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<document id="2569743EF761E6A20CA2556853FE7B18" ID-DOI="10.11646/zootaxa.3637.1.3" ID-GBIF-Dataset="4cb7cf94-c063-40dc-9488-e1e76d440214" ID-ISSN="1175-5326" ID-Zenodo-Dep="283584" ID-ZooBank="BA78F913-9C29-4B80-A328-229529EB1961" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460590524900" checkinUser="plazi" docAuthor="Bilton, David T." docDate="2013" docId="03CC87EEFFFDFFC00995FB74FE88AA66" docLanguage="en" docName="zt03637p038.pdf" docOrigin="Zootaxa 3637 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydraena lotti Bilton, 2013, sp. nov." docType="treatment" docVersion="5" lastPageNumber="37" masterDocId="FFF5FF96FFFCFFC80902FFB8FFCAA85A" masterDocTitle="Hydraena lotti sp. nov., a new member of the “ Haenydra ” lineage from the Peloponnese (Greece), with additional records of Hydraena species in the region (Coleoptera, Hydraenidae)" masterLastPageNumber="38" masterPageNumber="29" pageNumber="30" updateTime="1698325792539" updateUser="plazi">
<mods:mods id="8482CA4187936C76BECC3A9D00F75712" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="05FEEDBDB62E8E657A166586851EA89E">
<mods:title id="F797A6C961AEF98EBD9FF411C910F84C">Hydraena lotti sp. nov., a new member of the “ Haenydra ” lineage from the Peloponnese (Greece), with additional records of Hydraena species in the region (Coleoptera, Hydraenidae)</mods:title>
</mods:titleInfo>
<mods:name id="B1E788465F14E1E798ADAAFFE7119EC7" type="personal">
<mods:role id="3C6B11FBF92BE23B2AD93611DE3272B5">
<mods:roleTerm id="4375DE4C0A926EF514DB3E0763FC8801">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C2C7330F27DB7A8E8C3D5329326F6B96">Bilton, David T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C095ED670A0A24C2115D0F4D5F50E4C6">text</mods:typeOfResource>
<mods:relatedItem id="561B289E4D2B8FED7C8A7E8321256BE8" type="host">
<mods:titleInfo id="0C183062A1ED42393424D1E2E55633E8">
<mods:title id="42CB46F1C465CA73E30B9C29C247DAD2">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F64C8802E55DBAB4F3C75C171F76938C">
<mods:date id="B157B989B6EF6D3B6A32E4C8E4DDE155">2013</mods:date>
<mods:detail id="E9ED2B95F83B721572904866A230CBBE" type="volume">
<mods:number id="5454E0F02F76F686225645FFE5B5C81A">3637</mods:number>
</mods:detail>
<mods:detail id="F2671A423CB4F13F741FDF5F69C773B7" type="issue">
<mods:number id="C63605D0931160D17D78A827D74AA05B">1</mods:number>
</mods:detail>
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<mods:start id="33F43A49E55105C0112282A7AFE048FE">29</mods:start>
<mods:end id="51DFF22903E97141587598945A5CDD97">38</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="317B828E7C5263FCCB5667C872CD20CB">journal article</mods:classification>
<mods:identifier id="8215FB9E2087FED4C3D25DD7569631DF" type="DOI">10.11646/zootaxa.3637.1.3</mods:identifier>
<mods:identifier id="D4F59DC709B248EA879B71138A01F6C6" type="GBIF-Dataset">4cb7cf94-c063-40dc-9488-e1e76d440214</mods:identifier>
<mods:identifier id="476508A73DB9B15BB86CF6C4CE6BAFA8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="75EC08341D851F4A310FB528C4BC09B2" type="Zenodo-Dep">283584</mods:identifier>
<mods:identifier id="930FD458AEE4D9D0351DB8AD33A3B305" type="ZooBank">BA78F913-9C29-4B80-A328-229529EB1961</mods:identifier>
</mods:mods>
<treatment id="03CC87EEFFFDFFC00995FB74FE88AA66" ID-DOI="http://doi.org/10.5281/zenodo.6151267" ID-GBIF-Taxon="127663737" ID-Zenodo-Dep="6151267" LSID="urn:lsid:plazi:treatment:03CC87EEFFFDFFC00995FB74FE88AA66" httpUri="http://treatment.plazi.org/id/03CC87EEFFFDFFC00995FB74FE88AA66" lastPageId="8" lastPageNumber="37" pageId="1" pageNumber="30">
<subSubSection id="C37F6573FFFDFFC90995FB74FEC1AD5C" pageId="1" pageNumber="30" type="nomenclature">
<paragraph id="8BDA36F8FFFDFFC90995FB74FE61ACBF" blockId="1.[151,427,1227,1286]" box="[151,427,1227,1254]" pageId="1" pageNumber="30">
<heading id="D0928194FFFDFFC90995FB74FE61ACBF" bold="true" box="[151,427,1227,1254]" fontSize="11" level="1" pageId="1" pageNumber="30" reason="1">
<emphasis id="B911EAEAFFFDFFC90995FB74FE61ACBF" bold="true" box="[151,427,1227,1254]" pageId="1" pageNumber="30">
<taxonomicName id="4C654D7BFFFDFFC90995FB74FE8CACBC" ID-CoL="3N2WC" box="[151,326,1228,1254]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="30" phylum="Arthropoda" rank="species" species="lotti" status="sp. nov.">
<emphasis id="B911EAEAFFFDFFC90995FB74FE8CACBC" bold="true" box="[151,326,1228,1254]" italics="true" pageId="1" pageNumber="30">Hydraena lotti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2225791FFFDFFC9084FFB73FE61ACBF" box="[333,427,1227,1253]" pageId="1" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA36F8FFFDFFC90995FB56FEC1AD5C" blockId="1.[151,427,1227,1286]" box="[151,267,1262,1286]" pageId="1" pageNumber="30">
(
<figureCitation id="135E2A7DFFFDFFC9099DFB56FECBAD5C" box="[159,257,1262,1286]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1926,1949]" captionStartId-1="3.[151,250,1173,1196]" captionStartId-2="4.[151,250,1465,1488]" captionStartId-3="5.[151,250,1909,1932]" captionStartId-4="6.[151,250,1960,1983]" captionTargetBox-1="[164,1435,200,1152]" captionTargetBox-2="[162,1434,486,1441]" captionTargetBox-3="[164,1435,454,1888]" captionTargetBox-4="[318,1275,193,1894]" captionTargetId-1="figure@3.[151,1436,193,1152]" captionTargetId-2="figure@4.[151,1436,481,1444]" captionTargetId-3="figure@5.[151,1436,445,1888]" captionTargetId-4="figure@6.[318,1275,193,1894]" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1. Habitus of Hydraena lotti sp. nov., male holotype. Scale bar = 0. 5 mm." captionText-1="FIGURE 2. Hydraena lotti sp. nov. A) male elytral apex; B) female elytral apex; C) female tergite X; D) gonocoxite." captionText-2="FIGURE 3. Hydraena lotti sp. nov. A) aedeagus, lateral view; B) aedeagus dorsal view; C) aedeagus dorsolateral view; D) spermatheca." captionText-3="FIGURE 4. Hydraena lotti sp. nov. A) – B) male metatibia; C) – D) female metatibia; E) – F) male mesotibia." captionText-4="FIGURE 5. Type locality of Hydraena lotti sp. nov. Greece, Peloponnese, Taygetos Mountains, Lagadha stream above Lagadha pass, 1,100 m. Photo D. T. Bilton." httpUri-1="https://zenodo.org/record/283585/files/figure.png" httpUri-2="https://zenodo.org/record/283586/files/figure.png" httpUri-3="https://zenodo.org/record/283587/files/figure.png" httpUri-4="https://zenodo.org/record/283588/files/figure.png" pageId="1" pageNumber="30">Figs 1–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37F6573FFFDFFC90995FA8FFE60AEA4" pageId="1" pageNumber="30" type="materials_examined">
<paragraph id="8BDA36F8FFFDFFC90995FA8FFCABAD28" blockId="1.[151,1436,1334,2006]" pageId="1" pageNumber="30">
<emphasis id="B911EAEAFFFDFFC90995FA8FFEFBAD15" bold="true" box="[151,305,1334,1359]" pageId="1" pageNumber="30">
<typeStatus id="54DE885AFFFDFFC90995FA8FFF1AAD15" box="[151,208,1335,1359]" pageId="1" pageNumber="30">Type</typeStatus>
locality
</emphasis>
.
<emphasis id="B911EAEAFFFDFFC90846FA8EFE52AD15" bold="true" box="[324,408,1334,1359]" pageId="1" pageNumber="30">
<collectingCountry id="F3727668FFFDFFC90846FA8EFE52AD15" box="[324,408,1334,1359]" name="Greece" pageId="1" pageNumber="30">Greece</collectingCountry>
</emphasis>
, Pelopennese, Taygetos Mountains, Lagadha stream above Lagadha Pass on EO82 road between Kalamata and Sparti,
<quantity id="4C9D9B1DFFFDFFC908ECFAE2FD81AD28" box="[494,587,1370,1394]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="30" unit="m" value="1100.0">1,100 m</quantity>
, D.T. Bilton leg. (
<figureCitation id="135E2A7DFFFDFFC90A17FAE2FC99AD28" box="[789,851,1370,1394]" captionStart="FIGURE 5" captionStartId="6.[151,250,1960,1983]" captionTargetBox="[318,1275,193,1894]" captionTargetId="figure@6.[318,1275,193,1894]" captionTargetPageId="6" captionText="FIGURE 5. Type locality of Hydraena lotti sp. nov. Greece, Peloponnese, Taygetos Mountains, Lagadha stream above Lagadha pass, 1,100 m. Photo D. T. Bilton." httpUri="https://zenodo.org/record/283588/files/figure.png" pageId="1" pageNumber="30">Fig 5</figureCitation>
).
</paragraph>
<paragraph id="8BDA36F8FFFDFFC909C5FAC7FF33AD84" blockId="1.[151,1436,1334,2006]" pageId="1" pageNumber="30">
<emphasis id="B911EAEAFFFDFFC909C5FAC7FEA1ADCD" bold="true" box="[199,363,1406,1431]" pageId="1" pageNumber="30">
<typeStatus id="54DE885AFFFDFFC909C5FAC7FECAADCD" box="[199,256,1407,1431]" pageId="1" pageNumber="30">Type</typeStatus>
material
</emphasis>
.
<emphasis id="B911EAEAFFFDFFC90879FAC7FE22ADCD" bold="true" box="[379,488,1407,1431]" pageId="1" pageNumber="30">
<typeStatus id="54DE885AFFFDFFC90879FAC7FE22ADCD" box="[379,488,1407,1431]" pageId="1" pageNumber="30" type="holotype">Holotype</typeStatus>
</emphasis>
(male): “
<date id="FFDB1038FFFDFFC90B52FAC6FD07ADCC" box="[592,717,1406,1430]" pageId="1" pageNumber="30" value="2012-08-03">3/viii/2012</date>
GR Peloponnese Taygetos Mts. Lagadha stream above Lagadha Pass on EO82 road
<quantity id="4C9D9B1DFFFDFFC90873FA1AFE07ADE0" box="[369,461,1442,1466]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="30" unit="m" value="1100.0">1,100 m</quantity>
D T Bilton leg.” (genitalia extracted and mounted on same card) and red
<typeStatus id="54DE885AFFFDFFC90DFFFA1AFA95ADE0" box="[1277,1375,1442,1466]" pageId="1" pageNumber="30" type="holotype">holotype</typeStatus>
label (NMW).
</paragraph>
<paragraph id="8BDA36F8FFFDFFC909C5FA53FE60AEA4" blockId="1.[151,1436,1334,2006]" pageId="1" pageNumber="30">
<emphasis id="B911EAEAFFFDFFC909C5FA53FEF5AE59" bold="true" box="[199,319,1515,1539]" pageId="1" pageNumber="30">
<typeStatus id="54DE885AFFFDFFC909C5FA53FEF5AE59" box="[199,319,1515,1539]" pageId="1" pageNumber="30" type="paratype">Paratypes</typeStatus>
</emphasis>
(97):
<emphasis id="B911EAEAFFFDFFC9088EFA52FE2AAE59" bold="true" box="[396,480,1514,1539]" pageId="1" pageNumber="30">
<collectingCountry id="F3727668FFFDFFC9088EFA52FE2AAE59" box="[396,480,1514,1539]" name="Greece" pageId="1" pageNumber="30">Greece</collectingCountry>
</emphasis>
: 22 3, 11 Ƥ “
<date id="FFDB1038FFFDFFC90B96FA52FCDBAE58" box="[660,785,1514,1538]" pageId="1" pageNumber="30" value="2012-08-03">3/viii/2012</date>
GR Peloponnese Taygetos Mts. Lagadha stream above Lagadha Pass on EO82 road
<quantity id="4C9D9B1DFFFDFFC908E1F9B6FD8BAE7C" box="[483,577,1550,1574]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="30" unit="m" value="1100.0">1,100 m</quantity>
D T Bilton leg.” (
<collectingCountry id="F3727668FFFDFFC90A16F9B6FC98AE7C" box="[788,850,1550,1574]" name="Central African Republic" pageId="1" pageNumber="30">CAR</collectingCountry>
, CDTB, CFA, CJF, IBE, NMPC, NMW); 2 3, 4 Ƥ “
<date id="FFDB1038FFFDFFC909A3F98AFED7AE10" box="[161,285,1586,1610]" pageId="1" pageNumber="30" value="2012-06-29">29/vi/2012</date>
GR Peloponnese Taygetos Mts. Lagadha stream above Lagadha Pass on EO82 road
<quantity id="4C9D9B1DFFFDFFC90C00F98AFAABAE10" box="[1282,1377,1586,1610]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="30" unit="m" value="1100.0">1,100 m</quantity>
D T Bilton leg. (CDTB); 20 3, 16 Ƥ “
<date id="FFDB1038FFFDFFC90B12F9EEFD44AE34" box="[528,654,1622,1646]" pageId="1" pageNumber="30" value="2012-08-04">4/viii/2012</date>
GR Peloponnese Taygetos Mts. headwater of Dzirorema stream ca.
<quantity id="4C9D9B1DFFFDFFC90C8CF9EEFF71AEC8" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="30" unit="km" value="2.0">2 km</quantity>
S of Neochori
<quantity id="4C9D9B1DFFFDFFC9086EF9C2FE00AEC8" box="[364,458,1658,1682]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="1" pageNumber="30" unit="m" value="1150.0">1,150 m</quantity>
D.T. Bilton leg.” (CTV, MCZ, NHM, NMW, OUMNH, ZSM); 3 3, 4 Ƥ “
<date id="FFDB1038FFFDFFC90C1CF9C2FA56AEC8" box="[1310,1436,1658,1682]" pageId="1" pageNumber="30" value="2012-08-04">4/viii/2012</date>
GR Peloponnese Taygetos Mts. Sitsovitko stream above Machalas
<quantity id="4C9D9B1DFFFDFFC90A81F926FC14AEEC" box="[899,990,1694,1718]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="30" unit="m" value="1100.0">1,100 m</quantity>
D.T. Bilton leg.” (CDTB); 7 3, 8 Ƥ “
<date id="FFDB1038FFFDFFC90C86F926FF36AE80" pageId="1" pageNumber="30" value="2012-08-04">4/ viii/2012</date>
GR Peloponnese Taygetos Mts. Boursos stream above Alaghonia
<quantity id="4C9D9B1DFFFDFFC90AE5F97AFB88AE80" box="[999,1090,1730,1754]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="30" unit="m" value="1300.0">1,300 m</quantity>
D.T. Bilton leg.” (CDTB). All with red
<typeStatus id="54DE885AFFFDFFC909F8F95FFE90AEA5" box="[250,346,1767,1791]" pageId="1" pageNumber="30" type="paratype">paratype</typeStatus>
labels.
</paragraph>
</subSubSection>
<subSubSection id="C37F6573FFFDFFCF09C5F8B2FBDFA93E" lastPageId="7" lastPageNumber="36" pageId="1" pageNumber="30" type="description">
<paragraph id="8BDA36F8FFFDFFC909C5F8B2FEB3AFD4" blockId="1.[151,1436,1334,2006]" pageId="1" pageNumber="30">
<emphasis id="B911EAEAFFFDFFC909C5F8B2FE9BAF79" bold="true" box="[199,337,1802,1827]" pageId="1" pageNumber="30">Description</emphasis>
. Size:
<typeStatus id="54DE885AFFFDFFC908A4F8B2FDD9AF78" box="[422,531,1802,1826]" pageId="1" pageNumber="30" type="holotype">Holotype</typeStatus>
: body length (to elytral apices)
<quantity id="4C9D9B1DFFFDFFC90A89F8B2FC2DAF78" box="[907,999,1802,1826]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="30" unit="mm" value="2.2">2.2 mm</quantity>
; max. width (elytra)
<quantity id="4C9D9B1DFFFDFFC90DE3F8B2FA81AF78" box="[1249,1355,1802,1826]" metricMagnitude="-4" metricUnit="m" metricValue="7.7" pageId="1" pageNumber="30" unit="mm" value="0.77">0.77 mm</quantity>
. Head, pronotum and elytra black to blackish brown, somewhat lighter at all margins. Venter, lateral hypomera and pseudepipleura paler, with darker mentum. Legs reddish-brown; tarsi and maxilliary palps paler, especially towards their apices (
<figureCitation id="135E2A7DFFFDFFC90824F8CEFEA2AFD4" box="[294,360,1910,1934]" captionStart="FIGURE 1" captionStartId="2.[151,250,1926,1949]" captionText="FIGURE 1. Habitus of Hydraena lotti sp. nov., male holotype. Scale bar = 0. 5 mm." pageId="1" pageNumber="30">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BDA36F8FFFDFFCA09C5F822FE38A995" blockId="1.[151,1436,1334,2006]" lastBlockId="2.[151,1436,151,463]" lastPageId="2" lastPageNumber="31" pageId="1" pageNumber="30">Head: Labrum with deep V-shaped sutural notch. Lobes each side of sutural notch shallowly concave, with reflexed margins. Concave areas of lobes with fine microsculpture and shallow punctures supplied with short reflexed setae, these setae being particularly evident close to outer margins. Central portion of labrum with a smooth, shiny triangular patch at clypeal margin. Clypeus with narrow, smooth and shiny anterior margin adjacent to this area. Remainder of clypeus with dense small, shallow punctures, and scattered deeper, slightly larger ones, some of which are supplied with short reflexed setae. Shallow punctures becoming much denser in lateral marginal fifths of the clypeus, giving these areas a rugose appearance. Frons with punctures which are both deeper and larger than those on the clypeus. Punctures on frons spaced approximately 1–2 puncture widths apart in centre, becoming confluent at lateral margins and around eyes, giving a rugose appearance to these areas. Each puncture on frons supplied with a small central reflexed seta; setae becoming longer towards lateral margins. Frons strongly shining in centre; outer regions less so.</paragraph>
<caption id="DF1A6670FFFEFFCA0995F83EFC26AFC6" box="[151,1004,1926,1949]" pageId="2" pageNumber="31">
<paragraph id="8BDA36F8FFFEFFCA0995F83EFC26AFC6" blockId="2.[151,1004,1926,1949]" box="[151,1004,1926,1949]" pageId="2" pageNumber="31">
<emphasis id="B911EAEAFFFEFFCA0995F83EFED9AFC1" bold="true" box="[151,275,1926,1949]" pageId="2" pageNumber="31">FIGURE 1.</emphasis>
Habitus of
<taxonomicName id="4C654D7BFFFEFFCA088BF83EFDD4AFC6" box="[393,542,1926,1948]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="lotti" status="sp. nov.">
<emphasis id="B911EAEAFFFEFFCA088BF83EFDD4AFC6" box="[393,542,1926,1948]" italics="true" pageId="2" pageNumber="31">Hydraena lotti</emphasis>
</taxonomicName>
<emphasis id="B911EAEAFFFEFFCA0B26F83FFDBEAFC6" bold="true" box="[548,628,1927,1948]" pageId="2" pageNumber="31">
<taxonomicNameLabel id="A2225791FFFEFFCA0B26F83FFDBEAFC6" box="[548,628,1927,1948]" pageId="2" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype. Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="DF1A6670FFFFFFCB0995FB2DFA8BACF6" box="[151,1345,1173,1196]" httpUri="https://zenodo.org/record/283585/files/figure.png" pageId="3" pageNumber="32" targetBox="[164,1435,200,1152]" targetPageId="3">
<paragraph id="8BDA36F8FFFFFFCB0995FB2DFA8BACF6" blockId="3.[151,1345,1173,1196]" box="[151,1345,1173,1196]" pageId="3" pageNumber="32">
<emphasis id="B911EAEAFFFFFFCB0995FB2DFED9ACF1" bold="true" box="[151,275,1173,1196]" pageId="3" pageNumber="32">FIGURE 2.</emphasis>
<taxonomicName id="4C654D7BFFFFFFCB081BFB2EFE64ACF6" box="[281,430,1174,1196]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="lotti" status="sp. nov.">
<emphasis id="B911EAEAFFFFFFCB081BFB2EFE64ACF6" box="[281,430,1174,1196]" italics="true" pageId="3" pageNumber="32">Hydraena lotti</emphasis>
</taxonomicName>
<emphasis id="B911EAEAFFFFFFCB08B6FB2EFDCEACF1" bold="true" box="[436,516,1174,1195]" pageId="3" pageNumber="32">
<taxonomicNameLabel id="A2225791FFFFFFCB08B6FB2EFDCEACF1" box="[436,516,1174,1195]" pageId="3" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A) male elytral apex; B) female elytral apex; C) female tergite X; D) gonocoxite.
</paragraph>
</caption>
<paragraph id="8BDA36F8FFFFFFCB09C5FB6CFCC9AE75" blockId="3.[151,1437,1235,2015]" pageId="3" pageNumber="32">
Pronotum: Elongately hexagonal, with weakly denticulate side margins. Scattered, shallow open punctures throughout, these being 1–
<date id="FFDB1038FFFFFFCB08CBFB4FFE12AD55" box="[457,472,1271,1295]" pageId="3" pageNumber="32">2</date>
x diameter of those on the frons. Front and hind areas of pronotum with punctures approximately 0.75–1 puncture widths apart over areas approximately ¼ of pronotal length. Puncture spacing similar in a narrow longitudinal band running down the centre of the pronotum. Lateral areas of pronotum with strong longitudinal fossae, which are deeper and broader in both front and hind thirds. Punctures on pronotum either side of fossae 1–2 puncture widths apart. Punctures in fossae shallow and sparse, giving a relatively smooth appearance to these areas. Outside fossae, pronotum smooth and shiny between punctures, with traces of reticulation developing towards front angles. Pronotal punctures supplied with fine recumbent setae, which are approximately two puncture widths long. Some setae lost, especially on disc. Wings reduced, approx. ½ length of elytra in specimens examined, and with weak venation.
</paragraph>
<paragraph id="8BDA36F8FFFFFFCB09C5F983FD31AF5D" blockId="3.[151,1437,1235,2015]" pageId="3" pageNumber="32">
Elytra: Elongate, subparallel, slightly rounded at sides and widest behind middle. Lateral margins explanate, particularly in posterior half, as in other members of the
<emphasis id="B911EAEAFFFFFFCB0A08F9D8FC20AE2D" box="[778,1002,1632,1655]" italics="true" pageId="3" pageNumber="32">H. emarginata-saga</emphasis>
clade. Margins apparently smooth, but weakly serrate at higher magnifications (&gt;
<date id="FFDB1038FFFFFFCB0B79F93BFD7AAEC1" box="[635,688,1667,1691]" pageId="3" pageNumber="32" value="1960-10">60 x</date>
). Elytral apex conjointly rounded, with marked sutural notch (
<figureCitation id="135E2A7DFFFFFFCB0C6DF93BFF6CAEE5" captionStart="FIGURE 2" captionStartId="3.[151,250,1173,1196]" captionTargetBox="[164,1435,200,1152]" captionTargetId="figure@3.[151,1436,193,1152]" captionTargetPageId="3" captionText="FIGURE 2. Hydraena lotti sp. nov. A) male elytral apex; B) female elytral apex; C) female tergite X; D) gonocoxite." httpUri="https://zenodo.org/record/283585/files/figure.png" pageId="3" pageNumber="32">Fig. 2</figureCitation>
A). Elytral surface smooth and shining between the nine regular puncture rows. Each puncture of these rows with a fine, flat seta arising from a small tubercle at the front end of the puncture (
<figureCitation id="135E2A7DFFFFFFCB0D4FF973FB5FAEB9" box="[1101,1173,1739,1763]" captionStart="FIGURE 2" captionStartId="3.[151,250,1173,1196]" captionTargetBox="[164,1435,200,1152]" captionTargetId="figure@3.[151,1436,193,1152]" captionTargetPageId="3" captionText="FIGURE 2. Hydraena lotti sp. nov. A) male elytral apex; B) female elytral apex; C) female tergite X; D) gonocoxite." httpUri="https://zenodo.org/record/283585/files/figure.png" pageId="3" pageNumber="32">Fig. 2</figureCitation>
A–B). Setae reaching backwards to front margin of next puncture in the row.
</paragraph>
<paragraph id="8BDA36F8FFFFFFCC09C5F8ACFB20A9F6" blockId="3.[151,1437,1235,2015]" lastBlockId="4.[151,1436,151,428]" lastPageId="4" lastPageNumber="33" pageId="3" pageNumber="32">Venter: Front margin of mentum with a thin ventrally reflexed portion, furnished with fine, close, curved setae. Mentum finely reticulate, with a super-imposed network of raised ridges, giving a roughened appearance overall. Submentum densely covered in small shallow punctures; shining between punctures and around all margins of sclerite. Genae shining, with strong crenulated lines and weak reticulation between these at margins. Genal ridge obsolete; shining. Gular region heavily punctured and reticulate, except in centre and at hind margins. Lateral hypomeron of pronotum smooth, with shallow, confluent pits. Hypomeral carinae smooth and shining. Mesal hypomeron with dense hydrofuge pubescence. Pseudepipleura of elytra smooth and shining, concave due to reflexed dorsal margins of elytra. Entire meso- and metaventrites, and abdominal sterna covered in dense hydrofuge pubescence, with the exception of the metaventral plaques and abdominal ventrite VIII and a large adjacent semicircular area of ventrite VII. Areas with hydrofuge pubescence apparently also finely reticulate beneath these hairs. Metaventral plaques smooth and shining, without reticulation. Areas of sternites lacking hydrofuge pubescence with regular reticulation of large, open, meshes, these meshes becoming larger on ventrite VII and towards the apex of ventrite VIII. Glabrous area of ventrite VII with a row of punctures, each of which bears a long seta. Similar punctures with long setae concentrated towards the apex of ventrite VIII.</paragraph>
<caption id="DF1A6670FFF8FFCC0995FA01FED7ADB4" httpUri="https://zenodo.org/record/283586/files/figure.png" pageId="4" pageNumber="33" targetBox="[162,1434,486,1441]" targetPageId="4">
<paragraph id="8BDA36F8FFF8FFCC0995FA01FED7ADB4" blockId="4.[151,1436,1465,1518]" pageId="4" pageNumber="33">
<emphasis id="B911EAEAFFF8FFCC0995FA01FEDEAD95" bold="true" box="[151,276,1465,1488]" pageId="4" pageNumber="33">FIGURE 3.</emphasis>
<taxonomicName id="4C654D7BFFF8FFCC081FFA02FE79AD8A" box="[285,435,1466,1488]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="lotti" status="sp. nov.">
<emphasis id="B911EAEAFFF8FFCC081FFA02FE79AD8A" box="[285,435,1466,1488]" italics="true" pageId="4" pageNumber="33">Hydraena lotti</emphasis>
</taxonomicName>
<emphasis id="B911EAEAFFF8FFCC08B9FA02FDC4AD95" bold="true" box="[443,526,1466,1487]" pageId="4" pageNumber="33">
<taxonomicNameLabel id="A2225791FFF8FFCC08B9FA02FDC4AD95" box="[443,526,1466,1487]" pageId="4" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A) aedeagus, lateral view; B) aedeagus dorsal view; C) aedeagus dorsolateral view; D) spermatheca.
</paragraph>
</caption>
<paragraph id="8BDA36F8FFF8FFCC09C5F9AFFA57AE0B" blockId="4.[151,1437,1558,2014]" pageId="4" pageNumber="33">
Legs: Mesotibia with stout setae and a row of small teeth on mesal face of posterior half (
<figureCitation id="135E2A7DFFF8FFCC0DC3F9AEFACEAE74" box="[1217,1284,1558,1582]" captionStart="FIGURE 1" captionStartId="2.[151,250,1926,1949]" captionText="FIGURE 1. Habitus of Hydraena lotti sp. nov., male holotype. Scale bar = 0. 5 mm." pageId="4" pageNumber="33">Fig. 1</figureCitation>
,
<figureCitation id="135E2A7DFFF8FFCC0C11F9AEFA9FAE74" box="[1299,1365,1558,1582]" captionStart="FIGURE 4" captionStartId="5.[151,250,1909,1932]" captionTargetBox="[164,1435,454,1888]" captionTargetId="figure@5.[151,1436,445,1888]" captionTargetPageId="5" captionText="FIGURE 4. Hydraena lotti sp. nov. A) – B) male metatibia; C) – D) female metatibia; E) – F) male mesotibia." httpUri="https://zenodo.org/record/283587/files/figure.png" pageId="4" pageNumber="33">Fig. 4</figureCitation>
E–F). Metatibia with a row of long setae on mesial face of posterior half (
<figureCitation id="135E2A7DFFF8FFCC0A80F981FC0EAE0B" box="[898,964,1593,1617]" captionStart="FIGURE 1" captionStartId="2.[151,250,1926,1949]" captionText="FIGURE 1. Habitus of Hydraena lotti sp. nov., male holotype. Scale bar = 0. 5 mm." pageId="4" pageNumber="33">Fig. 1</figureCitation>
,
<figureCitation id="135E2A7DFFF8FFCC0AD3F981FBD8AE0B" box="[977,1042,1593,1617]" captionStart="FIGURE 4" captionStartId="5.[151,250,1909,1932]" captionTargetBox="[164,1435,454,1888]" captionTargetId="figure@5.[151,1436,445,1888]" captionTargetPageId="5" captionText="FIGURE 4. Hydraena lotti sp. nov. A) – B) male metatibia; C) – D) female metatibia; E) – F) male mesotibia." httpUri="https://zenodo.org/record/283587/files/figure.png" pageId="4" pageNumber="33">Fig. 4</figureCitation>
A–B). Legs otherwise unmodified.
</paragraph>
<paragraph id="8BDA36F8FFF8FFCC09C5F9E6FB13AEC3" blockId="4.[151,1437,1558,2014]" pageId="4" pageNumber="33">
Aedeagus: Main piece drawn out into long, tapering finger-like projection, of characteristic shape (
<figureCitation id="135E2A7DFFF8FFCC0C31F9E6FABDAE2C" box="[1331,1399,1630,1654]" captionStart="FIGURE 3" captionStartId="4.[151,250,1465,1488]" captionTargetBox="[162,1434,486,1441]" captionTargetId="figure@4.[151,1436,481,1444]" captionTargetPageId="4" captionText="FIGURE 3. Hydraena lotti sp. nov. A) aedeagus, lateral view; B) aedeagus dorsal view; C) aedeagus dorsolateral view; D) spermatheca." httpUri="https://zenodo.org/record/283586/files/figure.png" pageId="4" pageNumber="33">Fig. 3</figureCitation>
A). Distal lobe reflexed over the main piece, with long, thin tubular terminal appendage (
<figureCitation id="135E2A7DFFF8FFCC0D53F939FB59AEC3" box="[1105,1171,1665,1689]" captionStart="FIGURE 3" captionStartId="4.[151,250,1465,1488]" captionTargetBox="[162,1434,486,1441]" captionTargetId="figure@4.[151,1436,481,1444]" captionTargetPageId="4" captionText="FIGURE 3. Hydraena lotti sp. nov. A) aedeagus, lateral view; B) aedeagus dorsal view; C) aedeagus dorsolateral view; D) spermatheca." httpUri="https://zenodo.org/record/283586/files/figure.png" pageId="4" pageNumber="33">Fig. 3</figureCitation>
A–C).
</paragraph>
<paragraph id="8BDA36F8FFF8FFCC09C5F91EFB51AF5C" blockId="4.[151,1437,1558,2014]" pageId="4" pageNumber="33">
Female: Elytral apices bitruncate (
<figureCitation id="135E2A7DFFF8FFCC0B45F91EFD41AEE4" box="[583,651,1702,1726]" captionStart="FIGURE 2" captionStartId="3.[151,250,1173,1196]" captionTargetBox="[164,1435,200,1152]" captionTargetId="figure@3.[151,1436,193,1152]" captionTargetPageId="3" captionText="FIGURE 2. Hydraena lotti sp. nov. A) male elytral apex; B) female elytral apex; C) female tergite X; D) gonocoxite." httpUri="https://zenodo.org/record/283585/files/figure.png" pageId="4" pageNumber="33">Fig. 2</figureCitation>
B); not rounded as in males. Apices slightly raised, roof-like, in lateral view. Meso- and metatibia (
<figureCitation id="135E2A7DFFF8FFCC08CEF971FDC4AEBB" box="[460,526,1737,1761]" captionStart="FIGURE 4" captionStartId="5.[151,250,1909,1932]" captionTargetBox="[164,1435,454,1888]" captionTargetId="figure@5.[151,1436,445,1888]" captionTargetPageId="5" captionText="FIGURE 4. Hydraena lotti sp. nov. A) – B) male metatibia; C) – D) female metatibia; E) – F) male mesotibia." httpUri="https://zenodo.org/record/283587/files/figure.png" pageId="4" pageNumber="33">Fig. 4</figureCitation>
C–D) without modifications seen in males. Gonocoxite and tergite X
<figureCitation id="135E2A7DFFF8FFCC0C1AF971FA93AEBB" box="[1304,1369,1737,1761]" captionStart="FIGURE 2" captionStartId="3.[151,250,1173,1196]" captionTargetBox="[164,1435,200,1152]" captionTargetId="figure@3.[151,1436,193,1152]" captionTargetPageId="3" captionText="FIGURE 2. Hydraena lotti sp. nov. A) male elytral apex; B) female elytral apex; C) female tergite X; D) gonocoxite." httpUri="https://zenodo.org/record/283585/files/figure.png" pageId="4" pageNumber="33">Fig. 2</figureCitation>
C–D; spermatheca
<figureCitation id="135E2A7DFFF8FFCC0828F956FEA1AF5C" box="[298,363,1774,1798]" captionStart="FIGURE 3" captionStartId="4.[151,250,1465,1488]" captionTargetBox="[162,1434,486,1441]" captionTargetId="figure@4.[151,1436,481,1444]" captionTargetPageId="4" captionText="FIGURE 3. Hydraena lotti sp. nov. A) aedeagus, lateral view; B) aedeagus dorsal view; C) aedeagus dorsolateral view; D) spermatheca." httpUri="https://zenodo.org/record/283586/files/figure.png" pageId="4" pageNumber="33">Fig. 3</figureCitation>
D. Setae on glabrous areas of ventrites V and VI long, dense and golden.
</paragraph>
<paragraph id="8BDA36F8FFF8FFCD09C5F8A9FC39A9DD" blockId="4.[151,1437,1558,2014]" lastBlockId="5.[151,1437,151,391]" lastPageId="5" lastPageNumber="34" pageId="4" pageNumber="33">
<emphasis id="B911EAEAFFF8FFCC09C5F8A9FE07AF70" bold="true" box="[199,461,1809,1834]" pageId="4" pageNumber="33">Differential diagnosis.</emphasis>
The new species is apparently a member of the
<taxonomicName id="4C654D7BFFF8FFCC0AE3F8ABFB49AF70" box="[993,1155,1811,1834]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="emarginata">
<emphasis id="B911EAEAFFF8FFCC0AE3F8ABFB49AF70" box="[993,1155,1811,1834]" italics="true" pageId="4" pageNumber="33">H. emarginata</emphasis>
</taxonomicName>
complex (
<emphasis id="B911EAEAFFF8FFCC0DFBF8ABFAFDAF70" box="[1273,1335,1811,1834]" italics="true" pageId="4" pageNumber="33">sensu</emphasis>
Trizzino
<emphasis id="B911EAEAFFF8FFCC0995F88FFF02AF14" box="[151,200,1846,1870]" italics="true" pageId="4" pageNumber="33">et al</emphasis>
., 2013b) of the “
<emphasis id="B911EAEAFFF8FFCC0884F88EFE3CAF14" box="[390,502,1846,1870]" italics="true" pageId="4" pageNumber="33">Haenydra</emphasis>
” lineage, sharing external and aedeagal characteristics with members of this group. Externally the new species most closely resembles
<taxonomicName id="4C654D7BFFF8FFCC0BD4F8E2FBD2AF2B" authority="Jach, 1995" authorityName="Jach" authorityYear="1995" box="[726,1048,1881,1906]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="pelops">
<emphasis id="B911EAEAFFF8FFCC0BD4F8E2FC5CAF28" box="[726,918,1882,1906]" italics="true" pageId="4" pageNumber="33">Hydraena pelops</emphasis>
Jäch, 1995
</taxonomicName>
, known from the south and east of the Taygetos range. The two species are similar in size, colour, overall appearance, and modifications to male meso- and metatibia, and males are best distinguished on genitalia, which, whilst sharing the same basic pattern of the
<taxonomicName id="4C654D7BFFF8FFCC09BDF87FFEABAF84" box="[191,353,1991,2014]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="emarginata">
<emphasis id="B911EAEAFFF8FFCC09BDF87FFEABAF84" box="[191,353,1991,2014]" italics="true" pageId="4" pageNumber="33">H. emarginata</emphasis>
</taxonomicName>
complex, differ markedly in the two species. Externally,
<taxonomicName id="4C654D7BFFF8FFCC0ADFF87FFB82AF84" box="[989,1096,1990,2014]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="pelops">
<emphasis id="B911EAEAFFF8FFCC0ADFF87FFB82AF84" box="[989,1096,1990,2014]" italics="true" pageId="4" pageNumber="33">H. pelops</emphasis>
</taxonomicName>
is more sparsely punctured on the frons than the new species, and the small punctures on the clypeus are also much shallower, both characters being present in both sexes. The elytral apices of males of the two species are almost identical, but females of
<taxonomicName id="4C654D7BFFF9FFCD0C83FF05FF0EA8A2" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="lotti">
<emphasis id="B911EAEAFFF9FFCD0C83FF05FF0EA8A2" italics="true" pageId="5" pageNumber="34">H. lotti</emphasis>
</taxonomicName>
can be distinguished from those of
<taxonomicName id="4C654D7BFFF9FFCD0B61FF59FD1AA8A2" box="[611,720,224,248]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="pelops">
<emphasis id="B911EAEAFFF9FFCD0B61FF59FD1AA8A2" box="[611,720,224,248]" italics="true" pageId="5" pageNumber="34">H. pelops</emphasis>
</taxonomicName>
on this character: whereas the bitruncate apices of
<taxonomicName id="4C654D7BFFF9FFCD0C1DFF59FABAA8A2" box="[1311,1392,224,248]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="lotti">
<emphasis id="B911EAEAFFF9FFCD0C1DFF59FABAA8A2" box="[1311,1392,224,248]" italics="true" pageId="5" pageNumber="34">H. lotti</emphasis>
</taxonomicName>
are almost straight, as in
<figureCitation id="135E2A7DFFF9FFCD0892FEBCFE1FA946" box="[400,469,260,284]" captionStart="FIGURE 2" captionStartId="3.[151,250,1173,1196]" captionTargetBox="[164,1435,200,1152]" captionTargetId="figure@3.[151,1436,193,1152]" captionTargetPageId="3" captionText="FIGURE 2. Hydraena lotti sp. nov. A) male elytral apex; B) female elytral apex; C) female tergite X; D) gonocoxite." httpUri="https://zenodo.org/record/283585/files/figure.png" pageId="5" pageNumber="34">Fig. 2</figureCitation>
B, those of
<taxonomicName id="4C654D7BFFF9FFCD0B5EFEBDFD00A946" box="[604,714,260,284]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="pelops">
<emphasis id="B911EAEAFFF9FFCD0B5EFEBDFD00A946" box="[604,714,260,284]" italics="true" pageId="5" pageNumber="34">H. pelops</emphasis>
</taxonomicName>
slope inwards slightly towards the elytral suture. The truncate apices of the elytra of
<taxonomicName id="4C654D7BFFF9FFCD08A3FE91FDDAA91A" box="[417,528,296,320]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="pelops">
<emphasis id="B911EAEAFFF9FFCD08A3FE91FDDAA91A" box="[417,528,296,320]" italics="true" pageId="5" pageNumber="34">H. pelops</emphasis>
</taxonomicName>
are also typically narrower than in the new species, and more clearly serrate. Females of
<taxonomicName id="4C654D7BFFF9FFCD0819FEF5FE4DA93E" box="[283,391,332,356]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="pelops">
<emphasis id="B911EAEAFFF9FFCD0819FEF5FE4DA93E" box="[283,391,332,356]" italics="true" pageId="5" pageNumber="34">H. pelops</emphasis>
</taxonomicName>
have shorter hairs on the glabrous areas of ventrites V and VI than the new species, and the lateral apical margins of the gonocoxite are more concave, with longer setae.
</paragraph>
<caption id="DF1A6670FFF9FFCD0995F8CDFB3FAFD1" box="[151,1269,1909,1932]" httpUri="https://zenodo.org/record/283587/files/figure.png" pageId="5" pageNumber="34" targetBox="[164,1435,454,1888]" targetPageId="5">
<paragraph id="8BDA36F8FFF9FFCD0995F8CDFB3FAFD1" blockId="5.[151,1269,1909,1932]" box="[151,1269,1909,1932]" pageId="5" pageNumber="34">
<emphasis id="B911EAEAFFF9FFCD0995F8CDFED8AFD1" bold="true" box="[151,274,1909,1932]" pageId="5" pageNumber="34">FIGURE 4.</emphasis>
<taxonomicName id="4C654D7BFFF9FFCD081BF8CEFE64AFD6" box="[281,430,1910,1932]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="lotti" status="sp. nov.">
<emphasis id="B911EAEAFFF9FFCD081BF8CEFE64AFD6" box="[281,430,1910,1932]" italics="true" pageId="5" pageNumber="34">Hydraena lotti</emphasis>
</taxonomicName>
<emphasis id="B911EAEAFFF9FFCD08B6F8CEFDCEAFD1" bold="true" box="[436,516,1910,1931]" pageId="5" pageNumber="34">
<taxonomicNameLabel id="A2225791FFF9FFCD08B6F8CEFDCEAFD1" box="[436,516,1910,1931]" pageId="5" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A) – B) male metatibia; C) – D) female metatibia; E)– F) male mesotibia.
</paragraph>
</caption>
<caption id="DF1A6670FFFAFFCE0995F810FD8DAF87" httpUri="https://zenodo.org/record/283588/files/figure.png" pageId="6" pageNumber="35" targetBox="[318,1275,193,1894]" targetPageId="6">
<paragraph id="8BDA36F8FFFAFFCE0995F810FD8DAF87" blockId="6.[151,1436,1960,2013]" pageId="6" pageNumber="35">
<emphasis id="B911EAEAFFFAFFCE0995F810FEDDAFE4" bold="true" box="[151,279,1960,1983]" pageId="6" pageNumber="35">FIGURE 5.</emphasis>
Type locality of
<taxonomicName id="4C654D7BFFFAFFCE08DAF811FDB8AFE5" box="[472,626,1961,1983]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="lotti" status="sp. nov.">
<emphasis id="B911EAEAFFFAFFCE08DAF811FDB8AFE5" box="[472,626,1961,1983]" italics="true" pageId="6" pageNumber="35">Hydraena lotti</emphasis>
</taxonomicName>
<emphasis id="B911EAEAFFFAFFCE0B7FF811FD19AFE4" bold="true" box="[637,723,1961,1982]" pageId="6" pageNumber="35">
<taxonomicNameLabel id="A2225791FFFAFFCE0B7FF811FD19AFE4" box="[637,723,1961,1982]" pageId="6" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Greece, Peloponnese, Taygetos Mountains, Lagadha stream above Lagadha pass, 1,100 m. Photo D.T. Bilton.
</paragraph>
</caption>
<paragraph id="8BDA36F8FFFBFFCF09C5FF21FBDFA93E" blockId="7.[151,1436,151,1293]" pageId="7" pageNumber="36">
<taxonomicName id="4C654D7BFFFBFFCF09C5FF21FED2A8EA" box="[199,280,152,176]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="lotti">
<emphasis id="B911EAEAFFFBFFCF09C5FF21FED2A8EA" box="[199,280,152,176]" italics="true" pageId="7" pageNumber="36">H. lotti</emphasis>
</taxonomicName>
also resembles
<taxonomicName id="4C654D7BFFFBFFCF08CDFF21FD19A8F5" authority="Jach, 1992" authorityName="Jach" authorityYear="1992" box="[463,723,151,176]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="pangaei">
<emphasis id="B911EAEAFFFBFFCF08CDFF21FD84A8EA" box="[463,590,153,176]" italics="true" pageId="7" pageNumber="36">H. pangaei</emphasis>
Jäch, 1992
</taxonomicName>
, endemic to small springs on Mount Pangaeon in northeastern
<collectingCountry id="F3727668FFFBFFCF0995FF04FF23A88E" box="[151,233,188,212]" name="Greece" pageId="7" pageNumber="36">Greece</collectingCountry>
, and
<taxonomicName id="4C654D7BFFFBFFCF082BFF05FD8CA88E" authority="Fiori, 1904" authorityName="Fiori" authorityYear="1904" box="[297,582,188,212]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="samnitica">
<emphasis id="B911EAEAFFFBFFCF082BFF05FE71A88E" box="[297,443,189,212]" italics="true" pageId="7" pageNumber="36">H. samnitica</emphasis>
Fiori, 1904
</taxonomicName>
, from the central Apennines, particularly in the form of its aedeagus. The shape of the finger-like projection of the main piece differs between these species, however, as does the detailed structure of the distal lobe. Externally both
<taxonomicName id="4C654D7BFFFBFFCF0B9FFEBDFCD4A946" box="[669,798,261,284]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="pangaei">
<emphasis id="B911EAEAFFFBFFCF0B9FFEBDFCD4A946" box="[669,798,261,284]" italics="true" pageId="7" pageNumber="36">H. pangaei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C654D7BFFFBFFCF0A5CFEBDFC3BA946" box="[862,1009,261,284]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="samnitica">
<emphasis id="B911EAEAFFFBFFCF0A5CFEBDFC3BA946" box="[862,1009,261,284]" italics="true" pageId="7" pageNumber="36">H. samnitica</emphasis>
</taxonomicName>
are somewhat larger at 2.5–2.7 and
<quantity id="4C9D9B1DFFFBFFCF0995FE9FFEE8A965" box="[151,290,295,319]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="7" pageNumber="36" unit="mm" value="2.5" valueMax="2.7" valueMin="2.3">2.3–2.7 mm</quantity>
in length respectively, and females of both species have more rounded elytral apices, those of
<taxonomicName id="4C654D7BFFFBFFCF0C83FE91FECEA93E" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="samnitica">
<emphasis id="B911EAEAFFFBFFCF0C83FE91FECEA93E" italics="true" pageId="7" pageNumber="36">H. samnitica</emphasis>
</taxonomicName>
much more so (see Trizzino
<emphasis id="B911EAEAFFFBFFCF0B4BFEF5FD48A93E" box="[585,642,332,356]" italics="true" pageId="7" pageNumber="36">et al.</emphasis>
, 2013b for details of these species).
</paragraph>
</subSubSection>
<subSubSection id="C37F6573FFFBFFCF09C5FED7FC51AA4D" pageId="7" pageNumber="36" type="distribution">
<paragraph id="8BDA36F8FFFBFFCF09C5FED7FC51AA4D" blockId="7.[151,1436,151,1293]" pageId="7" pageNumber="36">
<emphasis id="B911EAEAFFFBFFCF09C5FED7FE93A9D2" bold="true" box="[199,345,367,392]" pageId="7" pageNumber="36">Distribution</emphasis>
. Found in four headwater streams, situated on closely adjacent sides of the watershed in the northwest of the Taygetos massif, where it is likely endemic. The
<typeStatus id="54DE885AFFFBFFCF0A8AFE2DFC72A9F7" box="[904,952,405,429]" pageId="7" pageNumber="36">type</typeStatus>
locality, the Lagadha stream, drains to the east, forming part of the Evrotas basin. The three remaining localities all drain to the west, being part of the Strogli and Nedhon drainages respectively. The short flight wings suggest poor dispersal ability, characteristic of many narrow-range endemics (see Bilton
<emphasis id="B911EAEAFFFBFFCF0B24FDB9FD92AA42" box="[550,600,512,536]" italics="true" pageId="7" pageNumber="36">et al</emphasis>
., 2001, Arribas
<emphasis id="B911EAEAFFFBFFCF0A0FFDB9FCF5AA42" box="[781,831,512,536]" italics="true" pageId="7" pageNumber="36">et al</emphasis>
., 2011).
</paragraph>
</subSubSection>
<subSubSection id="C37F6573FFFBFFCF09C5FD9CFCC1AD57" pageId="7" pageNumber="36" type="etymology">
<paragraph id="8BDA36F8FFFBFFCF09C5FD9CFE0DAA05" blockId="7.[151,1436,151,1293]" pageId="7" pageNumber="36">
<emphasis id="B911EAEAFFFBFFCF09C5FD9CFE8CAA66" bold="true" box="[199,326,548,572]" pageId="7" pageNumber="36">Etymology</emphasis>
. Named after the late Dr Derek Lott, in recognition of his contribution to the study of both aquatic and staphyliniform beetles.
</paragraph>
<paragraph id="8BDA36F8FFFBFFCF09C5FDD4FCC1AD57" blockId="7.[151,1436,151,1293]" pageId="7" pageNumber="36">
<emphasis id="B911EAEAFFFBFFCF09C5FDD4FEEEAADE" bold="true" box="[199,292,620,644]" pageId="7" pageNumber="36">Ecology</emphasis>
. The new species is found in the upper reaches of swift flowing, first order streams, all four known localities being just above
<quantity id="4C9D9B1DFFFBFFCF08D5FD37FDF2AAFD" box="[471,568,655,679]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="36" unit="m" value="1000.0">1,000 m</quantity>
in altitude. Localities are apparently permanent, and mostly flow over a combination blue schist and metamorphosed siltstone. Most streams in the Taygetos flow over limestone, and geology may be important to the distribution of “
<emphasis id="B911EAEAFFFBFFCF0BC1FD60FCF9AAAA" box="[707,819,728,752]" italics="true" pageId="7" pageNumber="36">Haenydra</emphasis>
” in the region, although more data are required before firm conclusions can be drawn.
<taxonomicName id="4C654D7BFFFBFFCF0B03FD45FD9BAB4E" box="[513,593,764,788]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="lotti">
<emphasis id="B911EAEAFFFBFFCF0B03FD45FD9BAB4E" box="[513,593,764,788]" italics="true" pageId="7" pageNumber="36">H. lotti</emphasis>
</taxonomicName>
is most abundant in small shallow riffles, particularly under stones. In the
<typeStatus id="54DE885AFFFBFFCF0995FC99FF0DAB63" box="[151,199,801,825]" pageId="7" pageNumber="36">type</typeStatus>
locality above Lagadha Pass (
<figureCitation id="135E2A7DFFFBFFCF0B3FFCA7FD4CAB6D" box="[573,646,799,823]" captionStart="FIGURE 5" captionStartId="6.[151,250,1960,1983]" captionTargetBox="[318,1275,193,1894]" captionTargetId="figure@6.[318,1275,193,1894]" captionTargetPageId="6" captionText="FIGURE 5. Type locality of Hydraena lotti sp. nov. Greece, Peloponnese, Taygetos Mountains, Lagadha stream above Lagadha pass, 1,100 m. Photo D. T. Bilton." httpUri="https://zenodo.org/record/283588/files/figure.png" pageId="7" pageNumber="36">Fig. 5</figureCitation>
) it shared this microhabitat with
<taxonomicName id="4C654D7BFFFBFFCF0D23FC99FA52AB6D" authority="Fairmaire" authorityName="Fairmaire" box="[1057,1432,799,824]" class="Insecta" family="Dytiscidae" genus="Deronectes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="moestus">
<emphasis id="B911EAEAFFFBFFCF0D23FC99FACDAB62" box="[1057,1287,801,824]" italics="true" pageId="7" pageNumber="36">Deronectes moestus</emphasis>
(Fairmaire)
</taxonomicName>
,
<taxonomicName id="4C654D7BFFFBFFCF0995FCFCFE32AB06" authority="Drapiez" authorityName="Drapiez" box="[151,504,836,860]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="tesselatus">
<emphasis id="B911EAEAFFFBFFCF0995FCFCFE5EAB06" box="[151,404,836,860]" italics="true" pageId="7" pageNumber="36">Hydroporus tesselatus</emphasis>
Drapiez
</taxonomicName>
,
<taxonomicName id="4C654D7BFFFBFFCF0B0BFCFCFC8CAB06" authority="Olivier" authorityName="Olivier" box="[521,838,836,860]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="biguttatus">
<emphasis id="B911EAEAFFFBFFCF0B0BFCFCFD1DAB06" box="[521,727,836,860]" italics="true" pageId="7" pageNumber="36">Agabus biguttatus</emphasis>
(Olivier)
</taxonomicName>
,
<taxonomicName id="4C654D7BFFFBFFCF0A57FCFCFB47AB06" authority="Rey, H." authorityName="Rey, H." box="[853,1165,836,861]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="armipes">
<emphasis id="B911EAEAFFFBFFCF0A57FCFCFBE0AB06" box="[853,1066,836,860]" italics="true" pageId="7" pageNumber="36">Hydraena armipes</emphasis>
Rey,
<emphasis id="B911EAEAFFFBFFCF0D73FCFDFB47AB06" box="[1137,1165,837,860]" italics="true" pageId="7" pageNumber="36">H.</emphasis>
</taxonomicName>
christinae Audisio,
<collectingCountry id="F3727668FFFBFFCF0C79FCFDFA56AB06" box="[1403,1436,837,860]" name="South Georgia and the South Sandwich Islands" pageId="7" pageNumber="36">De</collectingCountry>
Biase &amp; Jäch,
<taxonomicName id="4C654D7BFFFBFFCF0835FCD1FD46ABDA" authority="Orchymont, H." authorityName="Orchymont, H." box="[311,652,871,896]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="georgiadesi">
<emphasis id="B911EAEAFFFBFFCF0835FCD1FE16ABDA" box="[311,476,872,896]" italics="true" pageId="7" pageNumber="36">H. georgiadesi</emphasis>
Orchymont,
<emphasis id="B911EAEAFFFBFFCF0B72FCD1FD46ABDA" box="[624,652,873,896]" italics="true" pageId="7" pageNumber="36">H.</emphasis>
</taxonomicName>
helena Orchymont,
<taxonomicName id="4C654D7BFFFBFFCF0A72FCD1FB5BABDA" authority="Pretner, H." authorityName="Pretner, H." box="[880,1169,872,896]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="meschniggi">
<emphasis id="B911EAEAFFFBFFCF0A72FCD1FBDBABDA" box="[880,1041,872,896]" italics="true" pageId="7" pageNumber="36">H. meschniggi</emphasis>
Pretner,
<emphasis id="B911EAEAFFFBFFCF0D77FCD1FB5BABDA" box="[1141,1169,873,896]" italics="true" pageId="7" pageNumber="36">H.</emphasis>
</taxonomicName>
morio Kiesenwetter,
<taxonomicName id="4C654D7BFFFBFFCF0C83FCD1FE44ABFE" authority="Orchymont" authorityName="Orchymont" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="subjuncta">
<emphasis id="B911EAEAFFFBFFCF0C83FCD1FECEABFE" italics="true" pageId="7" pageNumber="36">H. subjuncta</emphasis>
Orchymont
</taxonomicName>
,
<taxonomicName id="4C654D7BFFFBFFCF0898FC34FCB6ABFE" authority="Ganglbauer, O." authorityName="Ganglbauer, O." box="[410,892,908,932]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="montenegrinus">
<emphasis id="B911EAEAFFFBFFCF0898FC34FD02ABFE" box="[410,712,908,932]" italics="true" pageId="7" pageNumber="36">Ochthebius montenegrinus</emphasis>
Ganglbauer,
<emphasis id="B911EAEAFFFBFFCF0A63FC35FCB6ABFE" box="[865,892,909,932]" italics="true" pageId="7" pageNumber="36">
<collectingCountry id="F3727668FFFBFFCF0A63FC35FCB2ABFE" box="[865,888,909,932]" name="Iceland" pageId="7" pageNumber="36">O</collectingCountry>
.
</emphasis>
</taxonomicName>
striatus (Castelnau),
<taxonomicName id="4C654D7BFFFBFFCF0D72FC34FB7AABFE" box="[1136,1200,908,932]" class="Insecta" family="Elmidae" genus="Elmis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAEAFFFBFFCF0D72FC34FB7AABFE" box="[1136,1200,908,932]" italics="true" pageId="7" pageNumber="36">Elmis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C654D7BFFFBFFCF0DEBFC35FA89ABFE" box="[1257,1347,909,932]" class="Insecta" family="Elmidae" genus="Limnius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAEAFFFBFFCF0DEBFC35FA89ABFE" box="[1257,1347,909,932]" italics="true" pageId="7" pageNumber="36">Limnius</emphasis>
</taxonomicName>
species and
<taxonomicName id="4C654D7BFFFBFFCF09CFFC08FCB2AB92" authority="Nyholm. In Dzirorema and Boursos" authorityName="Nyholm. In Dzirorema and Boursos" box="[205,888,943,968]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="novaki">
<emphasis id="B911EAEAFFFBFFCF09CFFC08FE77AB92" box="[205,445,944,968]" italics="true" pageId="7" pageNumber="36">Hydrocyphon novaki</emphasis>
Nyholm. In Dzirorema and Boursos
</taxonomicName>
streams the species was found together with
<emphasis id="B911EAEAFFFBFFCF0995FC6DFBF7ABB6" box="[151,1085,980,1004]" italics="true" pageId="7" pageNumber="36">
<taxonomicName id="4C654D7BFFFBFFCF0995FC6DFEB3ABB6" box="[151,377,981,1004]" class="Insecta" family="Dytiscidae" genus="Deronectes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="moestus">Deronectes moestus</taxonomicName>
,
<taxonomicName id="4C654D7BFFFBFFCF0887FC6CFD48ABB6" box="[389,642,980,1004]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="tesselatus">Hydroporus tesselatus</taxonomicName>
,
<taxonomicName id="4C654D7BFFFBFFCF0B8CFC6CFC96ABB6" box="[654,860,980,1004]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="biguttatus">Agabus biguttatus</taxonomicName>
,
<taxonomicName id="4C654D7BFFFBFFCF0A65FC6CFBF7ABB6" authority="Ferro, H." authorityName="Ferro, H." box="[871,1085,980,1004]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="aroensis">Hydraena aroensis</taxonomicName>
</emphasis>
Ferro,
<emphasis id="B911EAEAFFFBFFCF0D8DFC6DFDECAC4A" italics="true" pageId="7" pageNumber="36">
<taxonomicName id="4C654D7BFFFBFFCF0D8DFC6DFAEEABB6" box="[1167,1316,980,1004]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="christinae">H. christinae</taxonomicName>
,
<taxonomicName id="4C654D7BFFFBFFCF0C2DFC6DFA52ABB6" box="[1327,1432,981,1004]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="morio">H. morio</taxonomicName>
,
<taxonomicName id="4C654D7BFFFBFFCF0995FC41FEE3AC4A" box="[151,297,1016,1040]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="subjuncta">H. subjuncta</taxonomicName>
,
<taxonomicName id="4C654D7BFFFBFFCF0831FC40FDECAC4A" authority="Frivaldzsky" authorityName="Frivaldzsky" box="[307,550,1016,1040]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="montanus">Ochthebius montanus</taxonomicName>
</emphasis>
Frivaldzsky,
<emphasis id="B911EAEAFFFBFFCF0BBEFC40FC38AC4A" box="[700,1010,1016,1040]" italics="true" pageId="7" pageNumber="36">
<taxonomicName id="4C654D7BFFFBFFCF0BBEFC40FC6DAC4A" box="[700,935,1016,1040]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="novaki">Hydrocyphon novaki</taxonomicName>
,
<taxonomicName id="4C654D7BFFFBFFCF0AB0FC40FC38AC4A" box="[946,1010,1016,1040]" class="Insecta" family="Elmidae" genus="Elmis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="genus">Elmis</taxonomicName>
</emphasis>
spp. and
<taxonomicName id="4C654D7BFFFBFFCF0D5EFC41FB7CAC4A" box="[1116,1206,1017,1040]" class="Insecta" family="Elmidae" genus="Limnius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAEAFFFBFFCF0D5EFC41FB7CAC4A" box="[1116,1206,1017,1040]" italics="true" pageId="7" pageNumber="36">Limnius</emphasis>
</taxonomicName>
spp. Accompanying species in the Sitsovitiko stream were
<emphasis id="B911EAEAFFFBFFCF0B5CFBA4FAE5AC6E" box="[606,1327,1052,1076]" italics="true" pageId="7" pageNumber="36">
<taxonomicName id="4C654D7BFFFBFFCF0B5CFBA4FCF0AC6E" box="[606,826,1052,1076]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="aroensis">Hydraena aroensis</taxonomicName>
,
<taxonomicName id="4C654D7BFFFBFFCF0A48FBA5FC77AC6E" box="[842,957,1052,1076]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="helena">H. helena</taxonomicName>
,
<taxonomicName id="4C654D7BFFFBFFCF0ACFFBA5FBF0AC6E" box="[973,1082,1053,1076]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="morio">H. morio</taxonomicName>
,
<taxonomicName id="4C654D7BFFFBFFCF0D4BFBA5FB15AC6E" box="[1097,1247,1052,1076]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="subjuncta">H. subjuncta</taxonomicName>
,
<taxonomicName id="4C654D7BFFFBFFCF0DEDFBA4FAE5AC6E" box="[1263,1327,1052,1076]" class="Insecta" family="Elmidae" genus="Elmis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="genus">Elmis</taxonomicName>
</emphasis>
spp. and
<taxonomicName id="4C654D7BFFFBFFCF0995FBF9FF3BAC02" box="[151,241,1089,1112]" class="Insecta" family="Elmidae" genus="Limnius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAEAFFFBFFCF0995FBF9FF3BAC02" box="[151,241,1089,1112]" italics="true" pageId="7" pageNumber="36">Limnius</emphasis>
</taxonomicName>
spp. In all localities
<taxonomicName id="4C654D7BFFFBFFCF08D8FBF9FDE3AC02" box="[474,553,1088,1112]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="lotti" status="sp. nov.">
<emphasis id="B911EAEAFFFBFFCF08D8FBF9FDE3AC02" box="[474,553,1088,1112]" italics="true" pageId="7" pageNumber="36">H. lotti</emphasis>
</taxonomicName>
<emphasis id="B911EAEAFFFBFFCF0B32FBF8FD43AC02" bold="true" box="[560,649,1088,1112]" pageId="7" pageNumber="36">
<taxonomicNameLabel id="A2225791FFFBFFCF0B32FBF8FD43AC02" box="[560,649,1088,1112]" pageId="7" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was the commonest
<taxonomicName id="4C654D7BFFFBFFCF0A76FBF8FC2EAC02" box="[884,996,1088,1112]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAEAFFFBFFCF0A76FBF8FC2EAC02" box="[884,996,1088,1112]" italics="true" pageId="7" pageNumber="36">Hydraena</emphasis>
</taxonomicName>
, with the exception of
<taxonomicName id="4C654D7BFFFBFFCF0DE7FBF9FA81AC02" box="[1253,1355,1089,1112]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B911EAEAFFFBFFCF0DE7FBF9FA81AC02" box="[1253,1355,1089,1112]" italics="true" pageId="7" pageNumber="36">H. morio</emphasis>
</taxonomicName>
and
<taxonomicName id="4C654D7BFFFBFFCF0C83FBF9FECEAC26" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="subjuncta">
<emphasis id="B911EAEAFFFBFFCF0C83FBF9FECEAC26" italics="true" pageId="7" pageNumber="36">H. subjuncta</emphasis>
</taxonomicName>
. The stream above Lagadha Pass flows through pine forest and is fringed with a riparian strip of
<taxonomicName id="4C654D7BFFFBFFCF0C3AFBDCFEC9ACFA" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="7" pageNumber="36" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="B911EAEAFFFBFFCF0C3AFBDCFEC9ACFA" italics="true" pageId="7" pageNumber="36">Platanus orientalis</emphasis>
</taxonomicName>
L., whilst other localities are in areas burned during fires earlier this century, when the forest cover of parts of the western Taygetos was damaged or lost. These sites are mostly fringed by remnant resprouting trees and secondary scrub growth. So far the new species has not been collected together with
<taxonomicName id="4C654D7BFFFBFFCF0D58FB69FB0DACB2" box="[1114,1223,1232,1256]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="pelops">
<emphasis id="B911EAEAFFFBFFCF0D58FB69FB0DACB2" box="[1114,1223,1232,1256]" italics="true" pageId="7" pageNumber="36">H. pelops</emphasis>
</taxonomicName>
, which to date has been found further south and east in the Taygetos range.
</paragraph>
</subSubSection>
<subSubSection id="C37F6573FFFBFFC00995FAE7FE88AA66" lastPageId="8" lastPageNumber="37" pageId="7" pageNumber="36" type="description">
<paragraph id="8BDA36F8FFFBFFCF0995FAE7FDC9AE15" blockId="7.[151,978,1375,1400]" lastBlockId="7.[151,1436,1591,1976]" pageId="7" pageNumber="36">
<emphasis id="B911EAEAFFFBFFCF0995FAE7FC18AD22" bold="true" box="[151,978,1375,1400]" pageId="7" pageNumber="36">
Annotated checklist of Peloponnese species of the “
<emphasis id="B911EAEAFFFBFFCF0BF4FAE7FCA3AD22" bold="true" box="[758,873,1375,1400]" italics="true" pageId="7" pageNumber="36">Haenydra</emphasis>
” lineage
</emphasis>
*
<taxonomicName id="4C654D7BFFFBFFCF09A7F980FDC9AE15" authority="Ferro, 1991" authorityName="Ferro" authorityYear="1991" box="[165,515,1591,1616]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="aroensis">
<emphasis id="B911EAEAFFFBFFCF09A7F980FEB0AE0A" box="[165,378,1592,1616]" italics="true" pageId="7" pageNumber="36">Hydraena aroensis</emphasis>
Ferro, 1991
</taxonomicName>
</paragraph>
<paragraph id="8BDA36F8FFFBFFCF0995FA1FFEC1AE5D" pageId="7" pageNumber="36">
<table id="F965C458FFFB00370995FA1FFA56AE5D" box="[151,1436,1447,1543]" gridcols="4" gridrows="3" pageId="7" pageNumber="36">
<tr id="355534BAFFFB00370995FA1FFA56AD9A" box="[151,1436,1447,1472]" gridrow="0" pageId="7" pageNumber="36">
<th id="76845DC6FFFB00370995FA1FFDC6AD9A" box="[151,524,1447,1472]" gridcol="0" gridrow="0" pageId="7" pageNumber="36">
With the description of
<taxonomicName id="4C654D7BFFFBFFCF08B6FA11FDC2AD9A" box="[436,520,1448,1472]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="lotti">
<emphasis id="B911EAEAFFFBFFCF08B6FA11FDC2AD9A" box="[436,520,1448,1472]" italics="true" pageId="7" pageNumber="36">H. lotti</emphasis>
</taxonomicName>
</th>
<th id="76845DC6FFFB00370B17FA1FFCBCAD9A" box="[533,886,1447,1472]" gridcol="1" gridrow="0" pageId="7" pageNumber="36">
<emphasis id="B911EAEAFFFBFFCF0B17FA10FDBEAD9A" bold="true" box="[533,628,1448,1472]" pageId="7" pageNumber="36">sp. nov.</emphasis>
, seven species of the
</th>
<th id="76845DC6FFFB00370A82FA1FFABAAD9A" box="[896,1392,1447,1472]" gridcol="2" gridrow="0" pageId="7" pageNumber="36">
“
<emphasis id="B911EAEAFFFBFFCF0A8CFA10FC34AD9A" box="[910,1022,1448,1472]" italics="true" pageId="7" pageNumber="36">Haenydra</emphasis>
” lineage are now known from
</th>
<th id="76845DC6FFFB00370C78FA1FFA56AD9A" box="[1402,1436,1447,1472]" gridcol="3" gridrow="0" pageId="7" pageNumber="36">the</th>
</tr>
<tr id="355534BAFFFB00370995FA74FA56ADBE" box="[151,1436,1484,1508]" gridrow="1" pageId="7" pageNumber="36">
<th id="76845DC6FFFB00370995FA74FDC6ADBE" box="[151,524,1484,1508]" gridcol="0" gridrow="1" pageId="7" pageNumber="36">Peloponnese. The distribution of</th>
<td id="76845DC6FFFB00370B17FA74FCBCADBE" box="[533,886,1484,1508]" gridcol="1" gridrow="1" pageId="7" pageNumber="36">these is summarized below, an</td>
<td id="76845DC6FFFB00370A82FA74FABAADBE" box="[896,1392,1484,1508]" gridcol="2" gridrow="1" pageId="7" pageNumber="36">asterisk indicating taxa that are endemic to</td>
<td id="76845DC6FFFB00370C78FA74FA56ADBE" box="[1402,1436,1484,1508]" gridcol="3" gridrow="1" pageId="7" pageNumber="36">the</td>
</tr>
<tr id="355534BAFFFB00370995FA57FA56AE5D" box="[151,1436,1519,1543]" gridrow="2" pageId="7" pageNumber="36" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76845DC6FFFB00370995FA57FDC6AE5D" box="[151,524,1519,1543]" gridcol="0" gridrow="2" pageId="7" pageNumber="36">peninsula:</th>
</tr>
</table>
</paragraph>
<paragraph id="8BDA36F8FFFBFFCF09C5F9E4FCEBAE85" blockId="7.[151,1436,1591,1976]" pageId="7" pageNumber="36">
Previously known only from the northern Peloponnese, (Audisio
<emphasis id="B911EAEAFFFBFFCF0A9DF9E5FC1DAE2E" box="[927,983,1628,1652]" italics="true" pageId="7" pageNumber="36">et al.</emphasis>
, 1996; Trizzino
<emphasis id="B911EAEAFFFBFFCF0D8DF9E5FB75AE2E" box="[1167,1215,1628,1652]" italics="true" pageId="7" pageNumber="36">et al</emphasis>
., 2013b). Recorded together with
<taxonomicName id="4C654D7BFFFBFFCF0861F939FE79AEC2" box="[355,435,1664,1688]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="lotti" status="sp. nov.">
<emphasis id="B911EAEAFFFBFFCF0861F939FE79AEC2" box="[355,435,1664,1688]" italics="true" pageId="7" pageNumber="36">H. lotti</emphasis>
</taxonomicName>
<emphasis id="B911EAEAFFFBFFCF08BEF938FDDDAEC2" bold="true" box="[444,535,1664,1688]" pageId="7" pageNumber="36">
<taxonomicNameLabel id="A2225791FFFBFFCF08BEF938FDDDAEC2" box="[444,535,1664,1688]" pageId="7" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
at three localities in the northwestern Taygetos in 2012 (see above), and
<date id="FFDB1038FFFBFFCF0C59F9C7FECAAEE6" pageId="7" pageNumber="36" value="2012-06-28">28/vi/ 2012</date>
,
<collectingCountry id="F3727668FFFBFFCF0810F91CFEAEAEE6" box="[274,356,1700,1724]" name="Greece" pageId="7" pageNumber="36">Greece</collectingCountry>
, Peloponnese, Taygetos Mts., Vrisiotiko Bk. nr. Kastorio,
<quantity id="4C9D9B1DFFFBFFCF0D29F91CFBB6AEE6" box="[1067,1148,1700,1724]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="36" unit="m" value="500.0">500 m</quantity>
, D.T. Bilton leg., these extending the range of the species south considerably.
</paragraph>
<paragraph id="8BDA36F8FFFBFFCF0995F954FD31AF5E" blockId="7.[151,1436,1591,1976]" box="[151,763,1772,1796]" pageId="7" pageNumber="36">
*
<taxonomicName id="4C654D7BFFFBFFCF09A7F954FD31AF5E" authority="Audisio, De Biase &amp; Jach, 1996" authorityName="Audisio, De Biase &amp; Jach" authorityYear="1996" box="[165,763,1772,1796]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="christinae">
<emphasis id="B911EAEAFFFBFFCF09A7F954FE46AF5E" box="[165,396,1772,1796]" italics="true" pageId="7" pageNumber="36">Hydraena christinae</emphasis>
Audisio,
<collectingCountry id="F3727668FFFBFFCF08FBF955FDD0AF5E" box="[505,538,1773,1796]" name="South Georgia and the South Sandwich Islands" pageId="7" pageNumber="36">De</collectingCountry>
Biase &amp; Jäch, 1996
</taxonomicName>
</paragraph>
<paragraph id="8BDA36F8FFFBFFCF09C5F8B7FE53AF17" blockId="7.[151,1436,1591,1976]" pageId="7" pageNumber="36">Widespread in mountain streams and rivers along the central spine of the Peloponnese, from Achaia to the southern Taygetos.</paragraph>
<paragraph id="8BDA36F8FFFBFFCF0995F8E0FE3AAF35" blockId="7.[151,1436,1591,1976]" box="[151,496,1879,1904]" pageId="7" pageNumber="36">
<taxonomicName id="4C654D7BFFFBFFCF0995F8E0FE3AAF35" authority="Ferro, 1985" authorityName="Ferro" authorityYear="1985" box="[151,496,1879,1904]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="epeirosi">
<emphasis id="B911EAEAFFFBFFCF0995F8E0FEADAF2A" box="[151,359,1880,1904]" italics="true" pageId="7" pageNumber="36">Hydraena epeirosi</emphasis>
Ferro, 1985
</taxonomicName>
</paragraph>
<paragraph id="8BDA36F8FFFBFFCF09C5F8C4FBC3AFE2" blockId="7.[151,1436,1591,1976]" pageId="7" pageNumber="36">
Cited from the Peloponnese by Trizzino
<emphasis id="B911EAEAFFFBFFCF0B9EF8C5FD1AAFCE" box="[668,720,1916,1940]" italics="true" pageId="7" pageNumber="36">et al</emphasis>
(2013b). I can add the following record –
<date id="FFDB1038FFFBFFCF0DC0F8C4FAF1AFCE" box="[1218,1339,1916,1940]" pageId="7" pageNumber="36" value="2012-06-24">24/vi/2012</date>
<collectingCountry id="F3727668FFFBFFCF0C47F8C4FA5DAFCE" box="[1349,1431,1916,1940]" name="Greece" pageId="7" pageNumber="36">Greece</collectingCountry>
, Peloponnese, Selinoudas River below Kato Vlasia,
<quantity id="4C9D9B1DFFFBFFCF0A04F818FC85AFE2" box="[774,847,1952,1976]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="36" unit="m" value="700.0">700 m</quantity>
, D.T. Bilton leg.
</paragraph>
<paragraph id="8BDA36F8FFF4FFC00995FF2FFD06A8F5" blockId="8.[151,1436,151,572]" box="[151,716,151,176]" pageId="8" pageNumber="37">
*
<taxonomicName id="4C654D7BFFF4FFC009A7FF20FD06A8F5" authority="Audisio &amp; De Biase, 1990" authorityName="Audisio &amp; De Biase" authorityYear="1990" box="[165,716,151,176]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="jaechiana">
<emphasis id="B911EAEAFFF4FFC009A7FF20FE40A8EA" box="[165,394,152,176]" italics="true" pageId="8" pageNumber="37">Hydraena jaechiana</emphasis>
(Audisio &amp;
<collectingCountry id="F3727668FFF4FFC00B15FF20FDF2A8F5" box="[535,568,152,175]" name="South Georgia and the South Sandwich Islands" pageId="8" pageNumber="37">De</collectingCountry>
Biase, 1990)
</taxonomicName>
</paragraph>
<paragraph id="8BDA36F8FFF4FFC009C5FF04FB57A88E" blockId="8.[151,1436,151,572]" box="[199,1181,188,212]" pageId="8" pageNumber="37">
The commonest “
<emphasis id="B911EAEAFFF4FFC0088CFF04FE34A88E" box="[398,510,188,212]" italics="true" pageId="8" pageNumber="37">Haenydra</emphasis>
” in the peninsula – widespread in streams and larger rivers.
</paragraph>
<paragraph id="8BDA36F8FFF4FFC00995FF67FE63A8A2" blockId="8.[151,1436,151,572]" box="[151,425,223,248]" pageId="8" pageNumber="37">
*
<taxonomicName id="4C654D7BFFF4FFC009A7FF58FE83A8A2" box="[165,329,224,248]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="lotti" status="sp. nov.">
<emphasis id="B911EAEAFFF4FFC009A7FF58FE83A8A2" box="[165,329,224,248]" italics="true" pageId="8" pageNumber="37">Hydraena lotti</emphasis>
</taxonomicName>
<emphasis id="B911EAEAFFF4FFC00852FF58FE63A8A2" bold="true" box="[336,425,224,248]" pageId="8" pageNumber="37">
<taxonomicNameLabel id="A2225791FFF4FFC00852FF58FE63A8A2" box="[336,425,224,248]" pageId="8" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDA36F8FFF4FFC009CCFEBDFEDDA965" blockId="8.[151,1436,151,572]" pageId="8" pageNumber="37">Four streams in the northwestern section of the Taygetos, all flowing over metamorphosed mudstones and schists.</paragraph>
<paragraph id="8BDA36F8FFF4FFC00995FEF4FE2EA93E" blockId="8.[151,1436,151,572]" box="[151,484,332,356]" pageId="8" pageNumber="37">
*
<taxonomicName id="4C654D7BFFF4FFC009A7FEF4FE2EA93E" authority="Jach, 1995" authorityName="Jach" authorityYear="1995" box="[165,484,332,356]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="pelops">
<emphasis id="B911EAEAFFF4FFC009A7FEF4FEAFA93E" box="[165,357,332,356]" italics="true" pageId="8" pageNumber="37">Hydraena pelops</emphasis>
Jäch, 1995
</taxonomicName>
</paragraph>
<paragraph id="8BDA36F8FFF4FFC009C5FEC8FE02A995" blockId="8.[151,1436,151,572]" pageId="8" pageNumber="37">
Known from the southern and eastern sections of the Taygetos range, in permanent sections of high altitude streams. I can add the following record –
<date id="FFDB1038FFF4FFC00B91FE2CFD06A9F6" box="[659,716,404,428]" bridgedPair="/" pageId="8" pageNumber="37" value="1930-06">30/vi</date>
&amp;
<date id="FFDB1038FFF4FFC00BEDFE2CFCA3A9F6" box="[751,873,404,428]" pageId="8" pageNumber="37" value="2012-08-03">3/viii/2012</date>
<collectingCountry id="F3727668FFF4FFC00A72FE2CFC08A9F6" box="[880,962,404,428]" name="Greece" pageId="8" pageNumber="37">Greece</collectingCountry>
, Peloponnese, Loga stream above Anogia,
<quantity id="4C9D9B1DFFF4FFC009C7FE0FFEC4A995" box="[197,270,439,463]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="37" unit="m" value="800.0">800 m</quantity>
, D.T. Bilton leg.
</paragraph>
<paragraph id="8BDA36F8FFF4FFC00995FE64FDE7A9AE" blockId="8.[151,1436,151,572]" box="[151,557,476,500]" pageId="8" pageNumber="37">
<taxonomicName id="4C654D7BFFF4FFC00995FE64FDE7A9AE" authority="Orchymont, 1931" authorityName="Orchymont" authorityYear="1931" box="[151,557,476,500]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="vedrasi">
<emphasis id="B911EAEAFFF4FFC00995FE64FEAAA9AE" box="[151,352,476,500]" italics="true" pageId="8" pageNumber="37">Hydraena vedrasi</emphasis>
Orchymont, 1931
</taxonomicName>
</paragraph>
<paragraph id="8BDA36F8FFF4FFC009C5FDB8FE88AA66" blockId="8.[151,1436,151,572]" pageId="8" pageNumber="37">A widespread SE European species, which in the Peloponnese is so far only recorded from the northern mountains.</paragraph>
</subSubSection>
</treatment>
</document>
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