AFE4EFA69" box="[332,414,1387,1413]" italics="true" pageId="190" pageNumber="191">P. anak</emphasis>
|
||
</taxonomicName>
|
||
, and are less numerous and less distinct in other Pleistocene Australian material of species of
|
||
<taxonomicName id="4C4C4D68FF52D58BFF2371EDFE92FA29" authorityName="Owen" authorityYear="1874" box="[171,322,1451,1477]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF52D58BFF2371EDFE92FA29" box="[171,322,1451,1477]" italics="true" pageId="190" pageNumber="191">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
. No discrete difference in the number or size of these ridgelets is discernible in the sample of Pleistocene Australian material when excluding that of
|
||
<taxonomicName id="4C4C4D68FF52D58BFD7D71AAFF6EF9C9" authorityName=", NHMUK PVM" authorityYear="1895" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF52D58BFD7D71AAFF6EF9C9" italics="true" pageId="190" pageNumber="191">P. anak</emphasis>
|
||
</taxonomicName>
|
||
. These ridgelets disappear almost completely with a moderate degree of wear, and so in older and thus often skeletally larger individuals with a greater degree of dental wear, the premolar appears increasingly less ornamented. It is possible that this biased
|
||
<bibRefCitation id="EFDD4B1AFF52D58BFD8672CDFCD9F949" author="Bartholomai, A." box="[526,777,1675,1701]" pageId="190" pageNumber="191" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai’s (1973)</bibRefCitation>
|
||
morphological analysis by causing larger individuals,
|
||
<emphasis id="B938EAF9FF52D58BFD6F72EAFCDAF929" box="[743,778,1708,1733]" italics="true" pageId="190" pageNumber="191">i.e.</emphasis>
|
||
those more likely to be assigned to
|
||
<taxonomicName id="4C4C4D68FF52D58BFD89728AFDA2F909" box="[513,626,1739,1765]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="roechus">
|
||
<emphasis id="B938EAF9FF52D58BFD89728AFDA2F909" box="[513,626,1739,1765]" italics="true" pageId="190" pageNumber="191">P. roechus</emphasis>
|
||
</taxonomicName>
|
||
, to seemingly possess p3s with fewer, less raised vertical ridgelets.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF52D58BFF30734DFE21F7EA" blockId="190.[136,780,171,2085]" pageId="190" pageNumber="191">
|
||
The condition of rounded, swollen posterior lophid bases is present in various lower molar specimens of
|
||
<taxonomicName id="4C4C4D68FF52D58BFF00730DFECFF889" authorityName="Owen" authorityYear="1874" box="[136,287,1867,1893]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF52D58BFF00730DFECFF889" box="[136,287,1867,1893]" italics="true" pageId="190" pageNumber="191">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
from the Pleistocene, including individuals of
|
||
<taxonomicName id="4C4C4D68FF52D58BFF21732AFF2DF869" authorityName=", NHMUK PVM" authorityYear="1895" box="[169,253,1899,1925]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF52D58BFF21732AFF2DF869" box="[169,253,1899,1925]" italics="true" pageId="190" pageNumber="191">P. anak</emphasis>
|
||
</taxonomicName>
|
||
, rather than solely individuals allocated to
|
||
<taxonomicName id="4C4C4D68FF52D58BFD7D732AFF30F849" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="roechus">
|
||
<emphasis id="B938EAF9FF52D58BFD7D732AFF30F849" italics="true" pageId="190" pageNumber="191">P. roechus</emphasis>
|
||
</taxonomicName>
|
||
. This condition is seen in molars with residual postcingulids and very occasionally in large molars with fully developed postcingulids (as in the
|
||
<typeStatus id="54F78849FF52D58BFDEC738DFD16F809" box="[612,710,1995,2021]" pageId="190" pageNumber="191" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4C4D68FF52D58BFD7D738AFE4CF7EA" authority=", NHMUKPV OR" authorityName="NHMUKPV OR" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="mimas">
|
||
<emphasis id="B938EAF9FF52D58BFD7D738AFF01F7E9" italics="true" pageId="190" pageNumber="191">P. mimas</emphasis>
|
||
, NHMUKPV OR
|
||
</taxonomicName>
|
||
43351).
