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<document id="1293EA2207C78C447339EFE0C832E74E" ID-DOI="10.11646/zootaxa.4996.3.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="5074778" ID-ZooBank="50282C10-075C-4B2C-B61C-3A772D5BC2F2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625557780024" checkinUser="plazi" docAuthor="Hsueh, Pan-Wen" docDate="2021" docId="03E587C6FFA9FF8885A0FB9BFAD4F71D" docLanguage="en" docName="zootaxa.4996.3.4.pdf" docOrigin="Zootaxa 4996 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudonereis trimaculata Horst 1924" docType="treatment" docVersion="3" lastPageNumber="507" masterDocId="FFDCFFBEFFA3FF878537FFA5FFB0F06D" masterDocTitle="New species and record of Pseudonereis (Annelida, Polychaeta, Nereididae) from Taiwan" masterLastPageNumber="512" masterPageNumber="492" pageNumber="502" updateTime="1698992174721" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="8D0AC7534EC49979DEB4FD9E4EBB4B93">New species and record of Pseudonereis (Annelida, Polychaeta, Nereididae) from Taiwan</mods:title>
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<mods:namePart id="B31265A2EA765F53383503F49F809F8D">Hsueh, Pan-Wen</mods:namePart>
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<paragraph id="8BF336D0FFA9FF8D85A0FB9BFDD5F435" blockId="10.[151,802,1086,1149]" box="[151,613,1086,1113]" pageId="10" pageNumber="502">
<heading id="D0BB81BCFFA9FF8D85A0FB9BFDD5F435" bold="true" box="[151,613,1086,1113]" fontSize="11" level="1" pageId="10" pageNumber="502" reason="1">
<taxonomicName id="4C4C4D53FFA9FF8D85A0FB9BFDD5F435" ID-CoL="4P4ZB" authority="Horst, 1924" authorityName="Horst" authorityYear="1924" box="[151,613,1086,1113]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="502" phylum="Annelida" rank="species" species="trimaculata">
<emphasis id="B938EAC2FFA9FF8D85A0FB9BFDD5F435" bold="true" box="[151,613,1086,1113]" pageId="10" pageNumber="502">
<emphasis id="B938EAC2FFA9FF8D85A0FB9BFE7CF434" bold="true" box="[151,460,1086,1113]" italics="true" pageId="10" pageNumber="502">Pseudonereis trimaculata</emphasis>
<bibRefCitation id="EFDD4B21FFA9FF8D84E4FB9AFDD5F435" author="Horst, R." box="[467,613,1086,1113]" pageId="10" pageNumber="502" pagination="145 - 198" refId="ref11486" refString="Horst, R. (1924) Polychaeta errantia of the Siboga Expedition. Part III. Nereidae and Hesionidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, 24 (1 c), 145 - 198." type="journal article" year="1924">Horst, 1924</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="C356655BFFA9FF8D85A0FBC6FC92F410" box="[151,802,1123,1149]" pageId="10" pageNumber="502" type="description">
<paragraph id="8BF336D0FFA9FF8D85A0FBC6FC92F410" blockId="10.[151,802,1086,1149]" box="[151,802,1123,1149]" pageId="10" pageNumber="502">
<figureCitation id="13772A55FFA9FF8D85A0FBC6FEBFF410" box="[151,271,1123,1149]" captionStart="FIGURE 8" captionStartId="11.[151,250,1041,1065]" captionTargetBox="[151,1437,199,1018]" captionTargetId="figure-33@11.[151,1437,199,1018]" captionTargetPageId="11" captionText="FIGURE 8. Pseudonereis trimaculata Horst, 1924; A, specimen NMNS 8383-29, BC, specimen NMNS 8383-26, D, speci- men NMNS 8383-33, EF, specimen NMNS 8383-27:A, anterior body, dorsal view; B, paragnaths of the pharynx, dorsal view; C, paragnaths of the pharynx, frontal view; D, anterior body, dorsal view; E, paragnaths of the pharynx, dorsal view; F, parag- naths of the pharynx, ventral view. Scale: A, D, 1.0 mm; BC, EF, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5074796" httpUri="https://zenodo.org/record/5074796/files/figure.png" pageId="10" pageNumber="502">Figs 8AF</figureCitation>
,
<figureCitation id="13772A55FFA9FF8D842DFBC6FEEAF410" box="[282,346,1123,1149]" captionStart="FIGURE 9" captionStartId="12.[151,250,1056,1080]" captionTargetBox="[154,1433,182,1032]" captionTargetId="figure-15@12.[154,1434,181,1032]" captionTargetPageId="12" captionText="FIGURE 9. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-29: A, right parapodium, anterior view, chaetiger 11; B, right parapodium, posterior view, chaetiger 11; C, right parapodium, chaetiger 35; D, right parapodium, chaetiger 50; E, right parapodium, chaetiger 60; F, right parapodium, chaetiger 76. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior lobe. Scale: AF, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074800" httpUri="https://zenodo.org/record/5074800/files/figure.png" pageId="10" pageNumber="502">9AF</figureCitation>
,
<figureCitation id="13772A55FFA9FF8D8451FBC6FE07F410" box="[358,439,1123,1149]" captionStart="FIGURE 10" captionStartId="13.[151,250,992,1016]" captionTargetBox="[151,1436,182,969]" captionTargetId="figure-33@13.[151,1436,181,969]" captionTargetPageId="13" captionText="FIGURE 10. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-29:A, notochaeta, homogomph spiniger, chaetiger 18; B, supra-acicular homogomph spiniger, chaetiger 45; C, supra-acicular short-bladed heterogomph falciger, chaetiger 45; D, subacicular heterogomph spiniger, chaetiger 45; E, subacicular short-bladed heterogomph falciger, chaetiger 45. Scale: AE, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.5074802" httpUri="https://zenodo.org/record/5074802/files/figure.png" pageId="10" pageNumber="502">10AE</figureCitation>
,
<figureCitation id="13772A55FFA9FF8D84F4FBC6FDA5F410" box="[451,533,1123,1149]" captionStart="FIGURE 11" captionStartId="14.[151,250,1918,1942]" captionTargetBox="[213,1375,237,1895]" captionTargetId="figure-38@14.[212,1375,237,1895]" captionTargetPageId="14" captionText="FIGURE 11. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, whole animal; B, anterior body, dorsal view; C, paragnaths of the pharynx, ventral view. Scale: A, 5.0 mm; B, 1.0 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5074804" httpUri="https://zenodo.org/record/5074804/files/figure.png" pageId="10" pageNumber="502">11AC</figureCitation>
,
<figureCitation id="13772A55FFA9FF8D8716FBC6FDDFF410" box="[545,623,1123,1149]" captionStart="FIGURE 12" captionStartId="15.[151,250,1051,1075]" captionTargetBox="[156,1430,181,1027]" captionTargetId="figure-33@15.[156,1430,181,1027]" captionTargetPageId="15" captionText="FIGURE 12. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, right parapodium, chaeti- ger 4; B, right parapodium, chaetiger 11; C, right parapodium, chaetiger 14; D, right parapodium, chaetiger 17; E, right para- podium, anterior view, chaetiger 54; F, right parapodium, anterior view, chaetiger 85. Abbreviation: DC, dorsal cirrus; DCLo, dorsal cirrus lobe; DCLa, dorsal cirrus lamellae; NeIA, neuropodial inferior acicular lobe; NePLa, neuropodial postchaetal lamellae; NeSA, neuropodial superior acicular lobe; NeVL, neuropodial ventral ligule; NoDL, notopodial dorsal ligule; NoVL, notopodial ventral ligule; VC; ventral cirrus; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AL, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074806" httpUri="https://zenodo.org/record/5074806/files/figure.png" pageId="10" pageNumber="502">12AF</figureCitation>
,
<figureCitation id="13772A55FFA9FF8D874DFBC6FD78F410" box="[634,712,1123,1149]" captionStart="FIGURE 13" captionStartId="18.[151,250,970,994]" captionTargetBox="[151,1436,181,946]" captionTargetId="figure-15@18.[151,1436,181,946]" captionTargetPageId="18" captionText="FIGURE 13. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, notochaeta, homogomph spiniger, chaetiger 10; B, supra-acicular homogomph spiniger, chaetiger 10; C, supra-acicular short-bladed heterogomph fal- ciger, chaetiger 10; D, parapodial chaetae, chaetiger 80; E, parapodial chaetae, chaetiger 54; F, subacicular short-bladed het- erogomph falciger, chaetiger 10. Abbreviation: NC, natatory chaetae; SubHeS, subacicular heterogomph spiniger; SubHoS, subacicular homogomph spiniger; SupHeF, supra-acicular heterogomph falciger. Scale: AC, F, 0.01 mm; DE, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5074808" httpUri="https://zenodo.org/record/5074808/files/figure.png" pageId="10" pageNumber="502">13AF</figureCitation>
,
<figureCitation id="13772A55FFA9FF8D87E3FBC6FC92F410" box="[724,802,1123,1149]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[241,1346,181,1843]" captionTargetId="figure-33@19.[241,1346,181,1843]" captionTargetPageId="19" captionText="FIGURE 14. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, male: A, anterior body, dorsal view; B, paragnaths of the pharynx, frontal view; C, left parapodium, posterior view, chaetiger 3; D, left parapodium, posterior view, chaetiger 10; E, left parapodium, posterior view, chaetiger 54; F, left parapodium, posterior view, chaetiger 78. Abbrevia- tion: DC, dorsal cirrus; DCLa, dorsal cirrus lamellae; NePLa, neuropodial postchaetal lamellae; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AB, 0.5 mm; CF, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5074810" httpUri="https://zenodo.org/record/5074810/files/figure.png" pageId="10" pageNumber="502">14AF</figureCitation>
</paragraph>
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<paragraph id="8BF336D0FFA9FF8D85A0FB0DFB17F4D2" blockId="10.[151,1191,1191,1277]" box="[151,1191,1191,1215]" pageId="10" pageNumber="502">
<treatmentCitationGroup id="AB5C11FEFFA9FF8D85A0FB0DFB17F4D2" box="[151,1191,1191,1215]" pageId="10" pageNumber="502">
<treatmentCitation id="0AED10C1FFA9FF8D85A0FB0DFD63F4D2" author="Horst, R." box="[151,723,1191,1215]" page="188" pageId="10" pageNumber="502" year="1924">
<taxonomicName id="4C4C4D53FFA9FF8D85A0FB0DFD63F4D2" ID-CoL="474G6" authority="Horst 1924: 187 - 188" authorityName="Horst" authorityPageNumber="187 - 188" authorityYear="1924" box="[151,723,1191,1215]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="502" phylum="Annelida" rank="species" species="trimaculata" subGenus="Pseudonereis">
<emphasis id="B938EAC2FFA9FF8D85A0FB0DFF68F4D2" box="[151,216,1192,1215]" italics="true" pageId="10" pageNumber="502">Nereis</emphasis>
(
<emphasis id="B938EAC2FFA9FF8D85D0FB02FEDFF4D2" box="[231,367,1191,1215]" italics="true" pageId="10" pageNumber="502">Pseudonereis</emphasis>
)
<emphasis id="B938EAC2FFA9FF8D844AFB02FE43F4D2" box="[381,499,1191,1215]" italics="true" pageId="10" pageNumber="502">trimaculata</emphasis>
<bibRefCitation id="EFDD4B21FFA9FF8D84CDFB0DFD63F4D2" author="Horst, R." box="[506,723,1191,1215]" pageId="10" pageNumber="502" pagination="145 - 198" refId="ref11486" refString="Horst, R. (1924) Polychaeta errantia of the Siboga Expedition. Part III. Nereidae and Hesionidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, 24 (1 c), 145 - 198." type="journal article" year="1924">Horst 1924: 187188</bibRefCitation>
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, pl. XXXVI, figs 8, 9. — Hartman 1959: 279.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF336D0FFA9FF8D85A0FB63FBE1F4B3" blockId="10.[151,1191,1191,1277]" box="[151,1105,1222,1246]" pageId="10" pageNumber="502">
<taxonomicName id="4C4C4D53FFA9FF8D85A0FB63FE2FF4B0" ID-CoL="6WFK6" baseAuthorityName="Augener" baseAuthorityYear="1913" box="[151,415,1222,1246]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="502" phylum="Annelida" rank="species" species="rottnestiana">
<emphasis id="B938EAC2FFA9FF8D85A0FB63FE2FF4B0" box="[151,415,1222,1246]" italics="true" pageId="10" pageNumber="502">Pseudonereis rottnestiana</emphasis>
</taxonomicName>
forma
<emphasis id="B938EAC2FFA9FF8D84DFFB63FDCDF4B3" box="[488,637,1222,1246]" italics="true" pageId="10" pageNumber="502">costatodentata</emphasis>
Hartmann-Schröder 1979: 119, figs 220222.
