269 lines
29 KiB
XML
269 lines
29 KiB
XML
<document id="5DDF9E2EAFEC57F4352C30F1F3779D37" ID-DOI="10.5281/zenodo.7666001" ID-ISSN="2305-2562" ID-Zenodo-Dep="7666001" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1677088844807" checkinUser="felipe" docAuthor="Plisko, Jadwiga Danuta" docDate="2002" docId="03C787ACBD1B57429597FD14FCC6BE61" docLanguage="en" docName="AfrInvertebr.43.1.183–203.pdf" docOrigin="African Invertebrates 43" docStyle="DocumentStyle:C20B10C002986E51083406A4EACF003A.3:AfrInvertebr.2001-2007.journal_article" docStyleId="C20B10C002986E51083406A4EACF003A" docStyleName="AfrInvertebr.2001-2007.journal_article" docStyleVersion="3" docTitle="Proandricus richerti Plisko 2002, sp. n." docType="treatment" docVersion="3" lastPageNumber="199" masterDocId="FFFEFFD4BD145753946DFF97FF8ABF06" masterDocTitle="Nine new earthworm species of Proandricus Plisko, 1992 from South Africa and Lesotho (Oligochaeta: Microchaetidae)" masterLastPageNumber="204" masterPageNumber="183" pageNumber="198" updateTime="1699455156175" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="172DB3A82CBCC738DE12CCDD014B08C2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="13ED6239E143A7AF949DF49294C561B0">
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<mods:title id="F5F44995953597B380BB138787095CDB">Nine new earthworm species of Proandricus Plisko, 1992 from South Africa and Lesotho (Oligochaeta: Microchaetidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="11E42100B12F751E52D10E4B50BE2F7B" type="personal">
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<mods:role id="68DE098C7BC1EAE339015EF08800037A">
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<mods:roleTerm id="3B03AC76BB581A4552A59A8A4EB5D358">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5A2228C96B2DA95ABB693A63EE80A7DA">Plisko, Jadwiga Danuta</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="4A035C8D21794BB1519860F693BE7123">text</mods:typeOfResource>
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<mods:relatedItem id="AA1787327D8DCD5E6D9297D7154D9BE6" type="host">
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<mods:titleInfo id="5A8E9BFF7499C3C4DA733114FAFB12CF">
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<mods:title id="4D19DCA99C394ACD826B0BBA80DD936A">African Invertebrates</mods:title>
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</mods:titleInfo>
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<mods:part id="F7D47D6DA210A07C6B4DF88329FB1D08">
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<mods:date id="EF182539F81103FFD24DE0DC9108C777">2002</mods:date>
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<mods:detail id="414715D45D41F8AB6FFC1FCC8A0E7E92" type="pubDate">
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<mods:number id="CDBEA90D52FDF0942FFA3166544BF6F6">2002-12-31</mods:number>
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</mods:detail>
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<mods:detail id="F67E9952BF276D9EF1E5FAC88FAAAF5C" type="volume">
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<mods:number id="BA4A3D48F3B9DB3B3B8371E8E58BF976">43</mods:number>
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</mods:detail>
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<mods:extent id="7B377741149FBE3F1E315995D8CDD440" unit="page">
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<mods:start id="73FE9AA45CE05B6BF40A1B7DFD54D926">183</mods:start>
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<mods:end id="247EF2A61472A0A803638722EB649C7B">204</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="CC40E6EC203DF00F897361FE9FA9A90C">journal article</mods:classification>
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<mods:identifier id="5FFAF3F3F62D672C275D42519CB6ADDF" type="DOI">10.5281/zenodo.7666001</mods:identifier>
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<mods:identifier id="4552CE4D675E602B2EED234E0F4F8C67" type="ISSN">2305-2562</mods:identifier>
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<mods:identifier id="8660112022B335CFD6B77EA96AD6B4AD" type="Zenodo-Dep">7666001</mods:identifier>
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</mods:mods>
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<treatment id="03C787ACBD1B57429597FD14FCC6BE61" ID-DOI="http://doi.org/10.5281/zenodo.7666045" ID-Zenodo-Dep="7666045" LSID="urn:lsid:plazi:treatment:03C787ACBD1B57429597FD14FCC6BE61" httpUri="http://treatment.plazi.org/id/03C787ACBD1B57429597FD14FCC6BE61" lastPageId="17" lastPageNumber="199" pageId="15" pageNumber="198">
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<subSubSection id="C3746531BD1B575C9597FD14FCBBBD9B" box="[506,817,643,669]" pageId="15" pageNumber="198" type="nomenclature">
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<paragraph id="8BD136BABD1B575C9597FD14FCBBBD9B" blockId="15.[181,1142,643,1669]" box="[506,817,643,669]" pageId="15" pageNumber="198">
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<heading id="D09981D6BD1B575C9597FD14FCBBBD9B" box="[506,817,643,669]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="198" reason="2">