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF52D58BFF307C4DFB24FCE9" blockId="190.[136,780,171,2085]" lastBlockId="190.[810,1452,171,1509]" pageId="190" pageNumber="191">
|
||
Our extensive survey of fossils of species of
|
||
<taxonomicName id="4C4C4D68FF52D58BFCA274EDFC11FF29" authorityName="Owen" authorityYear="1874" box="[810,961,171,197]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF52D58BFCA274EDFC11FF29" box="[810,961,171,197]" italics="true" pageId="190" pageNumber="191">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
from the Australian Pleistocene has revealed that there are indeed two Australian Pleistocene morphotypes of
|
||
<taxonomicName id="4C4C4D68FF52D58BFC7974ADFB58FEE9" authorityName="Owen" authorityYear="1874" box="[1009,1160,235,261]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF52D58BFC7974ADFB58FEE9" box="[1009,1160,235,261]" italics="true" pageId="190" pageNumber="191">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
aside from
|
||
<taxonomicName id="4C4C4D68FF52D58BFAAD74AAFAAEFEE9" authorityName=", NHMUK PVM" authorityYear="1895" box="[1317,1406,235,261]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF52D58BFAAD74AAFAAEFEE9" box="[1317,1406,235,261]" italics="true" pageId="190" pageNumber="191">P. anak</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C4C4D68FF52D58BFA1F74AAFC4FFEC9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF52D58BFA1F74AAFC4FFEC9" italics="true" pageId="190" pageNumber="191">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF52D58BFC2E754DFC2FFEC9" bold="true" box="[934,1023,267,293]" pageId="190" pageNumber="191">
|
||
<taxonomicNameLabel id="A20B5782FF52D58BFC2E754DFC2FFEC9" box="[934,1023,267,293]" pageId="190" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF52D58BFBBE754AFB45FEC9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1078,1173,268,293]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF52D58BFBBE754AFB45FEC9" box="[1078,1173,268,293]" italics="true" pageId="190" pageNumber="191">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF52D58BFB14754DFB25FEC9" bold="true" box="[1180,1269,267,293]" pageId="190" pageNumber="191">
|
||
<taxonomicNameLabel id="A20B5782FF52D58BFB14754DFB25FEC9" box="[1180,1269,267,293]" pageId="190" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
); however, they are primarily distinguishable on postcranial features. Neither can be delimited based on either the upper or lower cheek dentition and they have similar size ranges. As illustrated in
|
||
<figureCitation id="13772A6EFF52D58BFC7775CDFB73FE49" box="[1023,1187,395,421]" captionStart-0="FIGURE 116" captionStart-1="FIGURE 117" captionStart-2="FIGURE 118" captionStart-3="FIGURE 119" captionStart-4="FIGURE 120" captionStartId-0="198.[136,235,1068,1092]" captionStartId-1="198.[136,235,2028,2052]" captionStartId-2="199.[136,235,1068,1092]" captionStartId-3="199.[136,235,2028,2052]" captionStartId-4="200.[136,235,1068,1092]" captionTargetBox-0="[158,1429,201,1042]" captionTargetBox-1="[158,1429,1156,2002]" captionTargetBox-2="[158,1429,201,1042]" captionTargetBox-3="[158,1429,1161,2002]" captionTargetBox-4="[158,1429,201,1042]" captionTargetId-0="figure-27@198.[151,1436,194,1049]" captionTargetId-1="figure-65@198.[151,1436,1149,2009]" captionTargetId-2="figure-18@199.[151,1436,194,1049]" captionTargetId-3="figure-56@199.[151,1436,1154,2009]" captionTargetId-4="figure-29@200.[151,1436,194,1049]" captionTargetPageId-0="198" captionTargetPageId-1="198" captionTargetPageId-2="199" captionTargetPageId-3="199" captionTargetPageId-4="200" captionText-0="FIGURE 116. scatterplot of P3 anterior width versus length in specimens of P. mamkurra sp. nov., P. viator sp. nov., and specimens allocated to P. mamkurra sp. nov. or viator sp. nov." captionText-1="FIGURE 117. scatterplot of P3 posterior width versus length in specimens of P. mamkurra sp. nov., P. viator sp. nov., and specimens allocated to P. mamkurra sp. nov. or viator sp. nov." captionText-2="FIGURE 118. scatterplot of M1 anterior width versus length in specimens of P. mamkurra sp. nov., P. viator sp. nov., and specimens allocated to P. mamkurra sp. nov. or viator sp. nov." captionText-3="FIGURE 119. scatterplot of M2 