</paragraph>
<paragraph id="8BF336D0FFA9FF8D85A0FB41FCBBF490" blockId="10.[151,1191,1191,1277]" box="[151,779,1252,1277]" pageId="10" pageNumber="502">
<treatmentCitationGroup id="AB5C11FEFFA9FF8D85A0FB41FCBBF490" box="[151,779,1252,1277]" pageId="10" pageNumber="502">
<treatmentCitation id="0AED10C1FFA9FF8D85A0FB41FD20F491" author="Bakken, T." box="[151,656,1252,1277]" page="168" pageId="10" pageNumber="502" year="2007">
<taxonomicName id="4C4C4D53FFA9FF8D85A0FB41FD20F491" ID-CoL="4P4ZB" authority="Bakken 2007: 166 - 168" authorityName="Bakken" authorityPageNumber="166 - 168" authorityYear="2007" box="[151,656,1252,1277]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="502" phylum="Annelida" rank="species" species="trimaculata">
<emphasis id="B938EAC2FFA9FF8D85A0FB41FE2BF491" box="[151,411,1252,1276]" italics="true" pageId="10" pageNumber="502">Pseudonereis trimaculata</emphasis>
<bibRefCitation id="EFDD4B21FFA9FF8D8495FB41FD20F491" author="Bakken, T." box="[418,656,1252,1277]" pageId="10" pageNumber="502" pagination="145 - 176" refId="ref11104" refString="Bakken, T. (2007) Revision of Pseudonereis (Polychaeta, Nereididae). Zoological Journal of the Linnean Society, 150 (1), 145 - 176. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00289. x" type="journal article" year="2007">Bakken 2007: 166168</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 13AI.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C356655BFFA9FF8D85A0FA8FFDF8F608" pageId="10" pageNumber="502" type="materials_examined">
<paragraph id="8BF336D0FFA9FF8D85A0FA8FFDF8F608" blockId="10.[151,1437,1322,2033]" pageId="10" pageNumber="502">
<emphasis id="B938EAC2FFA9FF8D85A0FA8FFE33F529" bold="true" box="[151,387,1322,1348]" pageId="10" pageNumber="502">Material examined.</emphasis>
Atoke:
<materialsCitation id="3B243C8DFFA9FF8D84D4FA8EFCEFF504" collectingDate="2006-06-03" collectionCode="NMNS" collectorName="Oldman Rock" country="Taiwan" latitude="22.005" location="Lanyu Island" longLatPrecision="125" longitude="121.57833" pageId="10" pageNumber="502" specimenCode="NMNS 8383-26" specimenCount="1" stateProvince="Taitung County">
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(
<specimenCode id="DBEA9EABFFA9FF8D8741FA8EFC84F52B" box="[630,820,1323,1350]" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="10" pageNumber="502">NMNS 8383-26</specimenCode>
),
<collectorName id="26B95306FFA9FF8D867DFA8EFC59F528" box="[842,1001,1323,1349]" pageId="10" pageNumber="502">Oldman Rock</collectorName>
(
<geoCoordinate id="EE785017FFA9FF8D86CCFA8EFBEFF528" box="[1019,1119,1323,1349]" degrees="22" direction="north" minutes="0.3" orientation="latitude" pageId="10" pageNumber="502" precision="92" value="22.005">22°0.3'N</geoCoordinate>
<geoCoordinate id="EE785017FFA9FF8D815EFA8EFB59F528" box="[1129,1257,1323,1349]" degrees="121" direction="east" minutes="34.7" orientation="longitude" pageId="10" pageNumber="502" precision="92" value="121.57833">121°34.7'E</geoCoordinate>
),
<location id="8E93600BFFA9FF8D8037FA8EFA28F528" LSID="urn:lsid:plazi:treatment:03E587C6FFA9FF8885A0FB9BFAD4F71D:8E93600BFFA9FF8D8037FA8EFA28F528" box="[1280,1432,1323,1349]" country="Taiwan" latitude="22.005" longLatPrecision="125" longitude="121.57833" name="Lanyu Island" pageId="10" pageNumber="502" stateProvince="Taitung County">Lanyu Island</location>
,
<collectingRegion id="4988F832FFA9FF8D85A0FAEAFEF9F504" box="[151,329,1359,1385]" inferredBySuffix="true" pageId="10" pageNumber="502">Taitung County</collectingRegion>
,
<collectingCountry id="F35B7640FFA9FF8D8463FAEAFE19F504" box="[340,425,1359,1385]" name="Taiwan" pageId="10" pageNumber="502">Taiwan</collectingCountry>
, rocky intertidal habitats,
<date id="FFF21010FFA9FF8D87E6FAEAFCEFF504" box="[721,863,1359,1385]" pageId="10" pageNumber="502" value="2006-06-03">
<collectingDate id="EFB6E9F8FFA9FF8D87E6FAEAFCEFF504" box="[721,863,1359,1385]" pageId="10" pageNumber="502" value="2006-06-03">3 June 2006</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B243C8DFFA9FF8D865CFAEAFB86F5E0" collectingDate="2007-10-14" collectionCode="NMNS" collectorName="Jihuei" country="Taiwan" latitude="23.116167" location="Taitung County" longLatPrecision="12" longitude="121.40517" pageId="10" pageNumber="502" specimenCode="NMNS 8383-27" specimenCount="2" stateProvince="Taitung">
<specimenCount id="9D4AFD59FFA9FF8D865CFAEAFC46F504" box="[875,1014,1359,1385]" pageId="10" pageNumber="502" type="generic">2 specimens</specimenCount>
(
<specimenCode id="DBEA9EABFFA9FF8D8132FAEAFB71F504" box="[1029,1217,1359,1385]" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="10" pageNumber="502">NMNS 8383-27</specimenCode>
),
<collectorName id="26B95306FFA9FF8D81E1FAEAFAA9F504" box="[1238,1305,1359,1385]" pageId="10" pageNumber="502">Jihuei</collectorName>
(
<geoCoordinate id="EE785017FFA9FF8D801DFAEAFA2CF504" box="[1322,1436,1359,1385]" degrees="23" direction="north" minutes="6.97" orientation="latitude" pageId="10" pageNumber="502" precision="9" value="23.116167">23°6.97'N</geoCoordinate>
<geoCoordinate id="EE785017FFA9FF8D85A0FAD6FE95F5E0" box="[151,293,1395,1421]" degrees="121" direction="east" minutes="24.31" orientation="longitude" pageId="10" pageNumber="502" precision="9" value="121.40517">121°24.31'E</geoCoordinate>
),
<collectingRegion id="4988F832FFA9FF8D840DFAD6FE5BF5E0" box="[314,491,1395,1421]" country="Taiwan" name="Taitung" pageId="10" pageNumber="502">Taitung County</collectingRegion>
,
<collectingCountry id="F35B7640FFA9FF8D84C0FAD6FDFCF5E0" box="[503,588,1395,1421]" name="Taiwan" pageId="10" pageNumber="502">Taiwan</collectingCountry>
, rocky intertidal habitats,
<date id="FFF21010FFA9FF8D8643FAD6FB86F5E0" box="[884,1078,1395,1421]" pageId="10" pageNumber="502" value="2007-10-14">
<collectingDate id="EFB6E9F8FFA9FF8D8643FAD6FB86F5E0" box="[884,1078,1395,1421]" pageId="10" pageNumber="502" value="2007-10-14">14 October 2007</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B243C8DFFA9FF8D8174FAD6FDE9F5DC" collectingDate="2010-10-09" collectionCode="NMNS" country="Taiwan" location="Jihuei" pageId="10" pageNumber="502" specimenCode="NMNS 8383-28" specimenCount="1" stateProvince="Taitung">
<specimenCount id="9D4AFD59FFA9FF8D8174FAD6FB74F5E0" box="[1091,1220,1395,1421]" pageId="10" pageNumber="502" type="generic">1 specimen</specimenCount>
(
<specimenCode id="DBEA9EABFFA9FF8D81E3FAD6FA3FF5E0" box="[1236,1423,1395,1421]" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="10" pageNumber="502">NMNS 8383-28</specimenCode>
),
<location id="8E93600BFFA9FF8D85A0FA32FF6DF5DC" LSID="urn:lsid:plazi:treatment:03E587C6FFA9FF8885A0FB9BFAD4F71D:8E93600BFFA9FF8D85A0FA32FF6DF5DC" box="[151,221,1431,1457]" country="Taiwan" name="Jihuei" pageId="10" pageNumber="502" stateProvince="Taitung">Jihuei</location>
,
<collectingRegion id="4988F832FFA9FF8D85DFFA32FE29F5DC" box="[232,409,1431,1457]" country="Taiwan" name="Taitung" pageId="10" pageNumber="502">Taitung County</collectingRegion>
,
<date id="FFF21010FFA9FF8D8492FA32FDE9F5DC" box="[421,601,1431,1457]" pageId="10" pageNumber="502" value="2010-10-09">
<collectingDate id="EFB6E9F8FFA9FF8D8492FA32FDE9F5DC" box="[421,601,1431,1457]" pageId="10" pageNumber="502" value="2010-10-09">9 October 2010</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B243C8DFFA9FF8D8752FA32FA33F5DC" box="[613,1411,1431,1457]" collectingDate="2014-03-27" collectionCode="NMNS" country="Taiwan" location="Jihuei" pageId="10" pageNumber="502" specimenCode="NMNS 8383-29" specimenCount="1" stateProvince="Taitung">
<specimenCount id="9D4AFD59FFA9FF8D8752FA32FD55F5DC" box="[613,741,1431,1457]" pageId="10" pageNumber="502" type="generic">1 specimen</specimenCount>
(
<specimenCode id="DBEA9EABFFA9FF8D87C3FA32FC00F5DC" box="[756,944,1431,1457]" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="10" pageNumber="502">NMNS 8383-29</specimenCode>
),
<location id="8E93600BFFA9FF8D86F3FA32FBBAF5DC" LSID="urn:lsid:plazi:treatment:03E587C6FFA9FF8885A0FB9BFAD4F71D:8E93600BFFA9FF8D86F3FA32FBBAF5DC" box="[964,1034,1431,1457]" country="Taiwan" name="Jihuei" pageId="10" pageNumber="502" stateProvince="Taitung">Jihuei</location>
,
<collectingRegion id="4988F832FFA9FF8D8122FA32FB76F5DC" box="[1045,1222,1431,1457]" country="Taiwan" name="Taitung" pageId="10" pageNumber="502">Taitung County</collectingRegion>
,
<date id="FFF21010FFA9FF8D81E5FA32FA33F5DC" box="[1234,1411,1431,1457]" pageId="10" pageNumber="502" value="2014-03-27">
<collectingDate id="EFB6E9F8FFA9FF8D81E5FA32FA33F5DC" box="[1234,1411,1431,1457]" pageId="10" pageNumber="502" value="2014-03-27">27 March 2014</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B243C8DFFA9FF8D80B9FA32FBBEF5B8" collectingDate="2014-09-27" collectionCode="NMNS" country="Taiwan" location="Jihuei" pageId="10" pageNumber="502" specimenCode="NMNS 8383-28, 30" specimenCount="7" stateProvince="Taitung">
<specimenCount id="9D4AFD59FFA9FF8D80B9FA32FEBCF5B8" pageId="10" pageNumber="502" type="generic">7 specimens</specimenCount>
(
<specimenCode id="DBEA9EABFFA9FF8D842AFA1EFE68F5B8" box="[285,472,1467,1493]" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="10" pageNumber="502">NMNS 8383-28</specimenCode>
<specimenCode id="DBEA9EABFFA9FF8D84D0FA1EFDB4F5B8" box="[487,516,1467,1493]" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="10" pageNumber="502">30</specimenCode>
),
<location id="8E93600BFFA9FF8D872DFA1EFDD0F5B8" LSID="urn:lsid:plazi:treatment:03E587C6FFA9FF8885A0FB9BFAD4F71D:8E93600BFFA9FF8D872DFA1EFDD0F5B8" box="[538,608,1467,1493]" country="Taiwan" name="Jihuei" pageId="10" pageNumber="502" stateProvince="Taitung">Jihuei</location>
,
<collectingRegion id="4988F832FFA9FF8D875BFA1EFCAFF5B8" box="[620,799,1467,1493]" country="Taiwan" name="Taitung" pageId="10" pageNumber="502">Taitung County</collectingRegion>
,
<date id="FFF21010FFA9FF8D861BFA1EFBBEF5B8" box="[812,1038,1467,1493]" pageId="10" pageNumber="502" value="2014-09-27">
<collectingDate id="EFB6E9F8FFA9FF8D861BFA1EFBBEF5B8" box="[812,1038,1467,1493]" pageId="10" pageNumber="502" value="2014-09-27">27 September 2014</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B243C8DFFA9FF8D812CFA1EFD9DF594" collectingDate="2017-04-29" collectionCode="NMNS" country="Taiwan" location="Taitung County" pageId="10" pageNumber="502" specimenCode="NMNS 8383-31" specimenCount="1" stateProvince="Taitung">
<specimenCount id="9D4AFD59FFA9FF8D812CFA1EFB2DF5B8" box="[1051,1181,1467,1493]" pageId="10" pageNumber="502" type="generic">1 specimen</specimenCount>
(
<specimenCode id="DBEA9EABFFA9FF8D819AFA1EFADAF5B8" box="[1197,1386,1467,1493]" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="10" pageNumber="502">NMNS 8383-31</specimenCode>
), Ji- huei,
<collectingRegion id="4988F832FFA9FF8D85E2FA7AFE34F594" box="[213,388,1503,1529]" country="Taiwan" name="Taitung" pageId="10" pageNumber="502">Taitung County</collectingRegion>
,
<date id="FFF21010FFA9FF8D84B8FA7AFD9DF594" box="[399,557,1503,1529]" pageId="10" pageNumber="502" value="2017-04-29">
<collectingDate id="EFB6E9F8FFA9FF8D84B8FA7AFD9DF594" box="[399,557,1503,1529]" pageId="10" pageNumber="502" value="2017-04-29">29 April 2017</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B243C8DFFA9FF8D870FFA7BFA33F594" box="[568,1411,1502,1529]" collectingDate="2019-11-17" collectionCode="NMNS" country="Taiwan" location="Jihuei" pageId="10" pageNumber="502" specimenCode="NMNS 8383-32" specimenCount="5" stateProvince="Taitung">
<specimenCount id="9D4AFD59FFA9FF8D870FFA7BFD72F594" box="[568,706,1502,1529]" pageId="10" pageNumber="502" type="generic">5 specimens</specimenCount>
(
<specimenCode id="DBEA9EABFFA9FF8D87E7FA7AFC3BF594" box="[720,907,1503,1529]" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="10" pageNumber="502">NMNS 8383-32</specimenCode>
),
<location id="8E93600BFFA9FF8D86A9FA7AFC54F594" LSID="urn:lsid:plazi:treatment:03E587C6FFA9FF8885A0FB9BFAD4F71D:8E93600BFFA9FF8D86A9FA7AFC54F594" box="[926,996,1503,1529]" country="Taiwan" name="Jihuei" pageId="10" pageNumber="502" stateProvince="Taitung">Jihuei</location>
,
<collectingRegion id="4988F832FFA9FF8D86D9FA7AFB2EF594" box="[1006,1182,1503,1529]" country="Taiwan" name="Taitung" pageId="10" pageNumber="502">Taitung County</collectingRegion>
,
<date id="FFF21010FFA9FF8D819FFA7AFA33F594" box="[1192,1411,1503,1529]" pageId="10" pageNumber="502" value="2019-11-17">
<collectingDate id="EFB6E9F8FFA9FF8D819FFA7AFA33F594" box="[1192,1411,1503,1529]" pageId="10" pageNumber="502" value="2019-11-17">17 November 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B243C8DFFA9FF8D80B9FA7BFEE8F62C" collectingDate="2020-10-17" collectionCode="NMNS" collectorName="Kentin" country="Taiwan" latitude="20.911667" location="Pingtung County" longLatPrecision="126" longitude="121.846664" pageId="10" pageNumber="502" specimenCode="NMNS 8383-33" specimenCount="5" stateProvince="Pingtung">
<specimenCount id="9D4AFD59FFA9FF8D80B9FA7BFEBCF670" pageId="10" pageNumber="502" type="generic">5 specimens</specimenCount>
(
<specimenCode id="DBEA9EABFFA9FF8D842CF9A6FE67F670" box="[283,471,1539,1565]" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="10" pageNumber="502">NMNS 8383-33</specimenCode>
),
<collectorName id="26B95306FFA9FF8D84DDF9A6FD86F670" box="[490,566,1539,1565]" pageId="10" pageNumber="502">Kentin</collectorName>
((
<geoCoordinate id="EE785017FFA9FF8D8767F9A7FD73F670" box="[592,707,1538,1565]" degrees="20" direction="north" minutes="54.7" orientation="latitude" pageId="10" pageNumber="502" precision="92" value="20.911667">20°54.7'N</geoCoordinate>
<geoCoordinate id="EE785017FFA9FF8D87FDF9A7FCF9F670" box="[714,841,1538,1565]" degrees="121" direction="east" minutes="50.8" orientation="longitude" pageId="10" pageNumber="502" precision="92" value="121.846664">121°50.8'E</geoCoordinate>
),
<collectingRegion id="4988F832FFA9FF8D8669F9A6FBAFF670" box="[862,1055,1539,1565]" country="Taiwan" name="Pingtung" pageId="10" pageNumber="502">Pingtung County</collectingRegion>
,
<collectingCountry id="F35B7640FFA9FF8D811DF9A6FBCFF670" box="[1066,1151,1539,1565]" name="Taiwan" pageId="10" pageNumber="502">Taiwan</collectingCountry>
, rocky intertidal habitats,
<date id="FFF21010FFA9FF8D85A0F982FEE8F62C" box="[151,344,1575,1601]" pageId="10" pageNumber="502" value="2020-10-17">
<collectingDate id="EFB6E9F8FFA9FF8D85A0F982FEE8F62C" box="[151,344,1575,1601]" pageId="10" pageNumber="502" value="2020-10-17">17 October 2020</collectingDate>
</date>
</materialsCitation>
. Epitoke:
<materialsCitation id="3B243C8DFFA9FF8D84FEF982FDF5F608" collectingDate="2019-11-17" collectionCode="NMNS" collectorName="Jihuei" country="Taiwan" latitude="23.116167" location="Taitung County" longLatPrecision="12" longitude="121.40517" pageId="10" pageNumber="502" specimenCode="NMNS 8383-34" specimenCount="2" stateProvince="Taitung">
<specimenCount id="9D4AFD59FFA9FF8D84FEF982FDE4F62C" box="[457,596,1575,1601]" pageId="10" pageNumber="502" type="generic">2 specimens</specimenCount>
(
<specimenCode id="DBEA9EABFFA9FF8D8754F982FCAEF62C" box="[611,798,1575,1601]" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="10" pageNumber="502">NMNS 8383-34</specimenCode>
),
<collectorName id="26B95306FFA9FF8D8605F982FCC5F62C" box="[818,885,1575,1601]" pageId="10" pageNumber="502">Jihuei</collectorName>
(
<geoCoordinate id="EE785017FFA9FF8D86B2F982FC47F62C" box="[901,1015,1575,1601]" degrees="23" direction="north" minutes="6.97" orientation="latitude" pageId="10" pageNumber="502" precision="9" value="23.116167">23°6.97'N</geoCoordinate>
<geoCoordinate id="EE785017FFA9FF8D86C8F982FB3DF62C" box="[1023,1165,1575,1601]" degrees="121" direction="east" minutes="24.31" orientation="longitude" pageId="10" pageNumber="502" precision="9" value="121.40517">121°24.31'E</geoCoordinate>
),
<collectingRegion id="4988F832FFA9FF8D8196F982FAE1F62C" box="[1185,1361,1575,1601]" country="Taiwan" name="Taitung" pageId="10" pageNumber="502">Taitung County</collectingRegion>
, rocky intertidal habitats,
<date id="FFF21010FFA9FF8D845FF9EEFDF5F608" box="[360,581,1611,1637]" pageId="10" pageNumber="502" value="2019-11-17">
<collectingDate id="EFB6E9F8FFA9FF8D845FF9EEFDF5F608" box="[360,581,1611,1637]" pageId="10" pageNumber="502" value="2019-11-17">17 November 2019</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C356655BFFA9FF8985F0F9CBFDFCF0B8" lastPageId="14" lastPageNumber="506" pageId="10" pageNumber="502" type="description">
<paragraph id="8BF336D0FFA9FF8D85F0F9CBFAFFF6BC" blockId="10.[151,1437,1322,2033]" pageId="10" pageNumber="502">
<emphasis id="B938EAC2FFA9FF8D85F0F9CBFEE3F6E5" bold="true" box="[199,339,1646,1672]" pageId="10" pageNumber="502">Description.</emphasis>
Atoke: Based on specimen NMNS 8383-29, complete, body length
<quantity id="4CB49B35FFA9FF8D811BF9CBFB3BF6E4" box="[1068,1163,1646,1673]" metricMagnitude="-2" metricUnit="m" metricValue="3.55" pageId="10" pageNumber="502" unit="mm" value="35.5">35.5 mm</quantity>
with 83 chaetigers, chaetiger 10 width
<quantity id="4CB49B35FFA9FF8D8406F937FE35F6C0" box="[305,389,1682,1709]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="502" unit="mm" value="1.5">1.5 mm</quantity>
, excluding parapodia; beige in alcohol (
<figureCitation id="13772A55FFA9FF8D861BF936FC31F6C0" box="[812,897,1683,1709]" captionStart="FIGURE 8" captionStartId="11.[151,250,1041,1065]" captionTargetBox="[151,1437,199,1018]" captionTargetId="figure-33@11.[151,1437,199,1018]" captionTargetPageId="11" captionText="FIGURE 8. Pseudonereis trimaculata Horst, 1924; A, specimen NMNS 8383-29, BC, specimen NMNS 8383-26, D, speci- men NMNS 8383-33, EF, specimen NMNS 8383-27:A, anterior body, dorsal view; B, paragnaths of the pharynx, dorsal view; C, paragnaths of the pharynx, frontal view; D, anterior body, dorsal view; E, paragnaths of the pharynx, dorsal view; F, parag- naths of the pharynx, ventral view. Scale: A, D, 1.0 mm; BC, EF, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5074796" httpUri="https://zenodo.org/record/5074796/files/figure.png" pageId="10" pageNumber="502">Fig. 8A</figureCitation>
). Specimens NMNS 8383-2627, 33 with pharynx everted and used for illustrating paragnath patterns and shape of ridge pattern in Areas VIVVI of the pharynx.
</paragraph>
<paragraph id="8BF336D0FFA9FF8D85F0F97EFDDCF79C" blockId="10.[151,1437,1322,2033]" pageId="10" pageNumber="502">
Prostomium wider than long, lateral antennae antero-lateral, longer than palps, palpophores globose, palpostyles conical. Four pairs of tentacular cirri, longest one reaching chaetiger 4. Two pairs of eyes, in trapezoidal arrangement, subequal in size. Apodous anterior segment about 1.4 times longer than chaetiger 1. Pharynx with light brown jaws, each with 6 teeth; paragnath pattern: I=1; II=40 (left), 38 (right), in 4 comb-like rows; III=60, in 4 transverse comb-like rows; IV=78 (left), 72 (right), in 5 comb-like rows with additional cones and p-bars in sigmoid towards the jaws; V=3, in triangle; VI=1 (left), 1 (right), shield-shaped bars; VIIVIII=24, p-bars and cones interspaced, p-bar row slightly behind cone row, one paragnath on each furrow and ridge regions (
<figureCitation id="13772A55FFA9FF8D816CF816FB62F7A0" box="[1115,1234,1971,1997]" captionStart="FIGURE 8" captionStartId="11.[151,250,1041,1065]" captionTargetBox="[151,1437,199,1018]" captionTargetId="figure-33@11.[151,1437,199,1018]" captionTargetPageId="11" captionText="FIGURE 8. Pseudonereis trimaculata Horst, 1924; A, specimen NMNS 8383-29, BC, specimen NMNS 8383-26, D, speci- men NMNS 8383-33, EF, specimen NMNS 8383-27:A, anterior body, dorsal view; B, paragnaths of the pharynx, dorsal view; C, paragnaths of the pharynx, frontal view; D, anterior body, dorsal view; E, paragnaths of the pharynx, dorsal view; F, parag- naths of the pharynx, ventral view. Scale: A, D, 1.0 mm; BC, EF, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5074796" httpUri="https://zenodo.org/record/5074796/files/figure.png" pageId="10" pageNumber="502">Fig. 8BF</figureCitation>
). Ridge pattern of areas VIVVI, υ-shaped (
<figureCitation id="13772A55FFA9FF8D84F2F872FDD0F79C" box="[453,608,2007,2033]" captionStart="FIGURE 8" captionStartId="11.[151,250,1041,1065]" captionTargetBox="[151,1437,199,1018]" captionTargetId="figure-33@11.[151,1437,199,1018]" captionTargetPageId="11" captionText="FIGURE 8. Pseudonereis trimaculata Horst, 1924; A, specimen NMNS 8383-29, BC, specimen NMNS 8383-26, D, speci- men NMNS 8383-33, EF, specimen NMNS 8383-27:A, anterior body, dorsal view; B, paragnaths of the pharynx, dorsal view; C, paragnaths of the pharynx, frontal view; D, anterior body, dorsal view; E, paragnaths of the pharynx, dorsal view; F, parag- naths of the pharynx, ventral view. Scale: A, D, 1.0 mm; BC, EF, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5074796" httpUri="https://zenodo.org/record/5074796/files/figure.png" pageId="10" pageNumber="502">Fig. 8B, DE</figureCitation>
).
</paragraph>
<caption id="DF336658FFA8FF8C85A0FBB4FC2DF4F8" ID-DOI="http://doi.org/10.5281/zenodo.5074796" ID-Zenodo-Dep="5074796" httpUri="https://zenodo.org/record/5074796/files/figure.png" pageId="11" pageNumber="503" startId="11.[151,250,1041,1065]" targetBox="[151,1437,199,1018]" targetPageId="11">
<paragraph id="8BF336D0FFA8FF8C85A0FBB4FC2DF4F8" blockId="11.[151,1436,1041,1174]" pageId="11" pageNumber="503">
<emphasis id="B938EAC2FFA8FF8C85A0FBB4FEA4F444" bold="true" box="[151,276,1041,1065]" pageId="11" pageNumber="503">FIGURE 8.</emphasis>
<taxonomicName id="4C4C4D53FFA8FF8C842BFBB4FD15F447" authority="Horst, 1924" authorityName="Horst" authorityYear="1924" box="[284,677,1041,1066]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="503" phylum="Annelida" rank="species" species="trimaculata">
<emphasis id="B938EAC2FFA8FF8C842BFBB4FD91F444" box="[284,545,1041,1065]" italics="true" pageId="11" pageNumber="503">Pseudonereis trimaculata</emphasis>
<bibRefCitation id="EFDD4B21FFA8FF8C871EFBB7FD15F447" author="Horst, R." box="[553,677,1042,1066]" pageId="11" pageNumber="503" pagination="145 - 198" refId="ref11486" refString="Horst, R. (1924) Polychaeta errantia of the Siboga Expedition. Part III. Nereidae and Hesionidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, 24 (1 c), 145 - 198." type="journal article" year="1924">Horst, 1924</bibRefCitation>
</taxonomicName>
; A, specimen NMNS 8383-29, BC, specimen NMNS 8383-26, D, speci- men NMNS 8383-33, EF, specimen NMNS 8383-27: A, anterior body, dorsal view; B, paragnaths of the pharynx, dorsal view; C, paragnaths of the pharynx, frontal view; D, anterior body, dorsal view; E, paragnaths of the pharynx, dorsal view; F, parag- naths of the pharynx, ventral view. Scale: A, D, 1.0 mm; BC, EF, 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BF336D0FFA8FF8C85F0FB61FC8CF527" blockId="11.[151,1437,1220,2003]" pageId="11" pageNumber="503">
Dorsal cirri digitiform throughout, basally attached to dorsal ligule on anterior to mid-body chaetigers, about as long as dorsal ligule (
<figureCitation id="13772A55FFA8FF8C849DFB4DFD97F56F" box="[426,551,1256,1282]" captionStart="FIGURE 9" captionStartId="12.[151,250,1056,1080]" captionTargetBox="[154,1433,182,1032]" captionTargetId="figure-15@12.[154,1434,181,1032]" captionTargetPageId="12" captionText="FIGURE 9. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-29: A, right parapodium, anterior view, chaetiger 11; B, right parapodium, posterior view, chaetiger 11; C, right parapodium, chaetiger 35; D, right parapodium, chaetiger 50; E, right parapodium, chaetiger 60; F, right parapodium, chaetiger 76. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior lobe. Scale: AF, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074800" httpUri="https://zenodo.org/record/5074800/files/figure.png" pageId="11" pageNumber="503">Fig. 9AD</figureCitation>
), subdistally attached to dorsal ligule on mid-body to anterior half of posterior chaetigers, about 0.60.9 times longer than dorsal ligule (
<figureCitation id="13772A55FFA8FF8C861DFAA9FC11F54B" box="[810,929,1292,1318]" captionStart="FIGURE 9" captionStartId="12.[151,250,1056,1080]" captionTargetBox="[154,1433,182,1032]" captionTargetId="figure-15@12.[154,1434,181,1032]" captionTargetPageId="12" captionText="FIGURE 9. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-29: A, right parapodium, anterior view, chaetiger 11; B, right parapodium, posterior view, chaetiger 11; C, right parapodium, chaetiger 35; D, right parapodium, chaetiger 50; E, right parapodium, chaetiger 60; F, right parapodium, chaetiger 76. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior lobe. Scale: AF, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074800" httpUri="https://zenodo.org/record/5074800/files/figure.png" pageId="11" pageNumber="503">Fig. 9EF</figureCitation>
), distally attached to dorsal ligule on last 13 chaetigers, about 0.4 times as long as dorsal ligule (
<figureCitation id="13772A55FFA8FF8C87EDFA95FC9FF527" box="[730,815,1328,1354]" captionStart="FIGURE 9" captionStartId="12.[151,250,1056,1080]" captionTargetBox="[154,1433,182,1032]" captionTargetId="figure-15@12.[154,1434,181,1032]" captionTargetPageId="12" captionText="FIGURE 9. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-29: A, right parapodium, anterior view, chaetiger 11; B, right parapodium, posterior view, chaetiger 11; C, right parapodium, chaetiger 35; D, right parapodium, chaetiger 50; E, right parapodium, chaetiger 60; F, right parapodium, chaetiger 76. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior lobe. Scale: AF, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074800" httpUri="https://zenodo.org/record/5074800/files/figure.png" pageId="11" pageNumber="503">Fig. 9F</figureCitation>
).
</paragraph>
<paragraph id="8BF336D0FFA8FF8C85F0FAF1FC99F5B7" blockId="11.[151,1437,1220,2003]" pageId="11" pageNumber="503">
Dorsal ligule subconical on anterior chaetigers, subconical with blunt tips on anterior half of mid-body chaetigers, becoming small pennant-like lobe on posterior half of mid-body to posterior chaetigers (
<figureCitation id="13772A55FFA8FF8C81EDFADDFAE5F5FF" box="[1242,1365,1400,1426]" captionStart="FIGURE 9" captionStartId="12.[151,250,1056,1080]" captionTargetBox="[154,1433,182,1032]" captionTargetId="figure-15@12.[154,1434,181,1032]" captionTargetPageId="12" captionText="FIGURE 9. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-29: A, right parapodium, anterior view, chaetiger 11; B, right parapodium, posterior view, chaetiger 11; C, right parapodium, chaetiger 35; D, right parapodium, chaetiger 50; E, right parapodium, chaetiger 60; F, right parapodium, chaetiger 76. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior lobe. Scale: AF, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074800" httpUri="https://zenodo.org/record/5074800/files/figure.png" pageId="11" pageNumber="503">Fig. 9AF</figureCitation>
); base of dorsal ligule slightly elongate and broader on mid-body chaetigers, markedly elongate and broader on posterior chaetigers (
<figureCitation id="13772A55FFA8FF8C842FFA65FE21F5B7" box="[280,401,1472,1498]" captionStart="FIGURE 9" captionStartId="12.[151,250,1056,1080]" captionTargetBox="[154,1433,182,1032]" captionTargetId="figure-15@12.[154,1434,181,1032]" captionTargetPageId="12" captionText="FIGURE 9. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-29: A, right parapodium, anterior view, chaetiger 11; B, right parapodium, posterior view, chaetiger 11; C, right parapodium, chaetiger 35; D, right parapodium, chaetiger 50; E, right parapodium, chaetiger 60; F, right parapodium, chaetiger 76. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior lobe. Scale: AF, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074800" httpUri="https://zenodo.org/record/5074800/files/figure.png" pageId="11" pageNumber="503">Fig. 9DF</figureCitation>
). Notopodial prechaetal lobe absent.
</paragraph>
<paragraph id="8BF336D0FFA8FF8C85F0FA41FB97F64F" blockId="11.[151,1437,1220,2003]" pageId="11" pageNumber="503">
Median ligule subconical throughout, shorter than dorsal ligule and neuroacicular ligule on anterior chaetigers, longer than neuroacicular ligule on mid-body to posterior chaetigers (
<figureCitation id="13772A55FFA8FF8C8695F9ADFBABF64F" box="[930,1051,1544,1570]" captionStart="FIGURE 9" captionStartId="12.[151,250,1056,1080]" captionTargetBox="[154,1433,182,1032]" captionTargetId="figure-15@12.[154,1434,181,1032]" captionTargetPageId="12" captionText="FIGURE 9. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-29: A, right parapodium, anterior view, chaetiger 11; B, right parapodium, posterior view, chaetiger 11; C, right parapodium, chaetiger 35; D, right parapodium, chaetiger 50; E, right parapodium, chaetiger 60; F, right parapodium, chaetiger 76. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior lobe. Scale: AF, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074800" httpUri="https://zenodo.org/record/5074800/files/figure.png" pageId="11" pageNumber="503">Fig. 9AF</figureCitation>
).
</paragraph>
<paragraph id="8BF336D0FFA8FF8C85F0F989FCADF697" blockId="11.[151,1437,1220,2003]" pageId="11" pageNumber="503">
Neuroacicular ligule with predominant inferior lobe on anterior to mid-body chaetigers, inferior and superior lobes subequal in length on posterior half of mid-body to posterior chaetigers, about 1.3 times as long as ventral ligule on anterior to mid-body chaetigers, about as long as ventral ligule on posterior chaetigers. Neuropodial postchaetal lobe present throughout. Ventral ligule subconical throughout. Ventral cirri mid-ventrally attached to ventral edge of parapodia, about 0.7 times as long as ventral ligule on anterior to mid-body chaetigers, about 0.5 times as long as ventral ligule on posterior chaetigers (
<figureCitation id="13772A55FFA8FF8C87AFF945FCA1F697" box="[664,785,1760,1786]" captionStart="FIGURE 9" captionStartId="12.[151,250,1056,1080]" captionTargetBox="[154,1433,182,1032]" captionTargetId="figure-15@12.[154,1434,181,1032]" captionTargetPageId="12" captionText="FIGURE 9. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-29: A, right parapodium, anterior view, chaetiger 11; B, right parapodium, posterior view, chaetiger 11; C, right parapodium, chaetiger 35; D, right parapodium, chaetiger 50; E, right parapodium, chaetiger 60; F, right parapodium, chaetiger 76. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior lobe. Scale: AF, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074800" httpUri="https://zenodo.org/record/5074800/files/figure.png" pageId="11" pageNumber="503">Fig. 9AF</figureCitation>
).
</paragraph>
<paragraph id="8BF336D0FFA8FF8C85F0F8A1FF47F7C3" blockId="11.[151,1437,1220,2003]" pageId="11" pageNumber="503">
Notochaetae present from chaetiger 3 to posterior chaetigers, homogomph spinigers present throughout (
<figureCitation id="13772A55FFA8FF8C8058F8A1FF7BF72F" captionStart="FIGURE 10" captionStartId="13.[151,250,992,1016]" captionTargetBox="[151,1436,182,969]" captionTargetId="figure-33@13.[151,1436,181,969]" captionTargetPageId="13" captionText="FIGURE 10. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-29:A, notochaeta, homogomph spiniger, chaetiger 18; B, supra-acicular homogomph spiniger, chaetiger 45; C, supra-acicular short-bladed heterogomph falciger, chaetiger 45; D, subacicular heterogomph spiniger, chaetiger 45; E, subacicular short-bladed heterogomph falciger, chaetiger 45. Scale: AE, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.5074802" httpUri="https://zenodo.org/record/5074802/files/figure.png" pageId="11" pageNumber="503">Fig. 10A</figureCitation>
). Supra-acicular fascicle of neuropodia: homogomph spinigers and short-bladed heterogomph falcigers with serrations present (
<figureCitation id="13772A55FFA8FF8C8458F8E9FE4AF70B" box="[367,506,1868,1894]" captionStart="FIGURE 10" captionStartId="13.[151,250,992,1016]" captionTargetBox="[151,1436,182,969]" captionTargetId="figure-33@13.[151,1436,181,969]" captionTargetPageId="13" captionText="FIGURE 10. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-29:A, notochaeta, homogomph spiniger, chaetiger 18; B, supra-acicular homogomph spiniger, chaetiger 45; C, supra-acicular short-bladed heterogomph falciger, chaetiger 45; D, subacicular heterogomph spiniger, chaetiger 45; E, subacicular short-bladed heterogomph falciger, chaetiger 45. Scale: AE, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.5074802" httpUri="https://zenodo.org/record/5074802/files/figure.png" pageId="11" pageNumber="503">Fig. 10BC</figureCitation>
). Subacicular fascicle of neuropodia: a single heterogomph spiniger present from chaetiger 45 to posterior chaetigers and short-bladed heterogomph falcigers with serrations present throughout (
<figureCitation id="13772A55FFA8FF8C8058F8D5FF5BF7C3" captionStart="FIGURE 10" captionStartId="13.[151,250,992,1016]" captionTargetBox="[151,1436,182,969]" captionTargetId="figure-33@13.[151,1436,181,969]" captionTargetPageId="13" captionText="FIGURE 10. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-29:A, notochaeta, homogomph spiniger, chaetiger 18; B, supra-acicular homogomph spiniger, chaetiger 45; C, supra-acicular short-bladed heterogomph falciger, chaetiger 45; D, subacicular heterogomph spiniger, chaetiger 45; E, subacicular short-bladed heterogomph falciger, chaetiger 45. Scale: AE, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.5074802" httpUri="https://zenodo.org/record/5074802/files/figure.png" pageId="11" pageNumber="503">Fig. 10DE</figureCitation>
).
</paragraph>
<paragraph id="8BF336D0FFA8FF8C85F0F81DFC7EF7BF" blockId="11.[151,1437,1220,2003]" box="[199,974,1976,2003]" pageId="11" pageNumber="503">Pygidium crenulated; anal cirri cirriform, as long as last 5 chaetigers.</paragraph>
<caption id="DF336658FFAFFF8B85A0FB85FBB3F4CE" ID-DOI="http://doi.org/10.5281/zenodo.5074800" ID-Zenodo-Dep="5074800" httpUri="https://zenodo.org/record/5074800/files/figure.png" pageId="12" pageNumber="504" startId="12.[151,250,1056,1080]" targetBox="[154,1433,182,1032]" targetPageId="12">
<paragraph id="8BF336D0FFAFFF8B85A0FB85FBB3F4CE" blockId="12.[151,1436,1056,1188]" pageId="12" pageNumber="504">
<emphasis id="B938EAC2FFAFFF8B85A0FB85FEA4F455" bold="true" box="[151,276,1056,1080]" pageId="12" pageNumber="504">FIGURE 9.</emphasis>
<taxonomicName id="4C4C4D53FFAFFF8B842BFB85FD16F455" authority="Horst, 1924" authorityName="Horst" authorityYear="1924" box="[284,678,1056,1080]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="504" phylum="Annelida" rank="species" species="trimaculata">
<emphasis id="B938EAC2FFAFFF8B842BFB85FD92F455" box="[284,546,1056,1080]" italics="true" pageId="12" pageNumber="504">Pseudonereis trimaculata</emphasis>
<bibRefCitation id="EFDD4B21FFAFFF8B871DFB85FD16F455" author="Horst, R." box="[554,678,1056,1080]" pageId="12" pageNumber="504" pagination="145 - 198" refId="ref11486" refString="Horst, R. (1924) Polychaeta errantia of the Siboga Expedition. Part III. Nereidae and Hesionidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, 24 (1 c), 145 - 198." type="journal article" year="1924">Horst, 1924</bibRefCitation>
</taxonomicName>
; specimen NMNS 8383-29: A, right parapodium, anterior view, chaetiger 11; B, right parapodium, posterior view, chaetiger 11; C, right parapodium, chaetiger 35; D, right parapodium, chaetiger 50; E, right parapodium, chaetiger 60; F, right parapodium, chaetiger 76. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior lobe. Scale: AF, 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BF336D0FFAFFF8B85F0FB77FB04F5CC" blockId="12.[151,1437,1234,1980]" pageId="12" pageNumber="504">
<emphasis id="B938EAC2FFAFFF8B85F0FB77FE8FF481" bold="true" box="[199,319,1234,1260]" pageId="12" pageNumber="504">Variation.</emphasis>
Longest tentacular cirrus reached chaetiger 310 (n= 20). Apodous anterior segment ranged about 1.3-1.6 (n= 10) times longer than chaetiger 1. Number of lateral teeth on cutting edge of the jaws ranged 510 (n= 22). Paragnath numbers in Area II ranged 2949 (left) and 1846 (right) in 45 rows (n= 22), Area III
<quantity id="4CB49B35FFAFFF8B803EFABFFADCF559" box="[1289,1388,1306,1333]" metricMagnitude="0" metricUnit="m" metricValue="1.8415" metricValueMax="2.3114" metricValueMin="1.3716" pageId="12" pageNumber="504" unit="in" value="72.5" valueMax="91.0" valueMin="54.0">5491 in</quantity>
35 rows (n= 22), Area IV 3693 (left) and 44112 (right) in 5 rows (n= 22), Area V 12 (n= 22), Area VIIVIII 2230 (n= 22), respectively. Heterogomph spinigers in the subacicular fascicle of the neuropodia commenced on chaetiger 3741 (n=2). Length of anal cirri cirriform ranged from as long as last 510 chaetigers (n= 6).
</paragraph>
<paragraph id="8BF336D0FFAFFF8B85F0FA0FFC17F660" blockId="12.[151,1437,1234,1980]" pageId="12" pageNumber="504">
Epitoke: Based on specimen NMNS 8383-34 (1), ovigerous female, complete, body depressed, body length 32.0 mm with 99 chaetigers; beige in alcohol (
<figureCitation id="13772A55FFAFFF8B87A0FA6BFD4DF585" box="[663,765,1486,1512]" captionStart="FIGURE 11" captionStartId="14.[151,250,1918,1942]" captionTargetBox="[213,1375,237,1895]" captionTargetId="figure-38@14.[212,1375,237,1895]" captionTargetPageId="14" captionText="FIGURE 11. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, whole animal; B, anterior body, dorsal view; C, paragnaths of the pharynx, ventral view. Scale: A, 5.0 mm; B, 1.0 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5074804" httpUri="https://zenodo.org/record/5074804/files/figure.png" pageId="12" pageNumber="504">Fig. 11A</figureCitation>
). Body divided into two regions: 1) pre-natatory region from chaetiger 1 to 13; 2) natatory region from chaetiger 14 to end of body.
</paragraph>
<paragraph id="8BF336D0FFAFFF8B85F0F9B3FCEAF689" blockId="12.[151,1437,1234,1980]" pageId="12" pageNumber="504">
Prostomium and apodous segment similar to atoke except enlarged eyes (
<figureCitation id="13772A55FFAFFF8B86C5F9B3FBE8F65D" box="[1010,1112,1558,1584]" captionStart="FIGURE 11" captionStartId="14.[151,250,1918,1942]" captionTargetBox="[213,1375,237,1895]" captionTargetId="figure-38@14.[212,1375,237,1895]" captionTargetPageId="14" captionText="FIGURE 11. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, whole animal; B, anterior body, dorsal view; C, paragnaths of the pharynx, ventral view. Scale: A, 5.0 mm; B, 1.0 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5074804" httpUri="https://zenodo.org/record/5074804/files/figure.png" pageId="12" pageNumber="504">Fig. 11A</figureCitation>
). Jaws similar to atoke. Pharynx with dark brown jaws, each with 34 lateral teeth; paragnath pattern: I=1; II=42 (left), 41 (right), in 5 comb-like rows; III=65, in 4 transverse comb-like rows; IV=77 (left), 77 (right), in 5 comb-like rows with additional cones and p-bars in sigmoid towards the jaws; V=3, in triangle; VI=1 (left), 1 (right), shield-shaped bars; VIIVIII=26, p-bars and cones interspaced, p-bar row slightly behind cone row, one paragnath on each furrow and ridge regions (
<figureCitation id="13772A55FFAFFF8B8058F903FF59F689" captionStart="FIGURE 11" captionStartId="14.[151,250,1918,1942]" captionTargetBox="[213,1375,237,1895]" captionTargetId="figure-38@14.[212,1375,237,1895]" captionTargetPageId="14" captionText="FIGURE 11. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, whole animal; B, anterior body, dorsal view; C, paragnaths of the pharynx, ventral view. Scale: A, 5.0 mm; B, 1.0 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5074804" httpUri="https://zenodo.org/record/5074804/files/figure.png" pageId="12" pageNumber="504">Fig. 11BC</figureCitation>
). Ridge pattern of areas VIVVI, υ-shaped (
<figureCitation id="13772A55FFAFFF8B87D0F96FFCFDF689" box="[743,845,1738,1764]" captionStart="FIGURE 11" captionStartId="14.[151,250,1918,1942]" captionTargetBox="[213,1375,237,1895]" captionTargetId="figure-38@14.[212,1375,237,1895]" captionTargetPageId="14" captionText="FIGURE 11. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, whole animal; B, anterior body, dorsal view; C, paragnaths of the pharynx, ventral view. Scale: A, 5.0 mm; B, 1.0 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5074804" httpUri="https://zenodo.org/record/5074804/files/figure.png" pageId="12" pageNumber="504">Fig. 11B</figureCitation>
).
</paragraph>
<paragraph id="8BF336D0FFAFFF8B85F0F94AFE85F7D1" blockId="12.[151,1437,1234,1980]" pageId="12" pageNumber="504">
Pre-natatory region: Dorsal cirri robust with swollen base from chaetiger 1 to 7, basally attached to dorsal ligule, about 1.2 times longer than dorsal ligule (
<figureCitation id="13772A55FFAFFF8B87AAF8B7FCB5F741" box="[669,773,1810,1836]" captionStart="FIGURE 12" captionStartId="15.[151,250,1051,1075]" captionTargetBox="[156,1430,181,1027]" captionTargetId="figure-33@15.[156,1430,181,1027]" captionTargetPageId="15" captionText="FIGURE 12. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, right parapodium, chaeti- ger 4; B, right parapodium, chaetiger 11; C, right parapodium, chaetiger 14; D, right parapodium, chaetiger 17; E, right para- podium, anterior view, chaetiger 54; F, right parapodium, anterior view, chaetiger 85. Abbreviation: DC, dorsal cirrus; DCLo, dorsal cirrus lobe; DCLa, dorsal cirrus lamellae; NeIA, neuropodial inferior acicular lobe; NePLa, neuropodial postchaetal lamellae; NeSA, neuropodial superior acicular lobe; NeVL, neuropodial ventral ligule; NoDL, notopodial dorsal ligule; NoVL, notopodial ventral ligule; VC; ventral cirrus; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AL, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074806" httpUri="https://zenodo.org/record/5074806/files/figure.png" pageId="12" pageNumber="504">Fig. 12A</figureCitation>
), digitiform on chaetiger 8 to 13, basally attached to dorsal ligule, about as long as dorsal ligule (
<figureCitation id="13772A55FFAFFF8B870FF893FD2EF73D" box="[568,670,1846,1872]" captionStart="FIGURE 12" captionStartId="15.[151,250,1051,1075]" captionTargetBox="[156,1430,181,1027]" captionTargetId="figure-33@15.[156,1430,181,1027]" captionTargetPageId="15" captionText="FIGURE 12. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, right parapodium, chaeti- ger 4; B, right parapodium, chaetiger 11; C, right parapodium, chaetiger 14; D, right parapodium, chaetiger 17; E, right para- podium, anterior view, chaetiger 54; F, right parapodium, anterior view, chaetiger 85. Abbreviation: DC, dorsal cirrus; DCLo, dorsal cirrus lobe; DCLa, dorsal cirrus lamellae; NeIA, neuropodial inferior acicular lobe; NePLa, neuropodial postchaetal lamellae; NeSA, neuropodial superior acicular lobe; NeVL, neuropodial ventral ligule; NoDL, notopodial dorsal ligule; NoVL, notopodial ventral ligule; VC; ventral cirrus; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AL, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074806" httpUri="https://zenodo.org/record/5074806/files/figure.png" pageId="12" pageNumber="504">Fig. 12B</figureCitation>
). Dorsal ligule subconical, about as long as median ligule (
<figureCitation id="13772A55FFAFFF8B8006F893FF1CF718" captionStart="FIGURE 12" captionStartId="15.[151,250,1051,1075]" captionTargetBox="[156,1430,181,1027]" captionTargetId="figure-33@15.[156,1430,181,1027]" captionTargetPageId="15" captionText="FIGURE 12. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, right parapodium, chaeti- ger 4; B, right parapodium, chaetiger 11; C, right parapodium, chaetiger 14; D, right parapodium, chaetiger 17; E, right para- podium, anterior view, chaetiger 54; F, right parapodium, anterior view, chaetiger 85. Abbreviation: DC, dorsal cirrus; DCLo, dorsal cirrus lobe; DCLa, dorsal cirrus lamellae; NeIA, neuropodial inferior acicular lobe; NePLa, neuropodial postchaetal lamellae; NeSA, neuropodial superior acicular lobe; NeVL, neuropodial ventral ligule; NoDL, notopodial dorsal ligule; NoVL, notopodial ventral ligule; VC; ventral cirrus; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AL, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074806" httpUri="https://zenodo.org/record/5074806/files/figure.png" pageId="12" pageNumber="504">Fig. 12A, B</figureCitation>
). Medial ligule subconical. Neuroacicular ligule with predominant inferior lobe. Neuropodial postchaetal lobe present. Ventral ligule subconical. Ventral cirri robust on chaetiger 17, becoming digitiform from chaetiger 8 to 13 (
<figureCitation id="13772A55FFAFFF8B8597F807FE9AF7D1" box="[160,298,1954,1980]" captionStart="FIGURE 12" captionStartId="15.[151,250,1051,1075]" captionTargetBox="[156,1430,181,1027]" captionTargetId="figure-33@15.[156,1430,181,1027]" captionTargetPageId="15" captionText="FIGURE 12. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, right parapodium, chaeti- ger 4; B, right parapodium, chaetiger 11; C, right parapodium, chaetiger 14; D, right parapodium, chaetiger 17; E, right para- podium, anterior view, chaetiger 54; F, right parapodium, anterior view, chaetiger 85. Abbreviation: DC, dorsal cirrus; DCLo, dorsal cirrus lobe; DCLa, dorsal cirrus lamellae; NeIA, neuropodial inferior acicular lobe; NePLa, neuropodial postchaetal lamellae; NeSA, neuropodial superior acicular lobe; NeVL, neuropodial ventral ligule; NoDL, notopodial dorsal ligule; NoVL, notopodial ventral ligule; VC; ventral cirrus; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AL, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074806" httpUri="https://zenodo.org/record/5074806/files/figure.png" pageId="12" pageNumber="504">Fig. 12AB</figureCitation>
).
</paragraph>
<caption id="DF336658FFAEFF8A85A0FC45FF49F409" ID-DOI="http://doi.org/10.5281/zenodo.5074802" ID-Zenodo-Dep="5074802" httpUri="https://zenodo.org/record/5074802/files/figure.png" pageId="13" pageNumber="505" startId="13.[151,250,992,1016]" targetBox="[151,1436,182,969]" targetPageId="13">
<paragraph id="8BF336D0FFAEFF8A85A0FC45FF49F409" blockId="13.[151,1436,992,1125]" pageId="13" pageNumber="505">
<emphasis id="B938EAC2FFAEFF8A85A0FC45FEAFF394" bold="true" box="[151,287,992,1017]" pageId="13" pageNumber="505">FIGURE 10.</emphasis>
<taxonomicName id="4C4C4D53FFAEFF8A8412FC44FD18F394" authority="Horst, 1924" authorityName="Horst" authorityYear="1924" box="[293,680,993,1017]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="505" phylum="Annelida" rank="species" species="trimaculata">
<emphasis id="B938EAC2FFAEFF8A8412FC44FD98F394" box="[293,552,993,1017]" italics="true" pageId="13" pageNumber="505">Pseudonereis trimaculata</emphasis>
<bibRefCitation id="EFDD4B21FFAEFF8A8719FC44FD18F394" author="Horst, R." box="[558,680,993,1017]" pageId="13" pageNumber="505" pagination="145 - 198" refId="ref11486" refString="Horst, R. (1924) Polychaeta errantia of the Siboga Expedition. Part III. Nereidae and Hesionidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, 24 (1 c), 145 - 198." type="journal article" year="1924">Horst, 1924</bibRefCitation>
</taxonomicName>
; specimen NMNS 8383-29: A, notochaeta, homogomph spiniger, chaetiger 18; B, supra-acicular homogomph spiniger, chaetiger 45; C, supra-acicular short-bladed heterogomph falciger, chaetiger 45; D, subacicular heterogomph spiniger, chaetiger 45; E, subacicular short-bladed heterogomph falciger, chaetiger 45. Scale: AE, 0.01 mm.
</paragraph>
</caption>
<paragraph id="8BF336D0FFAEFF8A85F0FB36FD1EF550" blockId="13.[151,1437,1171,2025]" pageId="13" pageNumber="505">
Natatory region: Dorsal cirri digitiform to posterior end, basally attached to dorsal ligule on anterior to anterior half of posterior chaetigers, about as long as dorsal ligule, distally attached to dorsal ligule, about 0.4 times as long as dorsal ligule (
<figureCitation id="13772A55FFAEFF8A8460FB7EFE6EF498" box="[343,478,1243,1269]" captionStart="FIGURE 12" captionStartId="15.[151,250,1051,1075]" captionTargetBox="[156,1430,181,1027]" captionTargetId="figure-33@15.[156,1430,181,1027]" captionTargetPageId="15" captionText="FIGURE 12. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, right parapodium, chaeti- ger 4; B, right parapodium, chaetiger 11; C, right parapodium, chaetiger 14; D, right parapodium, chaetiger 17; E, right para- podium, anterior view, chaetiger 54; F, right parapodium, anterior view, chaetiger 85. Abbreviation: DC, dorsal cirrus; DCLo, dorsal cirrus lobe; DCLa, dorsal cirrus lamellae; NeIA, neuropodial inferior acicular lobe; NePLa, neuropodial postchaetal lamellae; NeSA, neuropodial superior acicular lobe; NeVL, neuropodial ventral ligule; NoDL, notopodial dorsal ligule; NoVL, notopodial ventral ligule; VC; ventral cirrus; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AL, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074806" httpUri="https://zenodo.org/record/5074806/files/figure.png" pageId="13" pageNumber="505">Fig. 12CF</figureCitation>
). Dorsal cirrus lobe present from chaetiger 14 (
<figureCitation id="13772A55FFAEFF8A8135FB7EFBDAF498" box="[1026,1130,1243,1269]" captionStart="FIGURE 12" captionStartId="15.[151,250,1051,1075]" captionTargetBox="[156,1430,181,1027]" captionTargetId="figure-33@15.[156,1430,181,1027]" captionTargetPageId="15" captionText="FIGURE 12. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, right parapodium, chaeti- ger 4; B, right parapodium, chaetiger 11; C, right parapodium, chaetiger 14; D, right parapodium, chaetiger 17; E, right para- podium, anterior view, chaetiger 54; F, right parapodium, anterior view, chaetiger 85. Abbreviation: DC, dorsal cirrus; DCLo, dorsal cirrus lobe; DCLa, dorsal cirrus lamellae; NeIA, neuropodial inferior acicular lobe; NePLa, neuropodial postchaetal lamellae; NeSA, neuropodial superior acicular lobe; NeVL, neuropodial ventral ligule; NoDL, notopodial dorsal ligule; NoVL, notopodial ventral ligule; VC; ventral cirrus; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AL, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074806" httpUri="https://zenodo.org/record/5074806/files/figure.png" pageId="13" pageNumber="505">Fig. 12C</figureCitation>
), becoming kidney-shaped lamellae from chaetiger 17 to mid-body chaetigers (
<figureCitation id="13772A55FFAEFF8A87E0FB5AFCEFF574" box="[727,863,1279,1305]" captionStart="FIGURE 12" captionStartId="15.[151,250,1051,1075]" captionTargetBox="[156,1430,181,1027]" captionTargetId="figure-33@15.[156,1430,181,1027]" captionTargetPageId="15" captionText="FIGURE 12. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, right parapodium, chaeti- ger 4; B, right parapodium, chaetiger 11; C, right parapodium, chaetiger 14; D, right parapodium, chaetiger 17; E, right para- podium, anterior view, chaetiger 54; F, right parapodium, anterior view, chaetiger 85. Abbreviation: DC, dorsal cirrus; DCLo, dorsal cirrus lobe; DCLa, dorsal cirrus lamellae; NeIA, neuropodial inferior acicular lobe; NePLa, neuropodial postchaetal lamellae; NeSA, neuropodial superior acicular lobe; NeVL, neuropodial ventral ligule; NoDL, notopodial dorsal ligule; NoVL, notopodial ventral ligule; VC; ventral cirrus; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AL, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074806" httpUri="https://zenodo.org/record/5074806/files/figure.png" pageId="13" pageNumber="505">Fig. 12DE</figureCitation>
), reduced progressively in size thereafter, absent on posterior half of posterior chaetigers (
<figureCitation id="13772A55FFAEFF8A8709FA86FD12F550" box="[574,674,1315,1341]" captionStart="FIGURE 12" captionStartId="15.[151,250,1051,1075]" captionTargetBox="[156,1430,181,1027]" captionTargetId="figure-33@15.[156,1430,181,1027]" captionTargetPageId="15" captionText="FIGURE 12. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, right parapodium, chaeti- ger 4; B, right parapodium, chaetiger 11; C, right parapodium, chaetiger 14; D, right parapodium, chaetiger 17; E, right para- podium, anterior view, chaetiger 54; F, right parapodium, anterior view, chaetiger 85. Abbreviation: DC, dorsal cirrus; DCLo, dorsal cirrus lobe; DCLa, dorsal cirrus lamellae; NeIA, neuropodial inferior acicular lobe; NePLa, neuropodial postchaetal lamellae; NeSA, neuropodial superior acicular lobe; NeVL, neuropodial ventral ligule; NoDL, notopodial dorsal ligule; NoVL, notopodial ventral ligule; VC; ventral cirrus; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AL, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074806" httpUri="https://zenodo.org/record/5074806/files/figure.png" pageId="13" pageNumber="505">Fig. 12F</figureCitation>
).
</paragraph>
<paragraph id="8BF336D0FFAEFF8A85F0FAE2FCDFF5E8" blockId="13.[151,1437,1171,2025]" pageId="13" pageNumber="505">
Dorsal ligule subconical with blunt tips on anterior to anterior half of posterior chaetigers (
<figureCitation id="13772A55FFAEFF8A81F6FAE2FAF6F50C" box="[1217,1350,1351,1377]" captionStart="FIGURE 12" captionStartId="15.[151,250,1051,1075]" captionTargetBox="[156,1430,181,1027]" captionTargetId="figure-33@15.[156,1430,181,1027]" captionTargetPageId="15" captionText="FIGURE 12. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, right parapodium, chaeti- ger 4; B, right parapodium, chaetiger 11; C, right parapodium, chaetiger 14; D, right parapodium, chaetiger 17; E, right para- podium, anterior view, chaetiger 54; F, right parapodium, anterior view, chaetiger 85. Abbreviation: DC, dorsal cirrus; DCLo, dorsal cirrus lobe; DCLa, dorsal cirrus lamellae; NeIA, neuropodial inferior acicular lobe; NePLa, neuropodial postchaetal lamellae; NeSA, neuropodial superior acicular lobe; NeVL, neuropodial ventral ligule; NoDL, notopodial dorsal ligule; NoVL, notopodial ventral ligule; VC; ventral cirrus; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AL, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074806" httpUri="https://zenodo.org/record/5074806/files/figure.png" pageId="13" pageNumber="505">Fig. 12CF</figureCitation>
), markedly elongate on posterior half of posterior chaetigers (
<figureCitation id="13772A55FFAEFF8A87C9FACEFCD2F5E8" box="[766,866,1387,1413]" captionStart="FIGURE 12" captionStartId="15.[151,250,1051,1075]" captionTargetBox="[156,1430,181,1027]" captionTargetId="figure-33@15.[156,1430,181,1027]" captionTargetPageId="15" captionText="FIGURE 12. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, right parapodium, chaeti- ger 4; B, right parapodium, chaetiger 11; C, right parapodium, chaetiger 14; D, right parapodium, chaetiger 17; E, right para- podium, anterior view, chaetiger 54; F, right parapodium, anterior view, chaetiger 85. Abbreviation: DC, dorsal cirrus; DCLo, dorsal cirrus lobe; DCLa, dorsal cirrus lamellae; NeIA, neuropodial inferior acicular lobe; NePLa, neuropodial postchaetal lamellae; NeSA, neuropodial superior acicular lobe; NeVL, neuropodial ventral ligule; NoDL, notopodial dorsal ligule; NoVL, notopodial ventral ligule; VC; ventral cirrus; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AL, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074806" httpUri="https://zenodo.org/record/5074806/files/figure.png" pageId="13" pageNumber="505">Fig. 12F</figureCitation>
).
</paragraph>
<paragraph id="8BF336D0FFAEFF8A85F0FA2AFE81F5A0" blockId="13.[151,1437,1171,2025]" pageId="13" pageNumber="505">
Median ligule subconical on anterior chaetigers to posterior end, longer than neuroacicular ligule throughout (
<figureCitation id="13772A55FFAEFF8A8597FA16FE94F5A0" box="[160,292,1459,1485]" captionStart="FIGURE 12" captionStartId="15.[151,250,1051,1075]" captionTargetBox="[156,1430,181,1027]" captionTargetId="figure-33@15.[156,1430,181,1027]" captionTargetPageId="15" captionText="FIGURE 12. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, right parapodium, chaeti- ger 4; B, right parapodium, chaetiger 11; C, right parapodium, chaetiger 14; D, right parapodium, chaetiger 17; E, right para- podium, anterior view, chaetiger 54; F, right parapodium, anterior view, chaetiger 85. Abbreviation: DC, dorsal cirrus; DCLo, dorsal cirrus lobe; DCLa, dorsal cirrus lamellae; NeIA, neuropodial inferior acicular lobe; NePLa, neuropodial postchaetal lamellae; NeSA, neuropodial superior acicular lobe; NeVL, neuropodial ventral ligule; NoDL, notopodial dorsal ligule; NoVL, notopodial ventral ligule; VC; ventral cirrus; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AL, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074806" httpUri="https://zenodo.org/record/5074806/files/figure.png" pageId="13" pageNumber="505">Fig. 12CF</figureCitation>
).
</paragraph>
<paragraph id="8BF336D0FFAEFF8A85F0FA72FED0F6EC" blockId="13.[151,1437,1171,2025]" pageId="13" pageNumber="505">
Neuroacicular ligule with predominant inferior lobe. Neuropodial postchaetal lobe present throughout, rounded on anterior chaetigers, lamellae present from mid-body to posterior chaetigers (
<figureCitation id="13772A55FFAEFF8A813FFA5EFB3DF678" box="[1032,1165,1531,1557]" captionStart="FIGURE 12" captionStartId="15.[151,250,1051,1075]" captionTargetBox="[156,1430,181,1027]" captionTargetId="figure-33@15.[156,1430,181,1027]" captionTargetPageId="15" captionText="FIGURE 12. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, right parapodium, chaeti- ger 4; B, right parapodium, chaetiger 11; C, right parapodium, chaetiger 14; D, right parapodium, chaetiger 17; E, right para- podium, anterior view, chaetiger 54; F, right parapodium, anterior view, chaetiger 85. Abbreviation: DC, dorsal cirrus; DCLo, dorsal cirrus lobe; DCLa, dorsal cirrus lamellae; NeIA, neuropodial inferior acicular lobe; NePLa, neuropodial postchaetal lamellae; NeSA, neuropodial superior acicular lobe; NeVL, neuropodial ventral ligule; NoDL, notopodial dorsal ligule; NoVL, notopodial ventral ligule; VC; ventral cirrus; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AL, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074806" httpUri="https://zenodo.org/record/5074806/files/figure.png" pageId="13" pageNumber="505">Fig. 12CF</figureCitation>
). Ventral ligule subconi- cal throughout. Ventral cirri digitiform throughout, basally attached to ventral edge of parapodia, about as long as ventral ligule; ventral cirrus lobe and lamellae present throughout, reduced progressively in size toward posterior end (
<figureCitation id="13772A55FFAEFF8A85F8F9C2FEE4F6EC" box="[207,340,1639,1665]" captionStart="FIGURE 12" captionStartId="15.[151,250,1051,1075]" captionTargetBox="[156,1430,181,1027]" captionTargetId="figure-33@15.[156,1430,181,1027]" captionTargetPageId="15" captionText="FIGURE 12. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, right parapodium, chaeti- ger 4; B, right parapodium, chaetiger 11; C, right parapodium, chaetiger 14; D, right parapodium, chaetiger 17; E, right para- podium, anterior view, chaetiger 54; F, right parapodium, anterior view, chaetiger 85. Abbreviation: DC, dorsal cirrus; DCLo, dorsal cirrus lobe; DCLa, dorsal cirrus lamellae; NeIA, neuropodial inferior acicular lobe; NePLa, neuropodial postchaetal lamellae; NeSA, neuropodial superior acicular lobe; NeVL, neuropodial ventral ligule; NoDL, notopodial dorsal ligule; NoVL, notopodial ventral ligule; VC; ventral cirrus; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AL, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074806" httpUri="https://zenodo.org/record/5074806/files/figure.png" pageId="13" pageNumber="505">Fig. 12CF</figureCitation>
).
</paragraph>
<paragraph id="8BF336D0FFAEFF8A85F0F92EFDBCF7CC" blockId="13.[151,1437,1171,2025]" pageId="13" pageNumber="505">
Notochaetal homogomph spinigers present from chaetiger 3 to 16 and from chaetiger 85 to posterior end (
<figureCitation id="13772A55FFAEFF8A8058F92EFF7AF6A4" captionStart="FIGURE 13" captionStartId="18.[151,250,970,994]" captionTargetBox="[151,1436,181,946]" captionTargetId="figure-15@18.[151,1436,181,946]" captionTargetPageId="18" captionText="FIGURE 13. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, notochaeta, homogomph spiniger, chaetiger 10; B, supra-acicular homogomph spiniger, chaetiger 10; C, supra-acicular short-bladed heterogomph fal- ciger, chaetiger 10; D, parapodial chaetae, chaetiger 80; E, parapodial chaetae, chaetiger 54; F, subacicular short-bladed het- erogomph falciger, chaetiger 10. Abbreviation: NC, natatory chaetae; SubHeS, subacicular heterogomph spiniger; SubHoS, subacicular homogomph spiniger; SupHeF, supra-acicular heterogomph falciger. Scale: AC, F, 0.01 mm; DE, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5074808" httpUri="https://zenodo.org/record/5074808/files/figure.png" pageId="13" pageNumber="505">Fig. 13A</figureCitation>
), homogomph spinigers replacing by natatory chaetae from chaetiger 17 to 84. Supra-acicular fascicle of neuropodia: homogomph spinigers and short-bladed heterogomph falcigers with serrations present from chaetiger 3 to 16 and from chaetiger 80 to posterior end (
<figureCitation id="13772A55FFAEFF8A874EF952FCB0F77C" box="[633,768,1783,1809]" captionStart="FIGURE 13" captionStartId="18.[151,250,970,994]" captionTargetBox="[151,1436,181,946]" captionTargetId="figure-15@18.[151,1436,181,946]" captionTargetPageId="18" captionText="FIGURE 13. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, notochaeta, homogomph spiniger, chaetiger 10; B, supra-acicular homogomph spiniger, chaetiger 10; C, supra-acicular short-bladed heterogomph fal- ciger, chaetiger 10; D, parapodial chaetae, chaetiger 80; E, parapodial chaetae, chaetiger 54; F, subacicular short-bladed het- erogomph falciger, chaetiger 10. Abbreviation: NC, natatory chaetae; SubHeS, subacicular heterogomph spiniger; SubHoS, subacicular homogomph spiniger; SupHeF, supra-acicular heterogomph falciger. Scale: AC, F, 0.01 mm; DE, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5074808" httpUri="https://zenodo.org/record/5074808/files/figure.png" pageId="13" pageNumber="505">Fig. 13BC</figureCitation>
), homogomph spinigers replacing by natatory chaetae from chaetiger 17 to 79 (
<figureCitation id="13772A55FFAEFF8A8440F8BEFDB1F758" box="[375,513,1819,1845]" captionStart="FIGURE 13" captionStartId="18.[151,250,970,994]" captionTargetBox="[151,1436,181,946]" captionTargetId="figure-15@18.[151,1436,181,946]" captionTargetPageId="18" captionText="FIGURE 13. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, notochaeta, homogomph spiniger, chaetiger 10; B, supra-acicular homogomph spiniger, chaetiger 10; C, supra-acicular short-bladed heterogomph fal- ciger, chaetiger 10; D, parapodial chaetae, chaetiger 80; E, parapodial chaetae, chaetiger 54; F, subacicular short-bladed het- erogomph falciger, chaetiger 10. Abbreviation: NC, natatory chaetae; SubHeS, subacicular heterogomph spiniger; SubHoS, subacicular homogomph spiniger; SupHeF, supra-acicular heterogomph falciger. Scale: AC, F, 0.01 mm; DE, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5074808" httpUri="https://zenodo.org/record/5074808/files/figure.png" pageId="13" pageNumber="505">Fig. 13DE</figureCitation>
). Subacicular fascicle of neuropodia: homogomph spinigers and natatory chaetae present from chaetiger 17 to 84 (
<figureCitation id="13772A55FFAEFF8A8727F89AFD2BF734" box="[528,667,1855,1881]" captionStart="FIGURE 13" captionStartId="18.[151,250,970,994]" captionTargetBox="[151,1436,181,946]" captionTargetId="figure-15@18.[151,1436,181,946]" captionTargetPageId="18" captionText="FIGURE 13. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, notochaeta, homogomph spiniger, chaetiger 10; B, supra-acicular homogomph spiniger, chaetiger 10; C, supra-acicular short-bladed heterogomph fal- ciger, chaetiger 10; D, parapodial chaetae, chaetiger 80; E, parapodial chaetae, chaetiger 54; F, subacicular short-bladed het- erogomph falciger, chaetiger 10. Abbreviation: NC, natatory chaetae; SubHeS, subacicular heterogomph spiniger; SubHoS, subacicular homogomph spiniger; SupHeF, supra-acicular heterogomph falciger. Scale: AC, F, 0.01 mm; DE, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5074808" httpUri="https://zenodo.org/record/5074808/files/figure.png" pageId="13" pageNumber="505">Fig. 13D, E</figureCitation>
), heterogomph spinigers present from chaetiger 80 to posterior end (
<figureCitation id="13772A55FFAEFF8A8597F8C6FEB8F710" box="[160,264,1891,1917]" captionStart="FIGURE 13" captionStartId="18.[151,250,970,994]" captionTargetBox="[151,1436,181,946]" captionTargetId="figure-15@18.[151,1436,181,946]" captionTargetPageId="18" captionText="FIGURE 13. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, notochaeta, homogomph spiniger, chaetiger 10; B, supra-acicular homogomph spiniger, chaetiger 10; C, supra-acicular short-bladed heterogomph fal- ciger, chaetiger 10; D, parapodial chaetae, chaetiger 80; E, parapodial chaetae, chaetiger 54; F, subacicular short-bladed het- erogomph falciger, chaetiger 10. Abbreviation: NC, natatory chaetae; SubHeS, subacicular heterogomph spiniger; SubHoS, subacicular homogomph spiniger; SupHeF, supra-acicular heterogomph falciger. Scale: AC, F, 0.01 mm; DE, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5074808" httpUri="https://zenodo.org/record/5074808/files/figure.png" pageId="13" pageNumber="505">Fig. 13D</figureCitation>
), short-bladed heterogomph falcigers with serrations present from chaetiger 3 to 16 and from chaetiger 80 to posterior end (
<figureCitation id="13772A55FFAEFF8A844EF822FE4FF7CC" box="[377,511,1927,1953]" captionStart="FIGURE 13" captionStartId="18.[151,250,970,994]" captionTargetBox="[151,1436,181,946]" captionTargetId="figure-15@18.[151,1436,181,946]" captionTargetPageId="18" captionText="FIGURE 13. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, notochaeta, homogomph spiniger, chaetiger 10; B, supra-acicular homogomph spiniger, chaetiger 10; C, supra-acicular short-bladed heterogomph fal- ciger, chaetiger 10; D, parapodial chaetae, chaetiger 80; E, parapodial chaetae, chaetiger 54; F, subacicular short-bladed het- erogomph falciger, chaetiger 10. Abbreviation: NC, natatory chaetae; SubHeS, subacicular heterogomph spiniger; SubHoS, subacicular homogomph spiniger; SupHeF, supra-acicular heterogomph falciger. Scale: AC, F, 0.01 mm; DE, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5074808" httpUri="https://zenodo.org/record/5074808/files/figure.png" pageId="13" pageNumber="505">Fig. 13DF</figureCitation>
).
</paragraph>
<paragraph id="8BF336D0FFAEFF8985F0F80EFDFCF0B8" blockId="13.[151,1437,1171,2025]" lastBlockId="14.[151,1436,151,213]" lastPageId="14" lastPageNumber="506" pageId="13" pageNumber="505">
<emphasis id="B938EAC2FFAEFF8A85F0F80EFE8FF7A8" bold="true" box="[199,319,1963,1989]" pageId="13" pageNumber="505">Variation.</emphasis>
Male epitoke (specimen NMNS 8383-34 (2)) has digitiform dorsal cirri commenced on chaetiger 7. Dorsal cirrus lobe has not expanded to lamellae on any chaetiger. Neuropodial postchaetal lobe has not broadly expanded and always shorter than median ligule. Parapodial chaetae have not been replaced by natatory chaetae in the natatory body region (
<figureCitation id="13772A55FFADFF89848EFF1EFDF0F0B8" box="[441,576,187,213]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[241,1346,181,1843]" captionTargetId="figure-33@19.[241,1346,181,1843]" captionTargetPageId="19" captionText="FIGURE 14. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, male: A, anterior body, dorsal view; B, paragnaths of the pharynx, frontal view; C, left parapodium, posterior view, chaetiger 3; D, left parapodium, posterior view, chaetiger 10; E, left parapodium, posterior view, chaetiger 54; F, left parapodium, posterior view, chaetiger 78. Abbrevia- tion: DC, dorsal cirrus; DCLa, dorsal cirrus lamellae; NePLa, neuropodial postchaetal lamellae; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AB, 0.5 mm; CF, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5074810" httpUri="https://zenodo.org/record/5074810/files/figure.png" pageId="14" pageNumber="506">Fig. 14AF</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF336658FFADFF8985A0F8DBFB0AF7D7" ID-DOI="http://doi.org/10.5281/zenodo.5074804" ID-Zenodo-Dep="5074804" httpUri="https://zenodo.org/record/5074804/files/figure.png" pageId="14" pageNumber="506" startId="14.[151,250,1918,1942]" targetBox="[213,1375,237,1895]" targetPageId="14">
<paragraph id="8BF336D0FFADFF8985A0F8DBFB0AF7D7" blockId="14.[151,1436,1918,1979]" pageId="14" pageNumber="506">
<emphasis id="B938EAC2FFADFF8985A0F8DBFEAFF7FB" bold="true" box="[151,287,1918,1942]" pageId="14" pageNumber="506">FIGURE 11.</emphasis>
<taxonomicName id="4C4C4D53FFADFF898411F8DAFD1DF7FA" authority="Horst, 1924" authorityName="Horst" authorityYear="1924" box="[294,685,1919,1943]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="14" pageNumber="506" phylum="Annelida" rank="species" species="trimaculata">
<emphasis id="B938EAC2FFADFF898411F8DAFD9AF7FA" box="[294,554,1919,1943]" italics="true" pageId="14" pageNumber="506">Pseudonereis trimaculata</emphasis>
<bibRefCitation id="EFDD4B21FFADFF898706F8DAFD1DF7FA" author="Horst, R." box="[561,685,1919,1943]" pageId="14" pageNumber="506" pagination="145 - 198" refId="ref11486" refString="Horst, R. (1924) Polychaeta errantia of the Siboga Expedition. Part III. Nereidae and Hesionidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, 24 (1 c), 145 - 198." type="journal article" year="1924">Horst, 1924</bibRefCitation>
</taxonomicName>
; specimen NMNS 8383-35, epitoke, female: A, whole animal; B, anterior body, dorsal view; C, paragnaths of the pharynx, ventral view. Scale: A, 5.0 mm; B, 1.0 mm; C, 0.5 mm.
</paragraph>
</caption>
<caption id="DF336658FFACFF8885A0FBBEFACFF48A" ID-DOI="http://doi.org/10.5281/zenodo.5074806" ID-Zenodo-Dep="5074806" httpUri="https://zenodo.org/record/5074806/files/figure.png" pageId="15" pageNumber="507" startId="15.[151,250,1051,1075]" targetBox="[156,1430,181,1027]" targetPageId="15">
<paragraph id="8BF336D0FFACFF8885A0FBBEFACFF48A" blockId="15.[151,1436,1051,1256]" pageId="15" pageNumber="507">
<emphasis id="B938EAC2FFACFF8885A0FBBEFE90F45E" bold="true" box="[151,288,1051,1075]" pageId="15" pageNumber="507">FIGURE 12.</emphasis>
<taxonomicName id="4C4C4D53FFACFF888410FBBEFD1EF459" authority="Horst, 1924" authorityName="Horst" authorityYear="1924" box="[295,686,1051,1076]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="507" phylum="Annelida" rank="species" species="trimaculata">
<emphasis id="B938EAC2FFACFF888410FBBEFD9CF45E" box="[295,556,1051,1075]" italics="true" pageId="15" pageNumber="507">Pseudonereis trimaculata</emphasis>
<bibRefCitation id="EFDD4B21FFACFF888704FBB9FD1EF459" author="Horst, R." box="[563,686,1052,1076]" pageId="15" pageNumber="507" pagination="145 - 198" refId="ref11486" refString="Horst, R. (1924) Polychaeta errantia of the Siboga Expedition. Part III. Nereidae and Hesionidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, 24 (1 c), 145 - 198." type="journal article" year="1924">Horst, 1924</bibRefCitation>
</taxonomicName>
; specimen NMNS 8383-35, epitoke, female: A, right parapodium, chaeti- ger 4; B, right parapodium, chaetiger 11; C, right parapodium, chaetiger 14; D, right parapodium, chaetiger 17; E, right para- podium, anterior view, chaetiger 54; F, right parapodium, anterior view, chaetiger 85. Abbreviation: DC, dorsal cirrus; DCLo, dorsal cirrus lobe; DCLa, dorsal cirrus lamellae; NeIA, neuropodial inferior acicular lobe; NePLa, neuropodial postchaetal lamellae; NeSA, neuropodial superior acicular lobe; NeVL, neuropodial ventral ligule; NoDL, notopodial dorsal ligule; NoVL, notopodial ventral ligule; VC; ventral cirrus; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AL, 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C356655BFFACFF8885F0FAB0FBFBF55D" box="[199,1099,1301,1328]" pageId="15" pageNumber="507" type="distribution">
<paragraph id="8BF336D0FFACFF8885F0FAB0FBFBF55D" blockId="15.[151,1437,1301,1904]" box="[199,1099,1301,1328]" pageId="15" pageNumber="507">
<emphasis id="B938EAC2FFACFF8885F0FAB0FED1F542" bold="true" box="[199,353,1301,1327]" pageId="15" pageNumber="507">Distribution.</emphasis>
Geser,
<collectingCountry id="F35B7640FFACFF888481FAB3FD96F55D" box="[438,550,1302,1328]" name="Indonesia" pageId="15" pageNumber="507">Indonesia</collectingCountry>
, Western Australia, eastern and southern
<collectingCountry id="F35B7640FFACFF8886C5FAB3FBF8F55D" box="[1010,1096,1302,1328]" name="Taiwan" pageId="15" pageNumber="507">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C356655BFFACFF8885F0FA9FFAD4F71D" pageId="15" pageNumber="507" type="discussion">
<paragraph id="8BF336D0FFACFF8885F0FA9FFD84F641" blockId="15.[151,1437,1301,1904]" pageId="15" pageNumber="507">
<emphasis id="B938EAC2FFACFF8885F0FA9FFE8BF539" bold="true" box="[199,315,1338,1364]" pageId="15" pageNumber="507">Remarks.</emphasis>
The present species is similar to
<taxonomicName id="4C4C4D53FFACFF88878EFA9EFCE8F539" authorityName="Horst" authorityYear="1924" box="[697,856,1338,1364]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="507" phylum="Annelida" rank="species" species="trimaculata">
<emphasis id="B938EAC2FFACFF88878EFA9EFCE8F539" box="[697,856,1338,1364]" italics="true" pageId="15" pageNumber="507">P. trimaculata</emphasis>
</taxonomicName>
in term of paragnath patterns of the pharynx, morphology of parapodia, presence of glands on all three ligules of in most of chaetigers, and chaetal patterns (
<bibRefCitation id="EFDD4B21FFACFF888072FAFBFEF2F5F1" author="Bakken, T." pageId="15" pageNumber="507" pagination="145 - 176" refId="ref11104" refString="Bakken, T. (2007) Revision of Pseudonereis (Polychaeta, Nereididae). Zoological Journal of the Linnean Society, 150 (1), 145 - 176. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00289. x" type="journal article" year="2007">Bakken 2007: 167168</bibRefCitation>
, fig. 13AE;
<figureCitation id="13772A55FFACFF8884EDFA27FDE5F5F1" box="[474,597,1410,1436]" captionStart="FIGURE 8" captionStartId="11.[151,250,1041,1065]" captionTargetBox="[151,1437,199,1018]" captionTargetId="figure-33@11.[151,1437,199,1018]" captionTargetPageId="11" captionText="FIGURE 8. Pseudonereis trimaculata Horst, 1924; A, specimen NMNS 8383-29, BC, specimen NMNS 8383-26, D, speci- men NMNS 8383-33, EF, specimen NMNS 8383-27:A, anterior body, dorsal view; B, paragnaths of the pharynx, dorsal view; C, paragnaths of the pharynx, frontal view; D, anterior body, dorsal view; E, paragnaths of the pharynx, dorsal view; F, parag- naths of the pharynx, ventral view. Scale: A, D, 1.0 mm; BC, EF, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5074796" httpUri="https://zenodo.org/record/5074796/files/figure.png" pageId="15" pageNumber="507">Figs 8BC</figureCitation>
,
<figureCitation id="13772A55FFACFF888756FA27FD11F5F1" box="[609,673,1410,1436]" captionStart="FIGURE 9" captionStartId="12.[151,250,1056,1080]" captionTargetBox="[154,1433,182,1032]" captionTargetId="figure-15@12.[154,1434,181,1032]" captionTargetPageId="12" captionText="FIGURE 9. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-29: A, right parapodium, anterior view, chaetiger 11; B, right parapodium, posterior view, chaetiger 11; C, right parapodium, chaetiger 35; D, right parapodium, chaetiger 50; E, right parapodium, chaetiger 60; F, right parapodium, chaetiger 76. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior lobe. Scale: AF, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074800" httpUri="https://zenodo.org/record/5074800/files/figure.png" pageId="15" pageNumber="507">9AF</figureCitation>
,
<figureCitation id="13772A55FFACFF88879BFA27FD4EF5F1" box="[684,766,1410,1436]" captionStart="FIGURE 10" captionStartId="13.[151,250,992,1016]" captionTargetBox="[151,1436,182,969]" captionTargetId="figure-33@13.[151,1436,181,969]" captionTargetPageId="13" captionText="FIGURE 10. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-29:A, notochaeta, homogomph spiniger, chaetiger 18; B, supra-acicular homogomph spiniger, chaetiger 45; C, supra-acicular short-bladed heterogomph falciger, chaetiger 45; D, subacicular heterogomph spiniger, chaetiger 45; E, subacicular short-bladed heterogomph falciger, chaetiger 45. Scale: AE, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.5074802" httpUri="https://zenodo.org/record/5074802/files/figure.png" pageId="15" pageNumber="507">10AE</figureCitation>
,
<tableCitation id="C6CE036BFFACFF88863EFA27FCEBF5F1" box="[777,859,1410,1436]" captionStart="TABLE 1" captionStartId="16.[151,239,500,526]" captionText="TABLE 1. Key morphological characters of all known Pseudonereis to the world. Data in the table is largely based on Bakken (2007) with additional information from new described, new combined and resurrected species since 2007 by Glasby et al. (2013), Conde-Vela (2018), Kara et al. (2018) and the present study. Abbreviation: HeF, heterogomph falciger; HeS, heterogomph, spiniger; HoF, homogomph falciger; HoS, homogomph spiniger; LPTC, chaetigers reached by longest tentacular cirri; NTJ, number of teeth on the jaws. na = not available. * Conde-Vela (personal communicate)." pageId="15" pageNumber="507">Table 1</tableCitation>
). However, minor variations are noted between the present specimens and
<taxonomicName id="4C4C4D53FFACFF8884ADFA02FD88F5AD" authorityName="Horst" authorityYear="1924" box="[410,568,1446,1472]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="507" phylum="Annelida" rank="species" species="trimaculata">
<emphasis id="B938EAC2FFACFF8884ADFA02FD88F5AD" box="[410,568,1446,1472]" italics="true" pageId="15" pageNumber="507">P. trimaculata</emphasis>
</taxonomicName>
: 1) the present specimens have a range of 1 to 3 conical paragnaths in Area V of the pharynx (versus 3); and 2) no blacks spots on parapodia in fresh specimens of the present study (versus pres- ence of black spots).
<bibRefCitation id="EFDD4B21FFACFF8884B5FA4BFD98F665" author="Bakken, T." box="[386,552,1518,1544]" pageId="15" pageNumber="507" pagination="145 - 176" refId="ref11104" refString="Bakken, T. (2007) Revision of Pseudonereis (Polychaeta, Nereididae). Zoological Journal of the Linnean Society, 150 (1), 145 - 176. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00289. x" type="journal article" year="2007">Bakken (2007)</bibRefCitation>
stated that those spots are glands commonly seen in parapodia of nereidids, but now vaguely visible in the
<typeStatus id="54F78872FFACFF8884F1F9B7FD80F641" box="[454,560,1554,1580]" pageId="15" pageNumber="507" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="8BF336D0FFACFF8885F0F993FAD4F71D" blockId="15.[151,1437,1301,1904]" pageId="15" pageNumber="507">
Epitokes of
<taxonomicName id="4C4C4D53FFACFF888479F992FE5DF63D" authorityName="Horst" authorityYear="1924" box="[334,493,1590,1616]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="507" phylum="Annelida" rank="species" species="trimaculata">
<emphasis id="B938EAC2FFACFF888479F992FE5DF63D" box="[334,493,1590,1616]" italics="true" pageId="15" pageNumber="507">P. trimaculata</emphasis>
</taxonomicName>
are described for the first time in the present study. Different sexes show significant differences in morphological modifications. The female epitoke has a greater number of chaetigers and greater degree of morphological modifications on these chaetigers than that of the male epitoke. These morphological differences include that the female epitoke has: 1) about 90% (versus about 50%) of chaetigers with at least some degree of morphological modifications; 2) a dorsal small lobe on notopodial ligules and neuropodial ligule on modi- fied chaetigers (versus absent); 3) well-developed of dorsal cirrus lamellae, neuropodial postchaetal lamellae, and neuropodial ventral cirrus lobe and lamellae on the corresponding body segments (versus all moderate developed); 4) natatory chaetae on natatory body region (versus absent); and 5) different arrangement of notochaetae and neu- rochaetae on natatory body region than that of atokous individuals (versus similar arrangement) (
<figureCitation id="13772A55FFACFF8881E1F8F3FAE7F71D" box="[1238,1367,1878,1904]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStart-3="FIGURE 14" captionStartId-0="14.[151,250,1918,1942]" captionStartId-1="15.[151,250,1051,1075]" captionStartId-2="18.[151,250,970,994]" captionStartId-3="19.[151,250,1866,1890]" captionTargetBox-0="[213,1375,237,1895]" captionTargetBox-1="[156,1430,181,1027]" captionTargetBox-2="[151,1436,181,946]" captionTargetBox-3="[241,1346,181,1843]" captionTargetId-0="figure-38@14.[212,1375,237,1895]" captionTargetId-1="figure-33@15.[156,1430,181,1027]" captionTargetId-2="figure-15@18.[151,1436,181,946]" captionTargetId-3="figure-33@19.[241,1346,181,1843]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="18" captionTargetPageId-3="19" captionText-0="FIGURE 11. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, whole animal; B, anterior body, dorsal view; C, paragnaths of the pharynx, ventral view. Scale: A, 5.0 mm; B, 1.0 mm; C, 0.5 mm." captionText-1="FIGURE 12. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, right parapodium, chaeti- ger 4; B, right parapodium, chaetiger 11; C, right parapodium, chaetiger 14; D, right parapodium, chaetiger 17; E, right para- podium, anterior view, chaetiger 54; F, right parapodium, anterior view, chaetiger 85. Abbreviation: DC, dorsal cirrus; DCLo, dorsal cirrus lobe; DCLa, dorsal cirrus lamellae; NeIA, neuropodial inferior acicular lobe; NePLa, neuropodial postchaetal lamellae; NeSA, neuropodial superior acicular lobe; NeVL, neuropodial ventral ligule; NoDL, notopodial dorsal ligule; NoVL, notopodial ventral ligule; VC; ventral cirrus; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AL, 0.2 mm." captionText-2="FIGURE 13. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, female: A, notochaeta, homogomph spiniger, chaetiger 10; B, supra-acicular homogomph spiniger, chaetiger 10; C, supra-acicular short-bladed heterogomph fal- ciger, chaetiger 10; D, parapodial chaetae, chaetiger 80; E, parapodial chaetae, chaetiger 54; F, subacicular short-bladed het- erogomph falciger, chaetiger 10. Abbreviation: NC, natatory chaetae; SubHeS, subacicular heterogomph spiniger; SubHoS, subacicular homogomph spiniger; SupHeF, supra-acicular heterogomph falciger. Scale: AC, F, 0.01 mm; DE, 0.1 mm." captionText-3="FIGURE 14. Pseudonereis trimaculata Horst, 1924; specimen NMNS 8383-35, epitoke, male: A, anterior body, dorsal view; B, paragnaths of the pharynx, frontal view; C, left parapodium, posterior view, chaetiger 3; D, left parapodium, posterior view, chaetiger 10; E, left parapodium, posterior view, chaetiger 54; F, left parapodium, posterior view, chaetiger 78. Abbrevia- tion: DC, dorsal cirrus; DCLa, dorsal cirrus lamellae; NePLa, neuropodial postchaetal lamellae; VCLa, ventral cirrus lamellae; VCLo, ventral cirrus lobe. Scale: AB, 0.5 mm; CF, 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5074804" figureDoi-1="http://doi.org/10.5281/zenodo.5074806" figureDoi-2="http://doi.org/10.5281/zenodo.5074808" figureDoi-3="http://doi.org/10.5281/zenodo.5074810" httpUri-0="https://zenodo.org/record/5074804/files/figure.png" httpUri-1="https://zenodo.org/record/5074806/files/figure.png" httpUri-2="https://zenodo.org/record/5074808/files/figure.png" httpUri-3="https://zenodo.org/record/5074810/files/figure.png" pageId="15" pageNumber="507">Figs 1114</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>