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<document id="A5AF77AEED27DB98B1604D566337D7F7" ID-CLB-Dataset="22297" ID-DOI="10.5852/ejt.2018.455" ID-GBIF-Dataset="44a2fcf6-cc1b-4c07-9554-dd6dd279a76d" ID-Zenodo-Dep="3817707" ID-ZooBank="C445E95B-446A-4601-AAA3-C1CCBAB627F9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1588963536872" checkinUser="valdenar" docAuthor="Páll-Gergely, Barna" docDate="2018" docId="03CC87A06C2A8A20FDD1FC0B467CFCF3" docLanguage="en" docName="ejt-455_pall-gergely.pdf.imf" docOrigin="European Journal of Taxonomy 455" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Plectopylidae Mollendorff 1898" docType="treatment" docVersion="6" lastPageNumber="8" masterDocId="FFF5FFD86C2C8A27FFF8FFC04709FFA9" masterDocTitle="Systematic revision of the Plectopylinae (Gastropoda, Pulmonata, Plectopylidae)" masterLastPageNumber="114" masterPageNumber="1" pageNumber="7" updateTime="1698839078568" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2381EDDA14E779F9BF36DA87E78BE037">Systematic revision of the Plectopylinae (Gastropoda, Pulmonata, Plectopylidae)</mods:title>
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<mods:namePart id="7668B0C3D873A20D0082981593D796B3">Páll-Gergely, Barna</mods:namePart>
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<mods:affiliation id="90C27B9C58B1CB8F2DB5945817AA9D32">Centre for Agricultural Research, Hungarian Academy of Sciences (MTA), Herman Ottó Street 15, Budapest, H- 1022, Hungary.</mods:affiliation>
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<mods:date id="43242BDDCBBB7384E2A5E6A1BF8D062D">2018</mods:date>
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<treatment id="03CC87A06C2A8A20FDD1FC0B467CFCF3" ID-DOI="http://doi.org/10.5281/zenodo.3818768" ID-GBIF-Taxon="163993770" ID-Zenodo-Dep="3818768" LSID="urn:lsid:plazi:treatment:03CC87A06C2A8A20FDD1FC0B467CFCF3" httpUri="http://treatment.plazi.org/id/03CC87A06C2A8A20FDD1FC0B467CFCF3" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<subSubSection id="C37F653D6C2A8A21FDD1FC0B4302FC4C" box="[553,1035,971,997]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BDA36B66C2A8A21FDD1FC0B4302FC4C" blockId="6.[553,1035,971,997]" box="[553,1035,971,997]" pageId="6" pageNumber="7">
<heading id="D09281DA6C2A8A21FDD1FC0B4302FC4C" box="[553,1035,971,997]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
Family
<taxonomicName id="4C654D356C2A8A21FD7BFC0B4302FC4C" ID-CoL="7NKJG" authority="Mollendorff, 1898" authorityName="Mollendorff" authorityYear="1898" box="[643,1035,971,997]" class="Gastropoda" family="Plectopylidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="family">
<emphasis id="B911EAA46C2A8A21FD7BFC0B4422FC4C" bold="true" box="[643,811,971,997]" pageId="6" pageNumber="7">Plectopylidae</emphasis>
<bibRefCitation id="EFF44B476C2A8A21FCCAFC0B4302FC4C" author="Mollendorff O. F. von" box="[818,1035,971,997]" pageId="6" pageNumber="7" pagination="25 - 208" refId="ref54802" refString="Mollendorff O. F. von 1898. Verzeichniss der auf den Philippinen lebenden Landmollusken. Abhandlungen der Naturforschenden Gesellschaft zu Gorlitz 22: 25 - 208." type="journal article" year="1898">Möllendorff, 1898</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C37F653D6C2A8A21FF45FBCC46F9FB3E" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="8BDA36B66C2A8A21FF45FBCC4576FB8E" blockId="6.[189,639,1036,1063]" box="[189,639,1036,1063]" pageId="6" pageNumber="7">
<taxonomicName id="4C654D356C2A8A21FF45FBCC4572FB8E" ID-CoL="7NKJG" authority="Mollendorff, 1898: 147" authorityName="Mollendorff" authorityPageNumber="147" authorityYear="1898" box="[189,635,1036,1063]" class="Gastropoda" family="Plectopylidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="family">
Plectopylidae
<bibRefCitation id="EFF44B476C2A8A21FE9CFBCC4572FB8E" author="Mollendorff O. F. von" box="[356,635,1036,1063]" pageId="6" pageNumber="7" pagination="25 - 208" refId="ref54802" refString="Mollendorff O. F. von 1898. Verzeichniss der auf den Philippinen lebenden Landmollusken. Abhandlungen der Naturforschenden Gesellschaft zu Gorlitz 22: 25 - 208." type="journal article" year="1898">Möllendorff, 1898: 147</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDA36B66C2A8A21FF45FB8E4643FBC1" blockId="6.[189,330,1102,1128]" box="[189,330,1102,1128]" pageId="6" pageNumber="7">
<heading id="D09281DA6C2A8A21FF45FB8E4643FBC1" bold="true" box="[189,330,1102,1128]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B911EAA46C2A8A21FF45FB8E4643FBC1" bold="true" box="[189,330,1102,1128]" pageId="6" pageNumber="7">
<typeStatus id="54DE88146C2A8A21FF45FB8E47F3FBC1" box="[189,250,1102,1128]" pageId="6" pageNumber="7">Type</typeStatus>
genus
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA36B66C2A8A21FF45FBBD46F9FB3E" blockId="6.[189,496,1149,1176]" box="[189,496,1149,1176]" pageId="6" pageNumber="7">
<taxonomicName id="4C654D356C2A8A21FF45FBBD46E2FB3E" ID-CoL="7PCTH" authority="Benson, 1860" authorityName="Benson" authorityYear="1860" box="[189,491,1149,1176]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="B911EAA46C2A8A21FF45FBBD4636FB3E" box="[189,319,1149,1175]" italics="true" pageId="6" pageNumber="7">Plectopylis</emphasis>
<bibRefCitation id="EFF44B476C2A8A21FEBFFBBE46E2FB3E" author="Benson W. H." box="[327,491,1149,1176]" pageId="6" pageNumber="7" pagination="243 - 247" refId="ref51737" refString="Benson W. H. 1860. Notes on Plectopylis, a group a Helicidae distinguished by several internal plicate Ephiphragms; with the Characters of a new Species. The Annals and Magazine of Natural History 3 (5): 243 - 247. Available from https: // biodiversitylibrary. org / page / 18658580 [accessed 30 Jul. 2018]." type="journal article" year="1860">Benson, 1860</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F653D6C2A8A20FF45FB7E45B3FDF4" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8BDA36B66C2A8A21FF45FB7E463FFB71" blockId="6.[189,310,1214,1240]" box="[189,310,1214,1240]" pageId="6" pageNumber="7">
<heading id="D09281DA6C2A8A21FF45FB7E463FFB71" bold="true" box="[189,310,1214,1240]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B911EAA46C2A8A21FF45FB7E463FFB71" bold="true" box="[189,310,1214,1240]" pageId="6" pageNumber="7">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA36B66C2A8A21FF45FB2D43A6F927" blockId="6.[189,1399,1261,1678]" pageId="6" pageNumber="7">
Shell sinistral or dextral, small to very large (
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); flat, depressed-conical or conical; dorsal surface concave, flat (even in those cases the protoconch usually protrudes above surface), conical or domed; protoconch sculpture variable between genera (ribbed, wrinkled, reticulated, tuberculated); dorsal surface usually with reticulated structure (consisting of spiral and radial lines); ventral surface usually with less prominent sculpture; deciduous periostracal folds might be present on the periphery of body whorl; aperture oblique to shell axis; peristome expanded, sometimes reflected; parietal callus present, V- or slightly S-shaped, apertural fold present or absent; multiple armatures formed during growth, the last one 0.250.75 whorls behind aperture; parietal wall with one or two vertical lamellae, usually with additional horizontal plicae above and below the lamellae or anterior to them; palatal wall with 67 plicae, which are variable in morphology: straight and horizontal, depressed Z-shaped, sometimes divided in the middle (most often when there is a single parietal lamella), or the fourth and fifth fuse to each other, forming a vertical plate (most often where there are two parietal lamellae).
</paragraph>
<paragraph id="8BDA36B66C2A8A20FF45F97546D5FEC5" blockId="6.[189,1399,1717,2027]" lastBlockId="7.[189,1399,267,364]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">Ommatophoral retractor (in sinistral specimens the left, in dextral specimens the right retractor) runs between the penis and vagina; atrium short; penis very short, vestigial to well-developed, without verge, usually with parallel folds internally; calcareous crystals may be present between folds; epiphallus usually present, bound to penis by weak fibres; penial caecum short or absent, retractor muscle inserts on penial caecum or, if caecum is absent, at the penis-epiphallus transition; vas deferens long, usually thickened before entering spermoviduct; vagina well-developed, usually U-shaped, sometimes with thickened vaginal bulb formed at turning point of U, attached to body wall with multiple, short muscle fibres; vagina internally with longitudinal folds; spermoviduct long; bursa copulatrix with slender stalk of variable length and usually an ovoid bursa; diverticulum usually present, short ovoid to very long, slender, usually originates near originating point of bursa copulatrix; all species ovoviviparous, embryo sac with several small calcareous crystals on its surface; talon normally developed, elongated, albumen gland of variable length.</paragraph>
<paragraph id="8BDA36B66C2B8A20FF45FE5345B3FDF4" blockId="7.[189,1399,403,606]" pageId="7" pageNumber="8">Radula elongated, but not very slender; teeth arranged in rows; laterals in straight rows perpendicular to central column; marginals in anteriorly pointed, slightly oblique rows, central tooth with a single, small to medium-sized cusp (sometimes absent), which is smaller or larger than ectocones of first laterals, but always smaller than endocones of first laterals; first laterals with a large endocone and a small ectocone; transition between laterals and marginals is gradual with endocone being gradually divided into two cusps, ectocones usually remain undivided.</paragraph>
</subSubSection>
<subSubSection id="C37F653D6C2B8A20FF45FD44467CFCF3" pageId="7" pageNumber="8" type="distribution">
<paragraph id="8BDA36B66C2B8A20FF45FD44465FFD37" blockId="7.[189,342,644,670]" box="[189,342,644,670]" pageId="7" pageNumber="8">
<heading id="D09281DA6C2B8A20FF45FD44465FFD37" bold="true" box="[189,342,644,670]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B911EAA46C2B8A20FF45FD44465FFD37" bold="true" box="[189,342,644,670]" pageId="7" pageNumber="8">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA36B66C2B8A20FF45FD73467CFCF3" blockId="7.[189,1399,691,859]" pageId="7" pageNumber="8">
The family
<taxonomicName id="4C654D356C2B8A20FEB0FD7346E1FD64" authorityName="Mollendorff" authorityYear="1898" box="[328,488,691,717]" class="Gastropoda" family="Plectopylidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="family">Plectopylidae</taxonomicName>
extends from the southern Himalayan region (
<collectingCountry id="F37276266C2B8A20FBEBFD734352FD64" box="[1043,1115,691,717]" name="Nepal" pageId="7" pageNumber="8">Nepal</collectingCountry>
and northeastern
<collectingCountry id="F37276266C2B8A20FAC8FD734267FD67" box="[1328,1390,691,718]" name="India" pageId="7" pageNumber="8">India</collectingCountry>
) throughout
<collectingCountry id="F37276266C2B8A20FEB0FD1746B5FD58" box="[328,444,727,753]" name="Myanmar" pageId="7" pageNumber="8">Myanmar</collectingCountry>
, the Malay Peninsula, southern and northern
<collectingCountry id="F37276266C2B8A20FC19FD164345FD59" box="[993,1100,726,752]" name="Thailand" pageId="7" pageNumber="8">Thailand</collectingCountry>
, northern
<collectingCountry id="F37276266C2B8A20FB31FD17420CFD58" box="[1225,1285,727,753]" name="Laos" pageId="7" pageNumber="8">Laos</collectingCountry>
, northern
<collectingCountry id="F37276266C2B8A20FF45FD39462AFCBA" box="[189,291,761,787]" name="Vietnam" pageId="7" pageNumber="8">Vietnam</collectingCountry>
, central, southern and eastern
<collectingCountry id="F37276266C2B8A20FD77FD3945DCFCBA" box="[655,725,761,787]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
(
<collectingRegion id="49A1F8546C2B8A20FD1EFD3A444CFCBD" box="[742,837,762,788]" country="China" name="Yunnan" pageId="7" pageNumber="8">Yunnan</collectingRegion>
,
<collectingRegion id="49A1F8546C2B8A20FCAAFD3944BAFCBA" box="[850,947,761,787]" country="China" name="Sichuan" pageId="7" pageNumber="8">Sichuan</collectingRegion>
,
<collectingRegion id="49A1F8546C2B8A20FC39FD394320FCBA" box="[961,1065,761,787]" country="China" name="Guizhou" pageId="7" pageNumber="8">Guizhou</collectingRegion>
,
<collectingRegion id="49A1F8546C2B8A20FBCEFD3943B2FCBA" box="[1078,1211,761,787]" country="China" name="Chongqing" pageId="7" pageNumber="8">Chongqing</collectingRegion>
,
<collectingRegion id="49A1F8546C2B8A20FB31FD39421DFCBA" box="[1225,1300,761,787]" country="China" name="Hubei" pageId="7" pageNumber="8">Hubei</collectingRegion>
,
<collectingRegion id="49A1F8546C2B8A20FAD9FD3A427BFCBD" box="[1313,1394,762,788]" country="China" name="Hunan" pageId="7" pageNumber="8">Hunan</collectingRegion>
,
<collectingRegion id="49A1F8546C2B8A20FF45FCDD462CFC9E" box="[189,293,797,823]" country="China" name="Guangxi" pageId="7" pageNumber="8">Guangxi</collectingRegion>
,
<collectingRegion id="49A1F8546C2B8A20FECAFCDD46B7FC9E" box="[306,446,797,823]" country="China" name="Guangdong" pageId="7" pageNumber="8">Guangdong</collectingRegion>
,
<collectingRegion id="49A1F8546C2B8A20FE32FCDD452BFC9E" box="[458,546,797,823]" country="China" name="Jiangxi" pageId="7" pageNumber="8">Jiangxi</collectingRegion>
,
<collectingRegion id="49A1F8546C2B8A20FDD6FCDD459FFC9E" box="[558,662,797,823]" country="China" name="Zhejiang" pageId="7" pageNumber="8">Zhejiang</collectingRegion>
and
<collectingRegion id="49A1F8546C2B8A20FD29FCDD4415FC9E" box="[721,796,797,823]" country="China" name="Fujian" pageId="7" pageNumber="8">Fujian</collectingRegion>
Provinces) to
<collectingCountry id="F37276266C2B8A20FC31FCDD4316FC9E" box="[969,1055,797,823]" name="Taiwan" pageId="7" pageNumber="8">Taiwan</collectingCountry>
and southern
<collectingCountry id="F37276266C2B8A20FB3EFCDD4200FC9E" box="[1222,1289,797,823]" name="Japan" pageId="7" pageNumber="8">Japan</collectingCountry>
(Miyako Island) (
<figureCitation id="135E2A336C2B8A20FEE6FC80466CFCF3" box="[286,357,832,858]" captionStart="Fig" captionStartId="7.[188,231,1930,1956]" captionTargetBox="[188,1399,1045,1890]" captionTargetId="figure@7.[186,1401,1042,1892]" captionTargetPageId="7" captionText="Fig. 3. Distribution of the extant Sinicolinae subfam. nov. (ruled area) and Plectopylinae Möllendorff, 1898 (grey shaded area). Mainly after Páll-Gergely &amp; Hunyadi (2013) and Páll-Gergely et al. (2015b, 2015c, 2016)." figureDoi="http://doi.org/10.5281/zenodo.3817713" httpUri="https://zenodo.org/record/3817713/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>