anterior width versus length in specimens of P. mamkurra sp. nov., P. viator sp. nov., and specimens allocated to P. mamkurra sp. nov. or viator sp. nov." captionText-4="FIGURE 120. scatterplot of M3 anterior width versus length in specimens of P. mamkurra sp. nov., P. viator sp. nov., and specimens allocated to P. mamkurra sp. nov. or viator sp. nov." figureDoi-0="http://doi.org/10.5281/zenodo.10994028" figureDoi-1="http://doi.org/10.5281/zenodo.10994030" figureDoi-2="http://doi.org/10.5281/zenodo.10994032" figureDoi-3="http://doi.org/10.5281/zenodo.10994034" figureDoi-4="http://doi.org/10.5281/zenodo.10994036" httpUri-0="https://zenodo.org/record/10994028/files/figure.png" httpUri-1="https://zenodo.org/record/10994030/files/figure.png" httpUri-2="https://zenodo.org/record/10994032/files/figure.png" httpUri-3="https://zenodo.org/record/10994034/files/figure.png" httpUri-4="https://zenodo.org/record/10994036/files/figure.png" pageId="190" pageNumber="191">Figs 116–120</figureCitation>
|
||
, any divide in terms of dental dimensions between the
|
||
<typeStatus id="54F78849FF52D58BFB4275EDFB2AFE29" box="[1226,1274,427,453]" pageId="190" pageNumber="191">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="4C4C4D68FF52D58BFA1F75EAFC4FFE09" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF52D58BFA1F75EAFC4FFE09" italics="true" pageId="190" pageNumber="191">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF52D58BFC23758DFBDAFE09" bold="true" box="[939,1034,459,485]" pageId="190" pageNumber="191">
|
||
<taxonomicNameLabel id="A20B5782FF52D58BFC23758DFBDAFE09" box="[939,1034,459,485]" pageId="190" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF52D58BFBC4758AFB60FE09" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1100,1200,460,485]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF52D58BFBC4758AFB60FE09" box="[1100,1200,460,485]" italics="true" pageId="190" pageNumber="191">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF52D58BFB35758DFACCFE09" bold="true" box="[1213,1308,459,485]" pageId="190" pageNumber="191">
|
||
<taxonomicNameLabel id="A20B5782FF52D58BFB35758DFACCFE09" box="[1213,1308,459,485]" pageId="190" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is filled by isolated dental or craniodental specimens allocated to
|
||
<taxonomicName id="4C4C4D68FF52D58BFA1F75AAFC4FFDC9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF52D58BFA1F75AAFC4FFDC9" italics="true" pageId="190" pageNumber="191">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF52D58BFC2D764DFC2EFDC9" bold="true" box="[933,1022,523,549]" pageId="190" pageNumber="191">
|
||
<taxonomicNameLabel id="A20B5782FF52D58BFC2D764DFC2EFDC9" box="[933,1022,523,549]" pageId="190" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
or
|
||
<taxonomicName id="4C4C4D68FF52D58BFBAA764AFBB5FDC9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1058,1125,524,549]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF52D58BFBAA764AFBB5FDC9" box="[1058,1125,524,549]" italics="true" pageId="190" pageNumber="191">viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF52D58BFBE4764DFB15FDC9" bold="true" box="[1132,1221,523,549]" pageId="190" pageNumber="191">
|
||
<taxonomicNameLabel id="A20B5782FF52D58BFBE4764DFB15FDC9" box="[1132,1221,523,549]" pageId="190" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
These specimens do not cluster in any distinct groups, but rather are spread between and around the groupings of
|
||
<taxonomicName id="4C4C4D68FF52D58BFB62760AFAAFFD89" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1258,1407,587,613]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF52D58BFB62760AFAAFFD89" box="[1258,1407,587,613]" italics="true" pageId="190" pageNumber="191">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF52D58BFA02760DFC8BFD69" bold="true" pageId="190" pageNumber="191">
|
||
<taxonomicNameLabel id="A20B5782FF52D58BFA02760DFC8BFD69" pageId="190" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF52D58BFC1B762AFC22FD69" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[915,1010,620,645]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF52D58BFC1B762AFC22FD69" box="[915,1010,620,645]" italics="true" pageId="190" pageNumber="191">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF52D58BFC71762DFB84FD69" bold="true" box="[1017,1108,619,645]" pageId="190" pageNumber="191">