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<taxonomicName id="4C6E4D39BD1B575C9597FD14FD64BD9A" ID-CoL="854HF" authority="Plisko, 2002" authorityName="Plisko" authorityYear="2002" box="[506,750,643,668]" class="Clitellata" family="Microchaetidae" genus="Proandricus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="15" pageNumber="198" phylum="Annelida" rank="species" species="richerti" status="sp. nov.">
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<emphasis id="B91AEAA8BD1B575C9597FD14FD64BD9A" bold="true" box="[506,750,643,668]" pageId="15" pageNumber="198">Proandricus richerti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22957D3BD1B575C9698FD13FCBBBD9B" box="[757,817,644,669]" pageId="15" pageNumber="198" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3746531BD1B57429606FD3BFCC6BE61" lastPageId="17" lastPageNumber="200" pageId="15" pageNumber="198" type="description">
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<paragraph id="8BD136BABD1B575C9606FD3BFD4ABDC3" blockId="15.[181,1142,643,1669]" box="[619,704,684,709]" pageId="15" pageNumber="198">
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||
(
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<figureCitation id="13552A3FBD1B575C9619FD3BFD3DBDC3" box="[628,695,684,709]" captionStart="Figs 8–9" captionStartId="13.[182,221,1623,1643]" captionTargetBox="[186,1130,1298,1603]" captionTargetId="figure-9@13.[185,1130,1283,1631]" captionTargetPageId="13" captionText="Figs 8–9. Proandricus species. Clitellar region, ventral view. 8. P. michelleae sp. n. 9. P. richerti sp. n. [C= clitellum; T = tuberculum pubertatis.]" figureDoi="http://doi.org/10.5281/zenodo.7666020" httpUri="https://zenodo.org/record/7666020/files/figure.png" pageId="15" pageNumber="198">Fig. 9</figureCitation>
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)
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</paragraph>
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<paragraph id="8BD136BABD1B575C94D8FD43FC6ABC0B" blockId="15.[181,1142,643,1669]" pageId="15" pageNumber="198">
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Etymology: Named for Mr S. Richert of the
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<collectingRegion id="49AAF858BD1B575C96A7FD43FCF6BDEB" box="[714,892,724,749]" country="South Africa" name="KwaZulu-Natal" pageId="15" pageNumber="198">KwaZulu-Natal</collectingRegion>
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Nature Conservation Service, who assisted Dr A. J. Armstrong in collecting the
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<typeStatus id="54D58818BD1B575C9728FD63FCFFBC0B" box="[837,885,756,781]" pageId="15" pageNumber="198">type</typeStatus>
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material.
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</paragraph>
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<paragraph id="8BD136BABD1B575C94D8FC8BFCB2BCB3" blockId="15.[181,1142,643,1669]" pageId="15" pageNumber="198">
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<materialsCitation id="3B063CE7BD1B575C94D8FC8BFCB2BCB3" collectingDate="2001-06" collectingDateMax="2001-12-31" collectingDateMin="2001-06" collectionCode="NMSA" collectorName="Olig. & A. J. Armstrong & S. Richert." country="South Africa" latitude="-29.561607" location="Vergelegen Nature Reserve" longLatPrecision="1" longitude="29.49349" municipality="Material" pageId="15" pageNumber="198" specimenCount="13" specimenCount-juvenile="13" stateProvince="KwaZulu-Natal" typeStatus="holotype">
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<collectingMunicipality id="6BB5ACC0BD1B575C94D8FC8BFE9EBC33" box="[181,276,796,821]" pageId="15" pageNumber="198">Material</collectingMunicipality>
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examined:
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<collectingCountry id="F379762ABD1B575C95F5FC8BFDECBC33" box="[408,614,796,821]" name="South Africa" pageId="15" pageNumber="198">SOUTH AFRICA</collectingCountry>
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:
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<emphasis id="B91AEAA8BD1B575C961CFC8BFC95BC32" box="[625,799,796,820]" italics="true" pageId="15" pageNumber="198">
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<collectingRegion id="49AAF858BD1B575C961CFC8BFC95BC32" box="[625,799,796,820]" country="South Africa" name="KwaZulu-Natal" pageId="15" pageNumber="198">KwaZulu-Natal</collectingRegion>
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</emphasis>
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:
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<typeStatus id="54D58818BD1B575C9742FC8BFC1DBC33" box="[815,919,796,821]" pageId="15" pageNumber="198" type="holotype">Holotype</typeStatus>
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<collectionCode id="ED7FAE7FBD1B575C97F3FC8BFC7BBC33" box="[926,1009,796,821]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="15" pageNumber="198" type="Museum">NMSA</collectionCode>
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/Olig.03540 clitellate,
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<location id="8EB16061BD1B575C9573FCABFDCCBC53" LSID="urn:lsid:plazi:treatment:03C787ACBD1B57429597FD14FCC6BE61:8EB16061BD1B575C9573FCABFDCCBC53" box="[286,582,828,853]" country="South Africa" latitude="-29.561607" longLatPrecision="1" longitude="29.49349" municipality="Material" name="Vergelegen Nature Reserve" pageId="15" pageNumber="198" stateProvince="KwaZulu-Natal">Vergelegen Nature Reserve</location>
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(
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<geoCoordinate id="EE5A507DBD1B575C963EFCABFC88BC53" box="[595,770,828,853]" degrees="29" direction="south" minutes="33" orientation="latitude" pageId="15" pageNumber="198" precision="1" seconds="41.786" value="-29.561607">29º33'41.786"S</geoCoordinate>
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:
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<geoCoordinate id="EE5A507DBD1B575C9767FCABFC30BC53" box="[778,954,828,853]" degrees="29" direction="east" minutes="29" orientation="longitude" pageId="15" pageNumber="198" precision="1" seconds="36.563" value="29.49349">29º29'36.563"E</geoCoordinate>
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), plateau covered by grass and herbs, burnt in
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<date id="FFD0107ABD1B575C9586FCCBFDEABC73" box="[491,608,860,885]" pageId="15" pageNumber="198" value="2001-06">
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<collectingDate id="EF94E992BD1B575C9586FCCBFDEABC73" box="[491,608,860,885]" pageId="15" pageNumber="198" value="2001-06">June 2001</collectingDate>
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</date>
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, in moist soil at depth of
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<quantity id="4C969B5FBD1B575C971EFCCBFC5CBC73" box="[883,982,860,885]" metricMagnitude="-1" metricUnit="m" metricValue="1.55" metricValueMax="3.0" metricValueMin="0.1" pageId="15" pageNumber="198" unit="cm" value="15.5" valueMax="30.0" valueMin="1.0">1–30 cm</quantity>
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,
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<date id="FFD0107ABD1B575C978DFCCBFF7ABC93" pageId="15" pageNumber="198" value="2001-12-31">
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<collectingDate id="EF94E992BD1B575C978DFCCBFF7ABC93" pageId="15" pageNumber="198" value="2001-12-31">31 December 2001</collectingDate>
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</date>
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.
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<typeStatus id="54D58818BD1B575C9490FCEBFEE0BC93" box="[253,362,892,917]" pageId="15" pageNumber="198" type="paratype">Paratypes</typeStatus>
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<collectionCode id="ED7FAE7FBD1B575C951FFCEBFE4CBC93" box="[370,454,892,917]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="15" pageNumber="198" type="Museum">NMSA</collectionCode>
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/
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<collectorName id="269B536CBD1B575C95A2FCEBFD8FBC93" box="[463,517,892,917]" pageId="15" pageNumber="198">Olig.</collectorName>
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03541 3 clitellate and
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<specimenCount id="9D68FD33BD1B575C976EFCEBFC07BC93" box="[771,909,892,917]" pageId="15" pageNumber="198" type="juvenile">13 juveniles</specimenCount>
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found together with
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<typeStatus id="54D58818BD1B575C94D8FC0BFE91BCB3" box="[181,283,924,949]" pageId="15" pageNumber="198" type="holotype">holotype</typeStatus>
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||
; all collected by
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<collectorName id="269B536CBD1B575C95BAFC0BFD03BCB3" box="[471,649,924,949]" pageId="15" pageNumber="198">A. J. Armstrong</collectorName>
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and
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<collectorName id="269B536CBD1B575C96ADFC0BFCB2BCB3" box="[704,824,924,949]" pageId="15" pageNumber="198">S. Richert.</collectorName>
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</materialsCitation>
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</paragraph>
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<paragraph id="8BD136BABD1B575C94D8FC53FE1FBCDB" blockId="15.[181,1142,643,1669]" box="[181,405,964,989]" pageId="15" pageNumber="198">External characters:</paragraph>
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<paragraph id="8BD136BABD1B575C94D8FC7BFEC5B903" blockId="15.[181,1142,643,1669]" pageId="15" pageNumber="198">
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<emphasis id="B91AEAA8BD1B575C94D8FC7BFE9BBB02" box="[181,273,1004,1028]" italics="true" pageId="15" pageNumber="198">General</emphasis>
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: Body cylindrical, extended laterally in area of tubercula pubertatis.
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<emphasis id="B91AEAA8BD1B575C9073FC7BFBE7BB02" box="[1054,1133,1004,1028]" italics="true" pageId="15" pageNumber="198">Colour</emphasis>
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: In life dorsally violet in preclitellar part, ventrally yellowish-white. Alcohol-preserved dorsally with violet tint, ventrally yellowish-grey.
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<emphasis id="B91AEAA8BD1B575C9692FBBBFC09BB42" box="[767,899,1068,1092]" italics="true" pageId="15" pageNumber="198">Dimensions</emphasis>
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:
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<typeStatus id="54D58818BD1B575C97F8FBBBFC77BB43" box="[917,1021,1068,1093]" pageId="15" pageNumber="198" type="holotype">Holotype</typeStatus>
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preserved slightly contracted, abscised 125+ mm long,
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<quantity id="4C969B5FBD1B575C96FBFBDBFD5EBB63" box="[662,724,1100,1125]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="15" pageNumber="198" unit="mm" value="7.0">7 mm</quantity>
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wide at segment
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<quantity id="4C969B5FBD1B575C97E0FBDBFC78BB63" box="[909,1010,1100,1125]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="15" pageNumber="198" unit="mm" value="10.9">10, 9 mm</quantity>
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at tubercula pubertatis.
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<typeStatus id="54D58818BD1B575C9559FBFBFE2FBB83" box="[308,421,1132,1157]" pageId="15" pageNumber="198" type="paratype">Paratypes</typeStatus>
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: Clitellate, abscised,
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<emphasis id="B91AEAA8BD1B575C96F5FBFBFD38BB82" box="[664,690,1132,1156]" italics="true" pageId="15" pageNumber="198">ca</emphasis>
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.
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<quantity id="4C969B5FBD1B575C96AFFBFBFCB4BB83" box="[706,830,1132,1157]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="15" pageNumber="198" unit="mm" value="60.0" valueMax="80.0" valueMin="40.0">40–80 mm</quantity>
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long; juvenile
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<quantity id="4C969B5FBD1B575C9780FBFBFBFCBB83" box="[1005,1142,1132,1157]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" metricValueMax="2.3" metricValueMin="0.3" pageId="15" pageNumber="198" unit="mm" value="130.0" valueMax="230.0" valueMin="30.0">30–230 mm</quantity>
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long.
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<emphasis id="B91AEAA8BD1B575C9493FB1BFE43BBA2" box="[254,457,1164,1188]" italics="true" pageId="15" pageNumber="198">Segment number</emphasis>
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: Abscised
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<typeStatus id="54D58818BD1B575C9638FB1BFD37BBA3" box="[597,701,1164,1189]" pageId="15" pageNumber="198" type="holotype">holotype</typeStatus>
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||
255+, juvenile
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<typeStatus id="54D58818BD1B575C97EAFB1BFC73BBA3" box="[903,1017,1164,1189]" pageId="15" pageNumber="198" type="paratype">paratypes</typeStatus>
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||
360–489.
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||
<emphasis id="B91AEAA8BD1B575C94D8FB3BFEB4BBC2" box="[181,318,1196,1220]" italics="true" pageId="15" pageNumber="198">Prostomium</emphasis>
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: Prolobous, moderate.
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<emphasis id="B91AEAA8BD1B575C9622FB3BFD62BBC2" box="[591,744,1196,1220]" italics="true" pageId="15" pageNumber="198">Segmentation</emphasis>
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: Secondary annulation present on preclitellar segments; 1 and 2 clearly separated one from another, short, simple, both with irregular longitudinal grooves; 3 simple, longer than 1 or 2; 4–8 with two simple ringlets, similar in size and appearance; 9 with two ringlets, second shorter than first; 10–12 simple; postclitellar simple, randomly annulated.
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<emphasis id="B91AEAA8BD1B575C9777FABBFCDCBA42" box="[794,854,1324,1348]" italics="true" pageId="15" pageNumber="198">Setae</emphasis>
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||
: Closely paired; moderate in size; first pairs on 3; on 4–9 on first ringlet; on postclitellar segments
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<emphasis id="B91AEAA8BD1B575C978DFADBFC76BA62" box="[992,1020,1356,1380]" italics="true" pageId="15" pageNumber="198">aa</emphasis>
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>
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<emphasis id="B91AEAA8BD1B575C9077FADBFBBEBA62" box="[1050,1076,1356,1380]" italics="true" pageId="15" pageNumber="198">bc</emphasis>
|
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,
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<emphasis id="B91AEAA8BD1B575C902EFADBFBD5BA62" box="[1091,1119,1356,1380]" italics="true" pageId="15" pageNumber="198">ab</emphasis>
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=
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<emphasis id="B91AEAA8BD1B575C94D8FAFBFF45BA82" box="[181,207,1388,1412]" italics="true" pageId="15" pageNumber="198">cd</emphasis>
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.
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<emphasis id="B91AEAA8BD1B575C948DFAFBFE2ABA82" box="[224,416,1388,1412]" italics="true" pageId="15" pageNumber="198">Nephridial pores</emphasis>
|
||
: Conspicuous, in
|
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<emphasis id="B91AEAA8BD1B575C961FFAFBFD06BA82" box="[626,652,1388,1412]" italics="true" pageId="15" pageNumber="198">cd</emphasis>
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setal lines; first pair in 2/3 intersegmental furrow.
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<emphasis id="B91AEAA8BD1B575C957CFA1BFE21BAA2" box="[273,427,1420,1444]" italics="true" pageId="15" pageNumber="198">Female pores</emphasis>
|
||
: Not observed.
|
||
<emphasis id="B91AEAA8BD1B575C960AFA1BFD6DBAA2" box="[615,743,1420,1444]" italics="true" pageId="15" pageNumber="198">Male pores</emphasis>
|
||
: Probably in 17/18 intersegmental furrow in area of tubercula pubertatis, where there are small depressions.
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<emphasis id="B91AEAA8BD1B575C97B6FA3BFF78BAE2" italics="true" pageId="15" pageNumber="198">Spermathecal pores</emphasis>
|
||
: Externally not detected; during dissection found in intersegmental furrows 11/ 12 and 12/13.
|
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</paragraph>
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<paragraph id="8BD136BABD1B574394BDF99BFF57BE01" blockId="15.[181,1142,643,1669]" lastBlockId="16.[144,1104,206,1671]" lastPageId="16" lastPageNumber="199" pageId="15" pageNumber="198">
|
||
Clitellar region (
|
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<figureCitation id="13552A3FBD1B575C95E2F99BFE5DB923" box="[399,471,1548,1573]" captionStart="Figs 8–9" captionStartId="13.[182,221,1623,1643]" captionTargetBox="[186,1130,1298,1603]" captionTargetId="figure-9@13.[185,1130,1283,1631]" captionTargetPageId="13" captionText="Figs 8–9. Proandricus species. Clitellar region, ventral view. 8. P. michelleae sp. n. 9. P. richerti sp. n. [C= clitellum; T = tuberculum pubertatis.]" figureDoi="http://doi.org/10.5281/zenodo.7666020" httpUri="https://zenodo.org/record/7666020/files/figure.png" pageId="15" pageNumber="198">Fig. 9</figureCitation>
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):
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<emphasis id="B91AEAA8BD1B575C9583F99BFDDCB922" box="[494,598,1548,1572]" italics="true" pageId="15" pageNumber="198">Clitellum</emphasis>
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: On
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||
<typeStatus id="54D58818BD1B575C96FEF99BFD7FB923" box="[659,757,1548,1573]" pageId="15" pageNumber="198" type="holotype">holotype</typeStatus>
|
||
and mature
|
||
<typeStatus id="54D58818BD1B575C97EAF99BFC78B923" box="[903,1010,1548,1573]" pageId="15" pageNumber="198" type="paratype">paratypes</typeStatus>
|
||
marked by brownish colour on segments 13–23; ventral edges in line of tubercula pubertatis.
|
||
<emphasis id="B91AEAA8BD1B575C94D8F9DBFE2BB962" box="[181,417,1612,1636]" italics="true" pageId="15" pageNumber="198">Tubercula pubertatis</emphasis>
|
||
: On
|
||
<typeStatus id="54D58818BD1B575C95B3F9DBFDCAB963" box="[478,576,1612,1637]" pageId="15" pageNumber="198" type="holotype">holotype</typeStatus>
|
||
large, rectangular pads with rounded corners, on 1/n16–20; rimmed at dorsal borders; on
|
||
<typeStatus id="54D58818BD1B575C96E0F9FBFD72B983" box="[653,760,1644,1669]" pageId="15" pageNumber="198" type="paratype">paratypes</typeStatus>
|
||
markedly less than on
|
||
<typeStatus id="54D58818BD1B575C9061F9FBFBFBB983" box="[1036,1137,1644,1669]" pageId="15" pageNumber="198" type="holotype">holotype</typeStatus>
|
||
, rimmed dorsally.
|
||
<emphasis id="B91AEAA8BD045743953DFF59FE25BFE0" box="[336,431,206,230]" italics="true" pageId="16" pageNumber="199">Papillae</emphasis>
|
||
: Very small tubercles in line of
|
||
<emphasis id="B91AEAA8BD0457439764FF59FCAFBFE0" box="[777,805,206,230]" italics="true" pageId="16" pageNumber="199">ab</emphasis>
|
||
setae on variable segments 10–12.
|
||
</paragraph>
|
||
<paragraph id="8BD136BABD04574394FDFE81FEE2BE29" blockId="16.[144,1104,206,1671]" box="[144,360,278,303]" pageId="16" pageNumber="199">Internal characters:</paragraph>
|
||
<paragraph id="8BD136BABD04574394FDFEA9FCC0BD91" blockId="16.[144,1104,206,1671]" pageId="16" pageNumber="199">
|
||
<emphasis id="B91AEAA8BD04574394FDFEA9FF44BE50" box="[144,206,318,342]" italics="true" pageId="16" pageNumber="199">Septa</emphasis>
|
||
: 4/5 5/6 6/7 very thin; 7/8 and 8/9 much thickened, muscular, firm, similar in size and appearance.
|
||
<emphasis id="B91AEAA8BD0457439525FEC9FE2ABE70" box="[328,416,350,374]" italics="true" pageId="16" pageNumber="199">Gizzard</emphasis>
|
||
: In 7, elongated, muscular.
|
||
<emphasis id="B91AEAA8BD04574396BDFEC9FC28BE70" box="[720,930,350,374]" italics="true" pageId="16" pageNumber="199">Calciferous glands</emphasis>
|
||
: In 9; moderate in size, laterally; separated dorsally and ventrally.
|
||
<emphasis id="B91AEAA8BD045743968DFEE9FCCBBE90" box="[736,833,382,406]" italics="true" pageId="16" pageNumber="199">Intestine</emphasis>
|
||
: Commences in 13; in segments 23–28 paired elongated diverticula.
|
||
<emphasis id="B91AEAA8BD04574396C9FE09FC97BEB0" box="[676,797,414,438]" italics="true" pageId="16" pageNumber="199">Typhlosole</emphasis>
|
||
: Commences immediately with intestine as thin tube, gradually enlarging and becoming U-shaped, thick structure; in abscised
|
||
<typeStatus id="54D58818BD045743957CFE49FEF9BEF1" box="[273,371,478,503]" pageId="16" pageNumber="199" type="holotype">holotype</typeStatus>
|
||
terminates in segment 178; in juvenile
|
||
<typeStatus id="54D58818BD0457439744FE49FC03BEF1" box="[809,905,478,503]" pageId="16" pageNumber="199" type="paratype">paratype</typeStatus>
|
||
consisting of 489 segments and terminates in 180, leaving over 300 segments with no typhlosole.
|
||
<emphasis id="B91AEAA8BD045743906FFE69FE93BD30" italics="true" pageId="16" pageNumber="199">Dorsal blood vessel</emphasis>
|
||
: In segments 4–7 double in posterior parts, separated; in 8 double, widely separated; in 9 double, large, cordiform; single when crossing septa, in 10 and the following segments; 4–8 thin tubes; in 9–11 enlarged, moniliform.
|
||
<taxonomicName id="4C6E4D39BD04574397A2FDC9FEB7BD91" authority=": Meganephridia" authorityName="Meganephridia" class="Insecta" family="Crabronidae" genus="Nephridia" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="199" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91AEAA8BD04574397A2FDC9FBCDBD70" box="[975,1095,606,630]" italics="true" pageId="16" pageNumber="199">Nephridia</emphasis>
|
||
: Meganephridia
|
||
</taxonomicName>
|
||
; coiled loops with short, thick V-shaped caeca.
|
||
</paragraph>
|
||
<paragraph id="8BD136BABD04574394C7FD09FCF2BB91" blockId="16.[144,1104,206,1671]" pageId="16" pageNumber="199">
|
||
Reproductive organs:
|
||
<emphasis id="B91AEAA8BD04574395F3FD09FD00BDB0" box="[414,650,670,694]" italics="true" pageId="16" pageNumber="199">Spermiductal funnels</emphasis>
|
||
: Proandric arrangement (in segment 10); in
|
||
<typeStatus id="54D58818BD04574394C0FD29FE85BDD1" box="[173,271,702,727]" pageId="16" pageNumber="199" type="holotype">holotype</typeStatus>
|
||
moderate in size, iridescent. Also in
|
||
<typeStatus id="54D58818BD04574396D9FD29FC9EBDD1" box="[692,788,702,727]" pageId="16" pageNumber="199" type="paratype">paratype</typeStatus>
|
||
with clitellum marked only by brown colour, a trace of iridescence was observed in small spermiductal funnels; both funnels closely connected with seminal sacs at posterior part of segment 10.
|
||
<emphasis id="B91AEAA8BD0457439076FD69FE8ABC30" italics="true" pageId="16" pageNumber="199">
|
||
<taxonomicName id="4C6E4D39BD0457439076FD69FBDABC10" authorityName="Agassiz" authorityYear="1842" box="[1051,1104,766,790]" class="Aves" genus="Vasa" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="199" phylum="Chordata" rank="genus">Vasa</taxonomicName>
|
||
deferentia
|
||
</emphasis>
|
||
: Single ducts commence at dorsal parts of spermiductal funnels, and run backwards almost medially at each side of the body to segment 17, where they enter body wall.
|
||
<emphasis id="B91AEAA8BD0457439567FCC9FE4BBC70" box="[266,449,862,886]" italics="true" pageId="16" pageNumber="199">Seminal vesicles</emphasis>
|
||
: One pair of moderate sacs, anteriorly connected at septum 10/11 with spermiductal funnels, extending backward to segments 11 and 12; in
|
||
<typeStatus id="54D58818BD0457439782FCE9FBDABC91" box="[1007,1104,894,919]" pageId="16" pageNumber="199" type="holotype">holotype</typeStatus>
|
||
both vesicles are elongated and of equal size, but when crossing septum 11/12 becoming narrower; in
|
||
<typeStatus id="54D58818BD0457439570FC29FEF7BCD1" box="[285,381,958,983]" pageId="16" pageNumber="199" type="paratype">paratype</typeStatus>
|
||
both vesicles smaller than in
|
||
<typeStatus id="54D58818BD04574396D2FC29FCAEBCD1" box="[703,804,958,983]" pageId="16" pageNumber="199" type="holotype">holotype</typeStatus>
|
||
, extending equally through segments 11 and 12 (probably the differences in size and shape can be accounted to different states of maturity).
|
||
<emphasis id="B91AEAA8BD04574395A1FC69FDE1BB10" box="[460,619,1022,1046]" italics="true" pageId="16" pageNumber="199">Spermathecae</emphasis>
|
||
: Close to septa 11/12 and 12/13; variable in shape and size, very small, bent ampullae; 3–4 at each side in segments 12 and 13.
|
||
<emphasis id="B91AEAA8BD04574394FDFBA9FF63BB50" box="[144,233,1086,1110]" italics="true" pageId="16" pageNumber="199">Ovaries</emphasis>
|
||
: Not observed.
|
||
<emphasis id="B91AEAA8BD04574395F7FBA9FDB5BB50" box="[410,575,1086,1110]" italics="true" pageId="16" pageNumber="199">Genital glands</emphasis>
|
||
: In segment 10 two simple, elongated, straight, narrow glands; in segment 21 similar in size to glands of segment 10, but twice-folded and bent posteriorly; in segment 22 only one gland, finger-shaped.
|
||
</paragraph>
|
||
<paragraph id="8BD136BABD04574394FDFB31FE1EBAD9" blockId="16.[144,1104,206,1671]" pageId="16" pageNumber="199">
|
||
Biological notes: The
|
||
<typeStatus id="54D58818BD04574395E4FB31FE33BBB9" box="[393,441,1190,1215]" pageId="16" pageNumber="199">type</typeStatus>
|
||
locality is located in the Drakensberg foothills at
|
||
<emphasis id="B91AEAA8BD0457439783FB31FB82BBB8" box="[1006,1032,1190,1214]" italics="true" pageId="16" pageNumber="199">ca</emphasis>
|
||
.
|
||
<quantity id="4C969B5FBD0457439075FB31FF22BBD9" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="199" unit="m" value="1500.0">1500 m</quantity>
|
||
, where there is a deep valley with steep hillsides covered by indigenous bushes and open grassveld (
|
||
<bibRefCitation id="EFFF4B4BBD0457439526FB71FDD8BBF9" author="POOLEY, T. & PLAYER, I." box="[331,594,1254,1279]" pageId="16" pageNumber="199" refId="ref12636" refString="POOLEY, T. & PLAYER, I. 1995. KwaZulu-Natal wildlife destinations. Halfway House: Southern Book Publishers (PTY) Ltd." type="book" year="1995">Pooley & Player 1995</bibRefCitation>
|
||
). During earlier years the area was used for grazing, but has never been ploughed or been arable land. The
|
||
<typeStatus id="54D58818BD045743972BFA91FC0BBA19" box="[838,897,1286,1311]" pageId="16" pageNumber="199">types</typeStatus>
|
||
were found on the plateau covered by grass and herbs, and were dug out at a depth of
|
||
<quantity id="4C969B5FBD0457439702FAB1FC45BA39" box="[879,975,1318,1343]" metricMagnitude="-1" metricUnit="m" metricValue="1.55" metricValueMax="3.0" metricValueMin="0.1" pageId="16" pageNumber="199" unit="cm" value="15.5" valueMax="30.0" valueMin="1.0">1–30 cm</quantity>
|
||
from moist soil and from between grass roots.A large part of the grassland had been burnt about six months previously. Earthworm presence between the roots of burnt grasses indicates an ability to migrate into deeper soil during unfavorable conditions, and a return to the top layer when conditions permit. The species is probably restricted to the outskirts of the Drakensberg range.
|
||
</paragraph>
|
||
<paragraph id="8BD136BABD04574294FDFA79FCC6BE61" blockId="16.[144,1104,206,1671]" lastBlockId="17.[181,1142,206,359]" lastPageId="17" lastPageNumber="200" pageId="16" pageNumber="199">
|
||
Discussion:
|
||
<taxonomicName id="4C6E4D39BD0457439577FA79FE01B900" authorityName="Plisko" authorityYear="2002" box="[282,395,1518,1542]" class="Clitellata" family="Microchaetidae" genus="Proandricus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="199" phylum="Annelida" rank="species" species="richerti">
|
||
<emphasis id="B91AEAA8BD0457439577FA79FE01B900" box="[282,395,1518,1542]" italics="true" pageId="16" pageNumber="199">P. richerti</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="4C6E4D39BD0457439656FA79FD23B900" baseAuthorityName="Beddard" baseAuthorityYear="1907" box="[571,681,1518,1543]" class="Clitellata" family="Microchaetidae" genus="Proandricus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="199" phylum="Annelida" rank="species" species="colletti">
|
||
<emphasis id="B91AEAA8BD0457439656FA79FDC6B900" box="[571,588,1518,1542]" italics="true" pageId="16" pageNumber="199">P</emphasis>
|
||
.
|
||
<emphasis id="B91AEAA8BD0457439636FA79FD23B900" box="[603,681,1518,1542]" italics="true" pageId="16" pageNumber="199">colletti</emphasis>
|
||
</taxonomicName>
|
||
species-group, having only septa 7/8 and 8/9 thickened, and spermathecal pores in intersegmental furrows 11/12 and 12/13. Similar to
|
||
<taxonomicName id="4C6E4D39BD045743956CF9B9FE04B940" authorityName="Plisko" authorityYear="2000" box="[257,398,1582,1606]" class="Clitellata" family="Microchaetidae" genus="Proandricus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="199" phylum="Annelida" rank="species" species="bergvillensis">
|
||
<emphasis id="B91AEAA8BD045743956CF9B9FE04B940" box="[257,398,1582,1606]" italics="true" pageId="16" pageNumber="199">bergvillensis</emphasis>
|
||
</taxonomicName>
|
||
in the backward extension of the seminal vesicles, and relatively short typhlosole. However, extension of the seminal vesicles in
|
||
<taxonomicName id="4C6E4D39BD045743973AF9D9FC21B960" authorityName="Plisko" authorityYear="2002" box="[855,939,1614,1638]" class="Clitellata" family="Microchaetidae" genus="Proandricus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="199" phylum="Annelida" rank="species" species="richerti">
|
||
<emphasis id="B91AEAA8BD045743973AF9D9FC21B960" box="[855,939,1614,1638]" italics="true" pageId="16" pageNumber="199">richerti</emphasis>
|
||
</taxonomicName>
|
||
probably does not cross septum 12/13.Also the shape of the tubercula pubertatis differs in both species. The length of the typhlosole in this new species is worthy of special note, as it is exceptionally short, occupying only a little more than one-third of the body segments. In the other species of the genus, the typhlosole extends to about two-thirds of the segment number, leaving only about one-third of segments without typhlosole. The presence of diverticula is also characteristic of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |