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<document id="860AEE72EF845B777521459FE6F749D8" ID-DOI="10.11646/zootaxa.3647.3.2" ID-GBIF-Dataset="235b8b0f-f1e2-4663-8522-6f40c1704588" ID-ISSN="1175-5326" ID-Zenodo-Dep="224235" ID-ZooBank="BF938988-855D-4596-93C8-09DA272341C5" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460605957267" checkinUser="plazi" docAuthor="Yasunaga, Tomohide, Yamada, Kazutaka &amp; Artchawakom, Taksin" docDate="2013" docId="03C787B4FFBA6C1EFF06FBA3FD04FD93" docLanguage="en" docName="zt03647p442.pdf" docOrigin="Zootaxa 3647 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alloeomimus muiri Schuh" docType="treatment" docVersion="9" lastPageNumber="437" masterDocId="FFFEFFCCFFBC6C16FF91FFB9FFE9FF84" masterDocTitle="New or little known taxa of the plant bug tribe Hallodapini (Hemiptera: Heteroptera: Miridae: Phylinae) from Thailand, with descriptions of three new species of the genus Acrorrhinium Noualhier" masterLastPageNumber="442" masterPageNumber="429" pageNumber="435" updateTime="1698327904738" updateUser="plazi">
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<mods:title id="C380274BB5A9C459FDB6B0C592F2A7D9">New or little known taxa of the plant bug tribe Hallodapini (Hemiptera: Heteroptera: Miridae: Phylinae) from Thailand, with descriptions of three new species of the genus Acrorrhinium Noualhier</mods:title>
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<mods:namePart id="70AC80CE155E55ABD4DE3CB82D1945A8">Yasunaga, Tomohide</mods:namePart>
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<mods:namePart id="B16AD6F4680B0382283D8831C3071A49">Yamada, Kazutaka</mods:namePart>
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<mods:namePart id="B3CA021FFF93E3498A56A4DDE93B6578">Artchawakom, Taksin</mods:namePart>
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<mods:title id="7E930C99B7B5373152430B0820513C1F">Zootaxa</mods:title>
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<mods:date id="76D9F566733FE525D68A6F8EE565F61E">2013</mods:date>
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<mods:number id="1E5AC2AE79BD3989406DD8D3A95067DC">3647</mods:number>
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<mods:identifier id="915EE38E337742FC1C4E61F457511508" type="DOI">10.11646/zootaxa.3647.3.2</mods:identifier>
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<treatment id="03C787B4FFBA6C1EFF06FBA3FD04FD93" ID-DOI="http://doi.org/10.5281/zenodo.6158907" ID-GBIF-Taxon="119567953" ID-Zenodo-Dep="6158907" LSID="urn:lsid:plazi:treatment:03C787B4FFBA6C1EFF06FBA3FD04FD93" httpUri="http://treatment.plazi.org/id/03C787B4FFBA6C1EFF06FBA3FD04FD93" lastPageId="8" lastPageNumber="437" pageId="6" pageNumber="435">
<subSubSection id="C3746529FFBA6C10FF06FBA3FE8AFBD1" pageId="6" pageNumber="435" type="nomenclature">
<paragraph id="8BD136A2FFBA6C10FF06FBA3FE3FFBB0" blockId="6.[151,470,1050,1109]" box="[151,470,1050,1076]" pageId="6" pageNumber="435">
<heading id="D09981CEFFBA6C10FF06FBA3FE3FFBB0" bold="true" box="[151,470,1050,1076]" fontSize="11" level="1" pageId="6" pageNumber="435" reason="1">
<taxonomicName id="4C6E4D21FFBA6C10FF06FBA3FE3FFBB0" ID-CoL="BWJ7" authority="Schuh" authorityName="Schuh" box="[151,470,1050,1076]" class="Insecta" family="Miridae" genus="Alloeomimus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="435" phylum="Arthropoda" rank="species" species="muiri">
<emphasis id="B91AEAB0FFBA6C10FF06FBA3FE3FFBB0" bold="true" box="[151,470,1050,1076]" pageId="6" pageNumber="435">
<emphasis id="B91AEAB0FFBA6C10FF06FBA3FE68FBB0" bold="true" box="[151,385,1050,1076]" italics="true" pageId="6" pageNumber="435">Alloeomimus muiri</emphasis>
Schuh
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD136A2FFBA6C10FF06FB84FE8AFBD1" blockId="6.[151,470,1050,1109]" box="[151,355,1085,1109]" pageId="6" pageNumber="435">
(
<figureCitation id="13552A27FFBA6C10FF0EFB84FF03FBD1" box="[159,234,1085,1109]" captionStart="FIGURE 3" captionStartId="7.[151,250,1678,1701]" captionTargetBox="[151,1435,195,1655]" captionTargetId="figure@7.[151,1436,195,1656]" captionTargetPageId="7" captionText="FIGURE 3. Habitus images of living adult individuals of hallodapines from Thailand. A. Alloeomimus muiri, male (AMNH _ PBI 00379623). B. Same, female (00379624). C. Clapmarius thailandana, male (00379627). D. Same, female (00379628). E. Hallodapus brunneus, male (SERS). F. H. albofasciatus, male (Nakhon Nayok). G. H. ravenar, male (Nakhon Nayok)." httpUri="https://zenodo.org/record/224237/files/figure.png" pageId="6" pageNumber="435">Figs. 3</figureCitation>
AB, 47)
</paragraph>
</subSubSection>
<subSubSection id="C3746529FFBA6C10FF06FB3CFCDCFA8D" pageId="6" pageNumber="435" type="diagnosis">
<paragraph id="8BD136A2FFBA6C10FF06FB3CFCDCFA8D" blockId="6.[151,1436,1157,1865]" pageId="6" pageNumber="435">
<emphasis id="B91AEAB0FFBA6C10FF06FB3CFEF8FB1A" bold="true" box="[151,273,1157,1182]" pageId="6" pageNumber="435">Diagnosis.</emphasis>
Recognized by the characters mentioned in generic diagnosis, macropterous female (see Remarks), two distinctive white maculae on the corium (anterior larger, triangular macula continuing to the clavus and posterior, wedge-like macula), small genital capsule with the apically tapered phallotheca (
<figureCitation id="13552A27FFBA6C10FBDAFB74FB7DFB61" box="[1099,1172,1229,1253]" captionStart="FIGURE 4" captionStartId="8.[151,250,1932,1955]" captionTargetBox="[196,1401,614,1882]" captionTargetId="figure@8.[151,1436,591,1911]" captionTargetPageId="8" captionText="FIGURE 4. Male genital segment (pygophore) and phallotheca of Thai hallodapines named on the figure. Scale bars 0.2 mm." httpUri="https://zenodo.org/record/224238/files/figure.png" pageId="6" pageNumber="435">Fig. 4</figureCitation>
), slender, simple male endosoma (
<figureCitation id="13552A27FFBA6C10FE89FB48FE6CFA8D" box="[280,389,1265,1289]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="9.[151,250,1754,1777]" captionStartId-1="10.[151,250,2014,2037]" captionTargetBox-0="[190,1378,212,1712]" captionTargetBox-1="[174,1392,603,1986]" captionTargetId-0="figure@9.[151,1436,177,1733]" captionTargetId-1="figure@10.[153,1438,575,2027]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURE 5. Male genitalia (left paramere and endosoma) of Thai hallodapines named on the figure. Scale bars 0.1 mm." captionText-1="FIGURE 6. Images of endosoma of Thai hallodapines named on the figure." httpUri-0="https://zenodo.org/record/224239/files/figure.png" httpUri-1="https://zenodo.org/record/224240/files/figure.png" pageId="6" pageNumber="435">Figs. 56</figureCitation>
), and narrow bursa copulatrix (
<figureCitation id="13552A27FFBA6C10FD73FB48FCCEFA8D" box="[738,807,1265,1289]" captionStart="FIGURE 7" captionStartId="11.[151,250,1469,1492]" captionTargetBox="[163,1390,237,1413]" captionTargetId="figure@11.[151,1436,175,1460]" captionTargetPageId="11" captionText="FIGURE 7. Images of female bursa copulatrix of Thai hallodapines named on the figure." httpUri="https://zenodo.org/record/224241/files/figure.png" pageId="6" pageNumber="435">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3746529FFBA6C10FF56FAACFC0EFA1D" pageId="6" pageNumber="435" type="description">
<paragraph id="8BD136A2FFBA6C10FF56FAACFC0EFA1D" blockId="6.[151,1436,1157,1865]" pageId="6" pageNumber="435">
<emphasis id="B91AEAB0FFBA6C10FF56FAACFE9FFAA9" bold="true" box="[199,374,1301,1325]" pageId="6" pageNumber="435">Measurements</emphasis>
. 3/Ƥ: Total body length 3.0/ 3.3; length from apex of clypeus to cuneal fracture 2.16/ 2.40; head width across eyes 0.60/ 0.62; vertex width 0.30/ 0.41; lengths of antennal segment IIV 0.27, 1.20, 1.05, 0.84/ 0.24, 1.32, 1.14, 0.75; labial length 1.56/ 1.53; basal pronotal width 0.75/ 0.75; width across hemelytron 0.74/ 0.90; and lengths of metafemur, tibia and tarsus 1.35, 2.10, 0.57/ 1.41, 2.13, 0.59.
</paragraph>
</subSubSection>
<subSubSection id="C3746529FFBA6C10FF56FA1CFDECF9F5" pageId="6" pageNumber="435" type="biology_ecology">
<paragraph id="8BD136A2FFBA6C10FF56FA1CFF01FA65" blockId="6.[151,1436,1157,1865]" pageId="6" pageNumber="435">
<emphasis id="B91AEAB0FFBA6C10FF56FA1CFEF6FA3A" bold="true" box="[199,287,1445,1470]" pageId="6" pageNumber="435">Biology</emphasis>
. No information is currently available, as all specimens used in this study were collected at light in March.
</paragraph>
<paragraph id="8BD136A2FFBA6C10FF56FA54FDECF9F5" blockId="6.[151,1436,1157,1865]" pageId="6" pageNumber="435">
<emphasis id="B91AEAB0FFBA6C10FF56FA54FD98F982" bold="true" box="[199,625,1517,1542]" pageId="6" pageNumber="435">
Specimens examined.
<collectingCountry id="F3797632FFBA6C10FE41FA54FD82F982" box="[464,619,1517,1542]" name="Thailand" pageId="6" pageNumber="435">THAILAND</collectingCountry>
:
</emphasis>
Nakhon Nayok Prov.: 13, Sarika (lodge),
<geoCoordinate id="EE5A5065FFBA6C10FBCDFA54FB08F981" box="[1116,1249,1517,1541]" direction="north" orientation="latitude" pageId="6" pageNumber="435" precision="15" value="14.310834">N14°18'39&quot;</geoCoordinate>
<geoCoordinate id="EE5A5065FFBA6C10FB78FA54FA92F981" box="[1257,1403,1517,1541]" direction="east" orientation="longitude" pageId="6" pageNumber="435" precision="15" value="101.3">E101°18'00&quot;</geoCoordinate>
, at light,
<date id="FFD01062FFBA6C10FF4AF9A8FE8FF9AD" box="[219,358,1553,1577]" pageId="6" pageNumber="435" value="2009-03-05">5 Mar 2009</date>
, T. Yasunaga (AMNH_PBI 00379623) (TYCN); 1Ƥ, same data except for date
<date id="FFD01062FFBA6C10FA97F9A8FA75F9AD" box="[1286,1436,1553,1577]" pageId="6" pageNumber="435" value="2009-03-14">14 Mar 2009</date>
(00379624) (
<collectingCountry id="F3797632FFBA6C10FEB8F98CFE8AF9C9" box="[297,355,1589,1613]" name="Spain" pageId="6" pageNumber="435">SUT</collectingCountry>
); 131Ƥ, Sarika,
<geoCoordinate id="EE5A5065FFBA6C10FDB3F98CFD4CF9C9" box="[546,677,1589,1613]" direction="north" orientation="latitude" pageId="6" pageNumber="435" precision="15" value="14.301945">N14°18'07&quot;</geoCoordinate>
<geoCoordinate id="EE5A5065FFBA6C10FD3CF98CFCD7F9C9" box="[685,830,1589,1613]" direction="east" orientation="longitude" pageId="6" pageNumber="435" precision="15" value="101.302505">E101°18'09&quot;</geoCoordinate>
, at light,
<date id="FFD01062FFBA6C10FC36F98CFB8FF9C9" box="[935,1126,1589,1613]" pageId="6" pageNumber="435" value="2010-03-21" valueMax="2010-03-23" valueMin="2010-03-21">2123 Mar 2010</date>
, T. Yasunaga &amp; K. Yamada (0037962500379626) (TYCN).
</paragraph>
</subSubSection>
<subSubSection id="C3746529FFBA6C11FF56F9C4FE95F8ED" lastPageId="7" lastPageNumber="436" pageId="6" pageNumber="435" type="discussion">
<paragraph id="8BD136A2FFBA6C10FF56F9C4FED4F8CD" blockId="6.[151,1436,1157,1865]" pageId="6" pageNumber="435">
<emphasis id="B91AEAB0FFBA6C10FF56F9C4FED2F911" bold="true" box="[199,315,1661,1685]" pageId="6" pageNumber="435">Remarks.</emphasis>
Although all specimens examined during this study correspond in overall appearance to the original description, the male endosoma is somewhat longer than that described and illustrated by Schuh (1984). Because the hallodapine endosoma may be sometimes variable in length and shape, the
<collectingCountry id="F3797632FFBA6C10FB86F97CFBA3F959" box="[1047,1098,1733,1757]" name="Thailand" pageId="6" pageNumber="435">Thai</collectingCountry>
population is herein regarded to be conspecific with typical
<taxonomicName id="4C6E4D21FFBA6C10FE65F953FDD8F885" box="[500,561,1770,1793]" class="Insecta" family="Miridae" genus="Alloeomimus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="435" phylum="Arthropoda" rank="species" species="muiri">
<emphasis id="B91AEAB0FFBA6C10FE65F953FDD8F885" box="[500,561,1770,1793]" italics="true" pageId="6" pageNumber="435">muiri</emphasis>
</taxonomicName>
(the
<typeStatus id="54D58800FFBA6C10FDE1F950FD3BF885" box="[624,722,1769,1793]" pageId="6" pageNumber="435" type="holotype">holotype</typeStatus>
male from Java). Both of two female adults examined in this study are macropterous (
<figureCitation id="13552A27FFBA6C10FE24F8B4FE13F8A1" box="[437,506,1805,1829]" captionStart="FIGURE 3" captionStartId="7.[151,250,1678,1701]" captionTargetBox="[151,1435,195,1655]" captionTargetId="figure@7.[151,1436,195,1656]" captionTargetPageId="7" captionText="FIGURE 3. Habitus images of living adult individuals of hallodapines from Thailand. A. Alloeomimus muiri, male (AMNH _ PBI 00379623). B. Same, female (00379624). C. Clapmarius thailandana, male (00379627). D. Same, female (00379628). E. Hallodapus brunneus, male (SERS). F. H. albofasciatus, male (Nakhon Nayok). G. H. ravenar, male (Nakhon Nayok)." httpUri="https://zenodo.org/record/224237/files/figure.png" pageId="6" pageNumber="435">Fig. 3</figureCitation>
B); however, Schuh (1984) mentioned that the
<typeStatus id="54D58800FFBA6C10FB8EF8B7FB96F8A2" box="[1055,1151,1806,1830]" pageId="6" pageNumber="435" type="paratype">paratype</typeStatus>
females (from Java) are brachypterous.
</paragraph>
<caption id="DF11662AFFBB6C11FF06F937FF03F884" httpUri="https://zenodo.org/record/224237/files/figure.png" pageId="7" pageNumber="436" targetBox="[151,1435,195,1655]" targetPageId="7">
<paragraph id="8BD136A2FFBB6C11FF06F937FF03F884" blockId="7.[151,1436,1678,1792]" pageId="7" pageNumber="436">
<emphasis id="B91AEAB0FFBB6C11FF06F937FEF3F927" bold="true" box="[151,282,1678,1701]" pageId="7" pageNumber="436">FIGURE 3.</emphasis>
Habitus images of living adult individuals of hallodapines from Thailand.
<emphasis id="B91AEAB0FFBB6C11FBF7F937FB96F927" bold="true" box="[1126,1151,1678,1699]" pageId="7" pageNumber="436">A.</emphasis>
<taxonomicName id="4C6E4D21FFBB6C11FB1CF937FABFF920" box="[1165,1366,1678,1700]" class="Insecta" family="Miridae" genus="Alloeomimus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="436" phylum="Arthropoda" rank="species" species="muiri">
<emphasis id="B91AEAB0FFBB6C11FB1CF937FABFF920" box="[1165,1366,1678,1700]" italics="true" pageId="7" pageNumber="436">Alloeomimus muiri</emphasis>
</taxonomicName>
, male (AMNH_PBI 00379623).
<emphasis id="B91AEAB0FFBB6C11FE3DF914FE2AF946" bold="true" box="[428,451,1709,1730]" pageId="7" pageNumber="436">B.</emphasis>
Same, female (00379624).
<emphasis id="B91AEAB0FFBB6C11FD60F915FCE0F947" bold="true" box="[753,777,1708,1731]" pageId="7" pageNumber="436">C.</emphasis>
<taxonomicName id="4C6E4D21FFBB6C11FC84F914FBFBF947" box="[789,1042,1709,1731]" class="Insecta" family="Miridae" genus="Clapmarius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="436" phylum="Arthropoda" rank="species" species="thailandana">
<emphasis id="B91AEAB0FFBB6C11FC84F914FBFBF947" box="[789,1042,1709,1731]" italics="true" pageId="7" pageNumber="436">Clapmarius thailandana</emphasis>
</taxonomicName>
, male (00379627).
<emphasis id="B91AEAB0FFBB6C11FB78F914FAE8F946" bold="true" box="[1257,1281,1709,1730]" pageId="7" pageNumber="436">D.</emphasis>
Same, female (00379628).
<emphasis id="B91AEAB0FFBB6C11FE8BF975FED8F965" bold="true" box="[282,305,1740,1761]" pageId="7" pageNumber="436">E.</emphasis>
<taxonomicName id="4C6E4D21FFBB6C11FEA9F975FDFAF966" box="[312,531,1740,1762]" class="Insecta" family="Miridae" genus="Hallodapus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="436" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B91AEAB0FFBB6C11FEA9F975FDFAF966" box="[312,531,1740,1762]" italics="true" pageId="7" pageNumber="436">Hallodapus brunneus</emphasis>
</taxonomicName>
, male (SERS).
<emphasis id="B91AEAB0FFBB6C11FD22F975FD2DF965" bold="true" box="[691,708,1740,1761]" pageId="7" pageNumber="436">F.</emphasis>
<taxonomicName id="4C6E4D21FFBB6C11FD5CF974FC9DF966" box="[717,884,1740,1762]" class="Insecta" family="Miridae" genus="Hallodapus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="436" phylum="Arthropoda" rank="species" species="albofasciatus">
<emphasis id="B91AEAB0FFBB6C11FD5CF974FC9DF966" box="[717,884,1740,1762]" italics="true" pageId="7" pageNumber="436">H. albofasciatus</emphasis>
</taxonomicName>
, male (Nakhon Nayok).
<emphasis id="B91AEAB0FFBB6C11FBE3F972FB6FF966" bold="true" box="[1138,1158,1739,1762]" pageId="7" pageNumber="436">G.</emphasis>
<taxonomicName id="4C6E4D21FFBB6C11FB1EF974FB17F966" box="[1167,1278,1741,1762]" class="Insecta" family="Miridae" genus="Hallodapus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="436" phylum="Arthropoda" rank="species" species="ravenar">
<emphasis id="B91AEAB0FFBB6C11FB1EF974FB17F966" box="[1167,1278,1741,1762]" italics="true" pageId="7" pageNumber="436">H. ravenar</emphasis>
</taxonomicName>
, male (Nakhon Nayok).
</paragraph>
</caption>
<paragraph id="8BD136A2FFBB6C11FF06F8E8FE95F8ED" blockId="7.[151,380,1872,1898]" box="[151,380,1872,1898]" pageId="7" pageNumber="436">
<taxonomicName id="4C6E4D21FFBB6C11FF06F8E8FE95F8ED" authority="Distant" authorityName="Distant" box="[151,380,1872,1898]" class="Insecta" family="Miridae" genus="Clapmarius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="436" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAB0FFBB6C11FF06F8E8FE95F8ED" bold="true" box="[151,380,1872,1898]" pageId="7" pageNumber="436">
<emphasis id="B91AEAB0FFBB6C11FF06F8E8FEF6F8EE" bold="true" box="[151,287,1873,1898]" italics="true" pageId="7" pageNumber="436">Clapmarius</emphasis>
Distant
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3746529FFBB6C1EFF06F821FC99FF50" lastPageId="8" lastPageNumber="437" pageId="7" pageNumber="436" type="diagnosis">
<paragraph id="8BD136A2FFBB6C1EFF06F821FC99FF50" blockId="7.[151,1437,1944,2006]" lastBlockId="8.[151,1436,151,535]" lastPageId="8" lastPageNumber="437" pageId="7" pageNumber="436">
<emphasis id="B91AEAB0FFBB6C11FF06F821FEF8F835" bold="true" box="[151,273,1944,1969]" pageId="7" pageNumber="436">Diagnosis.</emphasis>
Recognized by the elongate-oval body (nearly conventional form of the
<taxonomicName id="4C6E4D21FFBB6C11FBCFF820FB55F835" box="[1118,1212,1945,1969]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="436" phylum="Arthropoda" rank="family">Miridae</taxonomicName>
); brown-castaneous basic coloration; a mesal, longitudinal sulcus on vertex; lamellate, somewhat flattened antennal segment I; Cshaped, short but thick endosoma apically furnished with several sclerotized appendages (
<figureCitation id="13552A27FFB46C1EFB16FF2EFB1DFF2B" box="[1159,1268,151,175]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="9.[151,250,1754,1777]" captionStartId-1="10.[151,250,2014,2037]" captionTargetBox-0="[190,1378,212,1712]" captionTargetBox-1="[174,1392,603,1986]" captionTargetId-0="figure@9.[151,1436,177,1733]" captionTargetId-1="figure@10.[153,1438,575,2027]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURE 5. Male genitalia (left paramere and endosoma) of Thai hallodapines named on the figure. Scale bars 0.1 mm." captionText-1="FIGURE 6. Images of endosoma of Thai hallodapines named on the figure." httpUri-0="https://zenodo.org/record/224239/files/figure.png" httpUri-1="https://zenodo.org/record/224240/files/figure.png" pageId="8" pageNumber="437">Figs. 56</figureCitation>
); and basically toughened bursa copulatrix with distinct sclerotized ring (Fig. 8).
</paragraph>
</subSubSection>
<subSubSection id="C3746529FFB46C1EFF56FF66FD04FD93" pageId="8" pageNumber="437" type="discussion">
<paragraph id="8BD136A2FFB46C1EFF56FF66FD39FEBB" blockId="8.[151,1436,151,535]" pageId="8" pageNumber="437">
<emphasis id="B91AEAB0FFB46C1EFF56FF66FEA5FF7C" bold="true" box="[199,332,223,248]" pageId="8" pageNumber="437">Discussion.</emphasis>
<taxonomicName id="4C6E4D21FFB46C1EFEC8FF59FE34FF7C" box="[345,477,224,248]" class="Insecta" family="Miridae" genus="Clapmarius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAB0FFB46C1EFEC8FF59FE34FF7C" box="[345,477,224,248]" italics="true" pageId="8" pageNumber="437">Clapmarius</emphasis>
</taxonomicName>
is currently represented by four species, known from Peninsular
<collectingCountry id="F3797632FFB46C1EFB63FF66FAB2FF73" box="[1266,1371,223,247]" name="Malaysia" pageId="8" pageNumber="437">Malaysia</collectingCountry>
, East
<collectingCountry id="F3797632FFB46C1EFF06FEBDFF17FE98" box="[151,254,260,284]" name="Malaysia" pageId="8" pageNumber="437">Malaysia</collectingCountry>
(Borneo),
<collectingCountry id="F3797632FFB46C1EFEE8FEBDFE01FE98" box="[377,488,260,284]" name="Sri Lanka" pageId="8" pageNumber="437">Sri Lanka</collectingCountry>
and
<collectingCountry id="F3797632FFB46C1EFDB1FEBDFD6FFE98" box="[544,646,260,284]" name="Thailand" pageId="8" pageNumber="437">Thailand</collectingCountry>
, respectively, and two undescribed species additionally exist in Kuala Lumpur and Sabah (East
<collectingCountry id="F3797632FFB46C1EFE22FE9EFDF2FEBB" box="[435,539,295,319]" name="Malaysia" pageId="8" pageNumber="437">Malaysia</collectingCountry>
) (Schuh, 1984).
</paragraph>
<paragraph id="8BD136A2FFB46C1EFF56FEF5FD04FD93" blockId="8.[151,1436,151,535]" pageId="8" pageNumber="437">
Schuh also commented that the affinities of
<taxonomicName id="4C6E4D21FFB46C1EFD20FEF5FCDCFEE0" box="[689,821,332,356]" class="Insecta" family="Miridae" genus="Clapmarius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAB0FFB46C1EFD20FEF5FCDCFEE0" box="[689,821,332,356]" italics="true" pageId="8" pageNumber="437">Clapmarius</emphasis>
</taxonomicName>
within the
<taxonomicName id="4C6E4D21FFB46C1EFC22FEF5FBFCFEE0" box="[947,1045,332,356]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="437" phylum="Arthropoda" rank="subFamily" subFamily="Phylinae">Phylinae</taxonomicName>
can be determined by the flattened pronotal collar and sulcate vertex which may relate this genus to the group of genera including
<taxonomicName id="4C6E4D21FFB46C1EFA95FEC9FA75FE0C" box="[1284,1436,368,392]" class="Insecta" family="Miridae" genus="Acrorrhinium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAB0FFB46C1EFA95FEC9FA75FE0C" box="[1284,1436,368,392]" italics="true" pageId="8" pageNumber="437">Acrorrhinium</emphasis>
</taxonomicName>
within the
<taxonomicName id="4C6E4D21FFB46C1EFE9CFE2DFE7CFE28" box="[269,405,404,428]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="437" phylum="Arthropoda" rank="tribe" tribe="Hallodapini">Hallodapini</taxonomicName>
. However, the exact systematic position of
<taxonomicName id="4C6E4D21FFB46C1EFCE9FE2DFC15FE28" box="[888,1020,404,428]" class="Insecta" family="Miridae" genus="Clapmarius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAB0FFB46C1EFCE9FE2DFC15FE28" box="[888,1020,404,428]" italics="true" pageId="8" pageNumber="437">Clapmarius</emphasis>
</taxonomicName>
remains unclear, and requires further broader observations on larger numbers of specimens. On the basis of the form of the male endosoma and female bursa copulatrix,
<taxonomicName id="4C6E4D21FFB46C1EFEF0FE65FE0CFE70" box="[353,485,476,500]" class="Insecta" family="Miridae" genus="Clapmarius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAB0FFB46C1EFEF0FE65FE0CFE70" box="[353,485,476,500]" italics="true" pageId="8" pageNumber="437">Clapmarius</emphasis>
</taxonomicName>
is assumed to have the closest relationship with
<taxonomicName id="4C6E4D21FFB46C1EFBB8FE65FAC1FE70" authority="Schuh" authorityName="Schuh" box="[1065,1320,476,500]" class="Insecta" family="Miridae" genus="Cleotomiroides" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAB0FFB46C1EFBB8FE65FB3CFE70" box="[1065,1237,476,500]" italics="true" pageId="8" pageNumber="437">Cleotomiroides</emphasis>
Schuh
</taxonomicName>
, currently placed in the tribe
<taxonomicName id="4C6E4D21FFB46C1EFEF6FE46FD00FD93" authority="(Yasunaga, 2012a)" baseAuthorityName="Yasunaga" baseAuthorityYear="2012" box="[359,745,511,535]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="437" phylum="Arthropoda" rank="tribe" tribe="Auricillocorini">Auricillocorini (Yasunaga, 2012a)</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>