|
||
<taxonomicNameLabel id="A20B5782FF52D58BFC71762DFB84FD69" box="[1017,1108,619,645]" pageId="190" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
specimens. This demonstrates the inefficacy of diagnosing these two taxa based on the few small differences in dental dimension groupings, as these do not serve to delimit with any confidence the many remaining fragmentary specimens.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF52D58BFCD2774DFBF9FC09" blockId="190.[810,1452,171,1509]" pageId="190" pageNumber="191">
|
||
Because the
|
||
<typeStatus id="54F78849FF52D58BFC7B774DFBB0FCC9" box="[1011,1120,779,805]" pageId="190" pageNumber="191" type="holotype">holotypes</typeStatus>
|
||
of neither
|
||
<taxonomicName id="4C4C4D68FF52D58BFB63774AFA88FCC9" box="[1259,1368,779,805]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="brehus">
|
||
<emphasis id="B938EAF9FF52D58BFB63774AFA88FCC9" box="[1259,1368,779,805]" italics="true" pageId="190" pageNumber="191">P. brehus</emphasis>
|
||
</taxonomicName>
|
||
nor
|
||
<taxonomicName id="4C4C4D68FF52D58BFA1F774AFC52FCA9" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="roechus">
|
||
<emphasis id="B938EAF9FF52D58BFA1F774AFC52FCA9" italics="true" pageId="190" pageNumber="191">P. roechus</emphasis>
|
||
</taxonomicName>
|
||
can be unambiguously associated with either one of the two Pleistocene forms identifiable on the basis of partial and complete skeletons, neither nomina can be used to describe the taxa observed. We therefore recognise
|
||
<taxonomicName id="4C4C4D68FF52D58BFC1577EAFB7BFC29" authority="(Owen, 1874)" baseAuthorityName="Owen" baseAuthorityYear="1874" box="[925,1195,939,965]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="brehus">
|
||
<emphasis id="B938EAF9FF52D58BFC1577EAFBD5FC29" box="[925,1029,939,965]" italics="true" pageId="190" pageNumber="191">P. brehus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDD4B1AFF52D58BFB9E77EDFB73FC29" author="Owen, R." box="[1046,1187,939,965]" pageId="190" pageNumber="191" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen, 1874</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF52D58BFB6C77EAFCB2FC09" authority="Owen, 1874" authorityName="Owen" authorityYear="1874" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="roechus">
|
||
<emphasis id="B938EAF9FF52D58BFB6C77EAFA89FC29" box="[1252,1369,939,965]" italics="true" pageId="190" pageNumber="191">P. roechus</emphasis>
|
||
<bibRefCitation id="EFDD4B1AFF52D58BFAE977EDFCB2FC09" author="Owen, R." pageId="190" pageNumber="191" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen, 1874</bibRefCitation>
|
||
</taxonomicName>
|
||
as
|
||
<emphasis id="B938EAF9FF52D58BFC0F778AFBF2FC09" box="[903,1058,971,997]" italics="true" pageId="190" pageNumber="191">nomina dubia</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF52D58BFCD277ADFA47FA09" blockId="190.[810,1452,171,1509]" pageId="190" pageNumber="191">
|
||
This leaves
|
||
<taxonomicName id="4C4C4D68FF52D58BFC7377AAFACCFBE9" authority="Owen, 1874" authorityName="Owen" authorityYear="1874" box="[1019,1308,1003,1029]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="mimas">
|
||
<emphasis id="B938EAF9FF52D58BFC7377AAFBBEFBE9" box="[1019,1134,1004,1029]" italics="true" pageId="190" pageNumber="191">P. mimas</emphasis>
|
||
<bibRefCitation id="EFDD4B1AFF52D58BFB0B77ADFACCFBE9" author="Owen, R." box="[1155,1308,1003,1029]" pageId="190" pageNumber="191" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen, 1874</bibRefCitation>
|
||
</taxonomicName>
|
||
, previously synonymised with
|
||
<taxonomicName id="4C4C4D68FF52D58BFC75704AFBB4FBC9" baseAuthorityName="Owen" baseAuthorityYear="1874" box="[1021,1124,1035,1061]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="brehus">
|
||
<emphasis id="B938EAF9FF52D58BFC75704AFBB4FBC9" box="[1021,1124,1035,1061]" italics="true" pageId="190" pageNumber="191">P. brehus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C4C4D68FF52D58BFB2A704AFA77FBC9" authority="Owen, 1877" authorityName="Owen" authorityYear="1877" box="[1186,1447,1035,1061]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="antaeus">
|
||
<emphasis id="B938EAF9FF52D58BFB2A704AFAC4FBC9" box="[1186,1300,1036,1061]" italics="true" pageId="190" pageNumber="191">P. antaeus</emphasis>
|
||
<bibRefCitation id="EFDD4B1AFF52D58BFA93704DFA77FBC9" author="Owen, R." box="[1307,1447,1035,1061]" pageId="190" pageNumber="191" refId="ref152945" refString="Owen, R. (1877) Researches on the fossil remains of the extinct mammals of Australia; with a notice of the extinct marsupials of England. 1. J. Erxleben, London, 522 pp. https: // doi. org / 10.5962 / bhl. title. 77375" type="book" year="1877">Owen, 1877</bibRefCitation>
|
||
</taxonomicName>
|
||
, previously synonymised with
|
||
<taxonomicName id="4C4C4D68FF52D58BFB00706AFAD0FBA9" authorityName="Owen" authorityYear="1874" box="[1160,1280,1067,1093]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="roechus">
|
||
<emphasis id="B938EAF9FF52D58BFB00706AFAD0FBA9" box="[1160,1280,1067,1093]" italics="true" pageId="190" pageNumber="191">P. roechus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFDD4B1AFF52D58BFA9A706DFA7BFBA9" author="Owen, R." box="[1298,1451,1067,1093]" pageId="190" pageNumber="191" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen (1874)</bibRefCitation>
|
||
observed that
|
||
<taxonomicName id="4C4C4D68FF52D58BFC4D700AFBE6FB89" authorityName="Owen" authorityYear="1877" box="[965,1078,1100,1125]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="antaeus">
|
||
<emphasis id="B938EAF9FF52D58BFC4D700AFBE6FB89" box="[965,1078,1100,1125]" italics="true" pageId="190" pageNumber="191">P. antaeus</emphasis>
|
||
</taxonomicName>
|
||
differed from
|
||
<taxonomicName id="4C4C4D68FF52D58BFB5F700AFAEAFB89" authorityName="Owen" authorityYear="1874" box="[1239,1338,1100,1125]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="mimas">
|
||
<emphasis id="B938EAF9FF52D58BFB5F700AFAEAFB89" box="[1239,1338,1100,1125]" italics="true" pageId="190" pageNumber="191">P. mimas</emphasis>
|
||
</taxonomicName>
|
||
in its lack of a postcingulid and in having a larger p3 relative to its molars, and from
|
||
<taxonomicName id="4C4C4D68FF52D58BFC7070CAFBBEFB49" authorityName="Owen" authorityYear="1874" box="[1016,1134,1163,1189]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="roechus">
|
||
<emphasis id="B938EAF9FF52D58BFC7070CAFBBEFB49" box="[1016,1134,1163,1189]" italics="true" pageId="190" pageNumber="191">P. roechus</emphasis>
|
||
</taxonomicName>
|
||
in being smaller. However, because these traits and proportions vary within species (
|
||
<figureCitation id="13772A6EFF52D58BFCBA708DFC42FB09" box="[818,914,1227,1253]" captionStart="FIGURE 121" captionStartId="200.[136,235,2028,2052]" captionTargetBox="[158,1429,1160,2002]" captionTargetId="figure-67@200.[151,1436,1153,2009]" captionTargetPageId="200" captionText="FIGURE 121. boxplot of p3 length divided by m3 length in specimens of P. anak, P. mamkurra sp. nov., P. viator sp. nov., P. dawsonae sp. nov., P. snewini, P. otibandus, P. tumbuna, and four extrageneric taxa." figureDoi="http://doi.org/10.5281/zenodo.10994038" httpUri="https://zenodo.org/record/10994038/files/figure.png" pageId="190" pageNumber="191">Fig. 121</figureCitation>
|
||
) (see also
|
||
<bibRefCitation id="EFDD4B1AFF52D58BFB83708DFB09FB09" author="Bartholomai, A." box="[1035,1241,1227,1253]" pageId="190" pageNumber="191" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai 1973</bibRefCitation>
|
||
, p. 346) to a much greater degree than separates the
|
||
<typeStatus id="54F78849FF52D58BFB1770ADFADCFAE9" box="[1183,1292,1259,1285]" pageId="190" pageNumber="191" type="holotype">holotypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4C4D68FF52D58BFABA70AAFA74FAE9" authorityName="Owen" authorityYear="1877" box="[1330,1444,1260,1285]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="antaeus">
|
||
<emphasis id="B938EAF9FF52D58BFABA70AAFA74FAE9" box="[1330,1444,1260,1285]" italics="true" pageId="190" pageNumber="191">P. antaeus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4C4D68FF52D58BFCA2714AFC42FAC9" authorityName="Owen" authorityYear="1874" box="[810,914,1292,1317]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="mimas">
|
||
<emphasis id="B938EAF9FF52D58BFCA2714AFC42FAC9" box="[810,914,1292,1317]" italics="true" pageId="190" pageNumber="191">P. mimas</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF52D58BFC58714AFB97FAC9" authorityName="Owen" authorityYear="1874" box="[976,1095,1291,1317]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="roechus">
|
||
<emphasis id="B938EAF9FF52D58BFC58714AFB97FAC9" box="[976,1095,1291,1317]" italics="true" pageId="190" pageNumber="191">P. roechus</emphasis>
|
||
</taxonomicName>
|
||
, and because the
|
||
<typeStatus id="54F78849FF52D58BFA94714DFA59FAC9" box="[1308,1417,1291,1317]" pageId="190" pageNumber="191" type="holotype">holotypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4C4D68FF52D58BFCA2716AFC40FAA9" authorityName="Owen" authorityYear="1874" box="[810,912,1324,1349]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="mimas">
|
||
<emphasis id="B938EAF9FF52D58BFCA2716AFC40FAA9" box="[810,912,1324,1349]" italics="true" pageId="190" pageNumber="191">P. mimas</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF52D58BFC44716AFB90FAA9" authorityName="Owen" authorityYear="1877" box="[972,1088,1324,1349]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="antaeus">
|
||
<emphasis id="B938EAF9FF52D58BFC44716AFB90FAA9" box="[972,1088,1324,1349]" italics="true" pageId="190" pageNumber="191">P. antaeus</emphasis>
|
||
</taxonomicName>
|
||
, both partial left dentaries with p3–m4 with moderate wear (NHMUK PVOR43351 and NHMUK PVM2258, respectively), cannot be unambiguously associated with either
|
||
<taxonomicName id="4C4C4D68FF52D58BFB6171CAFAAFFA49" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1257,1407,1419,1445]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF52D58BFB6171CAFAAFFA49" box="[1257,1407,1419,1445]" italics="true" pageId="190" pageNumber="191">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF52D58BFA0271CDFC8BFA29" bold="true" pageId="190" pageNumber="191">
|
||
<taxonomicNameLabel id="A20B5782FF52D58BFA0271CDFC8BFA29" pageId="190" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
or
|
||
<taxonomicName id="4C4C4D68FF52D58BFC0E71EAFC37FA29" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[902,999,1452,1477]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF52D58BFC0E71EAFC37FA29" box="[902,999,1452,1477]" italics="true" pageId="190" pageNumber="191">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF52D58BFC7971EDFB9DFA29" bold="true" box="[1009,1101,1451,1477]" pageId="190" pageNumber="191">
|
||
<taxonomicNameLabel id="A20B5782FF52D58BFC7971EDFB9DFA29" box="[1009,1101,1451,1477]" pageId="190" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C4C4D68FF52D58BFBD671EAFA8BFA29" authority="Owen, 1874" authorityName="Owen" authorityYear="1874" box="[1118,1371,1451,1477]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="mimas">
|
||
<emphasis id="B938EAF9FF52D58BFBD671EAFB15FA29" box="[1118,1221,1452,1477]" italics="true" pageId="190" pageNumber="191">P. mimas</emphasis>
|
||
<bibRefCitation id="EFDD4B1AFF52D58BFB4771EDFA8BFA29" author="Owen, R." box="[1231,1371,1451,1477]" pageId="190" pageNumber="191" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen, 1874</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF52D58BFA1F71EAFBC1FA09" authority="Owen, 1877" authorityName="Owen" authorityYear="1877" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="190" pageNumber="191" phylum="Chordata" rank="species" species="antaeus">
|
||
<emphasis id="B938EAF9FF52D58BFA1F71EAFC51FA09" italics="true" pageId="190" pageNumber="191">P. antaeus</emphasis>
|
||
<bibRefCitation id="EFDD4B1AFF52D58BFC00718DFBC1FA09" author="Owen, R." box="[904,1041,1483,1509]" pageId="190" pageNumber="191" refId="ref152945" refString="Owen, R. (1877) Researches on the fossil remains of the extinct mammals of Australia; with a notice of the extinct marsupials of England. 1. J. Erxleben, London, 522 pp. https: // doi. org / 10.5962 / bhl. title. 77375" type="book" year="1877">Owen, 1877</bibRefCitation>
|
||
</taxonomicName>
|
||
are also considered
|
||
<emphasis id="B938EAF9FF52D58BFB7D718AFA40FA09" box="[1269,1424,1483,1509]" italics="true" pageId="190" pageNumber="191">nomina dubia</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |