300 lines
36 KiB
XML
300 lines
36 KiB
XML
<document id="0F7A0FF5B34AD759D5A0D374B2F92DB1" ID-CLB-Dataset="22877" ID-DOI="10.5852/ejt.2020.629" ID-GBIF-Dataset="3722aa65-b935-4294-ac20-0d1179da9015" ID-Zenodo-Dep="3762280" ID-ZooBank="ECEBD539-6E3E-45BE-A0CB-264DF3270CC0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587578640316" checkinUser="valdenar" docAuthor="Herbert, David G." docDate="2020" docId="03C787854611FFDC424141723450F914" docLanguage="en" docName="ejt-629_herbert.pdf.imf" docOrigin="European Journal of Taxonomy 629" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Costulodonta bidens Herbert 2020, gen. et sp. nov." docType="treatment" docUuid="C5C0F93F-7276-41F1-9B91-AB8C0578CB65" docUuidSource="ZooBank" docVersion="7" lastPageNumber="31" masterDocId="FFFEFFFD460CFFC2400243213138FFD9" masterDocTitle="Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies" masterLastPageNumber="55" masterPageNumber="1" pageNumber="30" updateTime="1698835565728" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="04431934601231A6B6D5D48EDFAE0968" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6EA978F1F16F266F938AC50F34F890EC">
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<mods:title id="521BA269C700EC44F331AF62B4D3C4C3">Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies</mods:title>
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<mods:name id="93AA06DF1E4924C201AFAE7A9B616E08" type="personal">
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<mods:roleTerm id="26CBBA2FC80916A4767FAEBCCD1EE2FD">Author</mods:roleTerm>
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<mods:namePart id="45A78945FA050FB4E25C9F146F893963">Herbert, David G.</mods:namePart>
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<mods:nameIdentifier id="273F5A9A9DB8D64F6C633E0A355A18FD" type="LSID">urn:lsid:zoobank.org:author:0C09EE45-6198-482E-857A-EF690C2A016F</mods:nameIdentifier>
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<mods:affiliation id="AEE39D6CD00C639F130AF86739A68B33">Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff, CF 10 3 NP, UK. Formerly at KwaZulu-Natal Museum, P. Bag 9070, Pietermaritzburg 3201, South Africa</mods:affiliation>
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<mods:nameIdentifier id="4377D2F30EF5E08656B4A67BA1A1A01A" type="email">phasianella@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="851DFBADF29B5A3C984E2BC742E250EB">text</mods:typeOfResource>
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<mods:titleInfo id="41F96D9DAFBAB947C68041AF5BE4DD87">
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<mods:title id="9B8239B8CF0149C6BD706EB4AF25738B">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="F2B503D033A273BD4F412B965A8957F1">
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<mods:date id="1811E4A347DC181D07DECA68DE64F202">2020</mods:date>
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<mods:number id="9ED50C8122FE7A9841B6C693A3E10E9B">2020-04-17</mods:number>
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<mods:detail id="88ADA5EDB2D1BF3D3E8C103C1F361334" type="volume">
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<mods:number id="4307B9183CBEA6BCBC7F557E8C9747A6">629</mods:number>
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<mods:classification id="F808E7D16EBB3602D5EAE1133030AF0F">journal article</mods:classification>
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<mods:identifier id="1D0A7F9CD63763051FC9D08AF1B25C61" type="CLB-Dataset">22877</mods:identifier>
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<mods:identifier id="F6FA4826861FD893E41C0114D1794FC4" type="DOI">10.5852/ejt.2020.629</mods:identifier>
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<mods:identifier id="58D6E1537E2B75C464CE81B15D0E78CA" type="GBIF-Dataset">3722aa65-b935-4294-ac20-0d1179da9015</mods:identifier>
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<mods:identifier id="18E1F04A56976D93AA17A726F2FB1FC2" type="Zenodo-Dep">3762280</mods:identifier>
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<treatment id="03C787854611FFDC424141723450F914" ID-DOI="http://doi.org/10.5281/zenodo.3804707" ID-GBIF-Taxon="163732379" ID-Zenodo-Dep="3804707" LSID="urn:lsid:zoobank.org:act:C5C0F93F-7276-41F1-9B91-AB8C0578CB65" httpUri="http://treatment.plazi.org/id/03C787854611FFDC424141723450F914" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
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<subSubSection id="C37465184611FFDF4241417235FAFD4B" pageId="29" pageNumber="30" type="nomenclature">
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<paragraph id="8BD136934611FFDF4241417232C9FDB7" blockId="29.[370,1218,595,693]" box="[579,1009,595,622]" pageId="29" pageNumber="30">
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<heading id="D09981FF4611FFDF4241417232C9FDB7" box="[579,1009,595,622]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="30" reason="2">
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<taxonomicName id="4C6E4D104611FFDF424141723201FDB7" ID-CoL="7QTRW" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[579,825,595,622]" class="Gastropoda" family="Charopidae" genus="Costulodonta" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="bidens" status="gen. et sp. nov.">
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<emphasis id="B91AEA814611FFDF424141723201FDB7" bold="true" box="[579,825,595,622]" italics="true" pageId="29" pageNumber="30">Costulodonta bidens</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22957FA4611FFDF4343417532C9FDB7" box="[833,1009,596,622]" pageId="29" pageNumber="30" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8BD136934611FFDF4170415635FAFD4B" blockId="29.[370,1218,595,693]" box="[370,1218,631,658]" pageId="29" pageNumber="30">
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<uri id="FFFF3A914611FFDF4170415635FAFD4B" box="[370,1218,631,658]" pageId="29" pageNumber="30">
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urn:lsid:zoobank.org:act:
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<uuid id="FFC80C464611FFDF4295415635FAFD4B" box="[663,1218,631,658]" pageId="29" pageNumber="30">C5C0F93F-7276-41F1-9B91-AB8C0578CB65</uuid>
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</uri>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37465184611FFDF428F41BA329FFD6C" box="[653,935,667,693]" pageId="29" pageNumber="30" type="description">
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<paragraph id="8BD136934611FFDF428F41BA329FFD6C" blockId="29.[370,1218,595,693]" box="[653,935,667,693]" pageId="29" pageNumber="30">
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<figureCitation id="13552A164611FFDF428F41BA33DBFD6C" box="[653,739,667,693]" captionStart="Fig" captionStartId="28.[189,232,1894,1920]" captionTargetBox="[202,1395,669,1835]" captionTargetId="figure@28.[189,1399,656,1854]" captionTargetPageId="28" captionText="Fig. 10. Shells of Costulodonta gen. nov. species. A–D. C. acinaces (Connolly, 1933) gen. et comb. nov., Van Reenen, KwaZulu-Natal, diameter 1.39 mm (NMSA A9183). E–H. C. bidens gen. et sp. nov., holotype, diameter 1.65 mm (NMSA V5678/T4264). I–K. C. burnupi (Connolly, 1933) gen. et comb. nov., Mkolombe Mtn, Weenen, KwaZulu-Natal, diameter 1.32 mm (NMSA A9187)." figureDoi="http://doi.org/10.5281/zenodo.3762300" httpUri="https://zenodo.org/record/3762300/files/figure.png" pageId="29" pageNumber="30">Figs 10</figureCitation>
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E–H, 11, 18K–L
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</paragraph>
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</subSubSection>
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<subSubSection id="C37465184611FFDF40BF41FC3048FC6F" pageId="29" pageNumber="30" type="diagnosis">
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<paragraph id="8BD136934611FFDF40BF41FC300EFD2E" blockId="29.[189,310,733,759]" box="[189,310,733,759]" pageId="29" pageNumber="30">
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<heading id="D09981FF4611FFDF40BF41FC300EFD2E" bold="true" box="[189,310,733,759]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
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<emphasis id="B91AEA814611FFDF40BF41FC300EFD2E" bold="true" box="[189,310,733,759]" pageId="29" pageNumber="30">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD136934611FFDF40BF402D3048FC6F" blockId="29.[189,1399,780,950]" pageId="29" pageNumber="30">
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Shell small, spire flat or at most slightly raised; protoconch for the most part sculptured by close-set axial riblets; teleoconch sculpture of close-set, compound axial riblets and microscopic spiral threads; aperture lacking parietal and columellar dentition; palatal region with two relatively small denticles, one at mid-whorl, the other basal. Shell translucent, corneous-brown to straw-brown when fresh; diameter up to
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<quantity id="4C969B764611FFDF410240BA3053FC6F" box="[256,363,923,950]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="29" pageNumber="30" unit="mm" value="1.65">1.65 mm</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37465184611FFDF40BF40FF35F8FBFE" box="[189,1216,990,1063]" pageId="29" pageNumber="30" type="etymology">
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<paragraph id="8BD136934611FFDF40BF40FF307CFC21" blockId="29.[189,324,990,1016]" box="[189,324,990,1016]" pageId="29" pageNumber="30">
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<heading id="D09981FF4611FFDF40BF40FF307CFC21" bold="true" box="[189,324,990,1016]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
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<emphasis id="B91AEA814611FFDF40BF40FF307CFC21" bold="true" box="[189,324,990,1016]" pageId="29" pageNumber="30">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD136934611FFDF40BF472C35F8FBFE" blockId="29.[189,1216,1037,1063]" box="[189,1216,1037,1063]" pageId="29" pageNumber="30">
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From the Latin
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<emphasis id="B91AEA814611FFDF4176472C30FEFBFE" box="[372,454,1037,1063]" italics="true" pageId="29" pageNumber="30">bis, bi-</emphasis>
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||
: two, and
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<emphasis id="B91AEA814611FFDF4240472C3340FBFE" box="[578,632,1037,1063]" italics="true" pageId="29" pageNumber="30">dens</emphasis>
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: a tooth; with reference to the apertural dentition.
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</paragraph>
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</subSubSection>
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<caption id="DF11661B4611FFDF40BF4449328FF834" ID-DOI="http://doi.org/10.5281/zenodo.3762302" ID-Zenodo-Dep="3762302" httpUri="https://zenodo.org/record/3762302/files/figure.png" pageId="29" pageNumber="30" startId="29.[189,232,1896,1922]" targetBox="[302,1285,1093,1856]" targetPageId="29">
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<paragraph id="8BD136934611FFDF40BF4449328FF834" blockId="29.[189,1399,1896,2029]" pageId="29" pageNumber="30">
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||
<emphasis id="B91AEA814611FFDF40BF4449302FF85B" bold="true" box="[189,279,1896,1922]" pageId="29" pageNumber="30">Fig. 11.</emphasis>
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Distribution of
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<taxonomicName id="4C6E4D104611FFDF41D244483356F85A" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[464,622,1897,1923]" class="Gastropoda" family="Charopidae" genus="Costulodonta" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus" status="gen. nov.">
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||
<emphasis id="B91AEA814611FFDF41D244483356F85A" box="[464,622,1897,1923]" italics="true" pageId="29" pageNumber="30">Costulodonta</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A22957FA4611FFDF4276444833E6F85A" box="[628,734,1897,1923]" pageId="29" pageNumber="30" rank="genus">gen. nov.</taxonomicNameLabel>
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species.
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<taxonomicName id="4C6E4D104611FFDF4345444835A4F85A" authority="(Connolly, 1933)" authorityName="Herbert" authorityYear="2020" baseAuthorityName="Connolly" baseAuthorityYear="1933" box="[839,1180,1896,1923]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="acinaces" status="gen. et comb. nov.">
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<emphasis id="B91AEA814611FFDF4345444832F7F85B" box="[839,975,1897,1923]" italics="true" pageId="29" pageNumber="30">C. acinaces</emphasis>
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(
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<bibRefCitation id="EFFF4B624611FFDF43DF444935ACF85A" author="Connolly M." box="[989,1172,1896,1923]" pageId="29" pageNumber="30" pagination="145 - 150" refId="ref26638" refString="Connolly M. 1933. On South African Endodontidae. Part 2. Annals of the Natal Museum 7 (2): 145 - 150." type="journal article" year="1933">Connolly, 1933</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A22957FA4611FFDF44A14448344FF85A" box="[1187,1399,1896,1923]" pageId="29" pageNumber="30" rank="species">gen. et comb. nov.</taxonomicNameLabel>
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(green square),
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<taxonomicName id="4C6E4D104611FFDF417144AD30D9F87F" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[371,481,1932,1958]" class="Gastropoda" family="Charopidae" genus="Costulodonta" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="bidens" status="gen. et sp. nov.">
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<emphasis id="B91AEA814611FFDF417144AD30D9F87F" box="[371,481,1932,1958]" italics="true" pageId="29" pageNumber="30">C. bidens</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22957FA4611FFDF41E544AD33ADF87F" box="[487,661,1932,1958]" pageId="29" pageNumber="30" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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(turquoise squares),
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<taxonomicName id="4C6E4D104611FFDF438A44AD35EBF87F" authority="(Connolly, 1933)" authorityName="Herbert" authorityYear="2020" baseAuthorityName="Connolly" baseAuthorityYear="1933" box="[904,1235,1932,1958]" class="Gastropoda" family="Charopidae" genus="Costulodonta" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="burnupi" status="gen. et comb. nov.">
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<emphasis id="B91AEA814611FFDF438A44AD353EF87F" box="[904,1030,1932,1958]" italics="true" pageId="29" pageNumber="30">C. burnupi</emphasis>
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(
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<bibRefCitation id="EFFF4B624611FFDF441644AD35F3F87F" author="Connolly M." box="[1044,1227,1932,1958]" pageId="29" pageNumber="30" pagination="145 - 150" refId="ref26638" refString="Connolly M. 1933. On South African Endodontidae. Part 2. Annals of the Natal Museum 7 (2): 145 - 150." type="journal article" year="1933">Connolly, 1933</bibRefCitation>
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||
)
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</taxonomicName>
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<taxonomicNameLabel id="A22957FA4611FFDF44D844AD31D7F813" pageId="29" pageNumber="30" rank="species">gen. et comb. nov.</taxonomicNameLabel>
|
||
(orange square)
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<taxonomicName id="4C6E4D104611FFDF41B7448E337EF810" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[437,582,1967,1993]" class="Gastropoda" family="Charopidae" genus="Costulodonta" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="pluridens" status="gen. et sp. nov.">
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||
<emphasis id="B91AEA814611FFDF41B7448E337EF810" box="[437,582,1967,1993]" italics="true" pageId="29" pageNumber="30">C. pluridens</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22957FA4611FFDF424F449133C6F813" box="[589,766,1968,1994]" pageId="29" pageNumber="30" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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(pink squares),
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<taxonomicName id="4C6E4D104611FFDF43B9448E3039F835" authority="(Melvill & Ponsonby, 1908)" authorityName="Herbert" authorityYear="2020" baseAuthorityName="Melvill & Ponsonby" baseAuthorityYear="1908" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="trilamellaris" status="gen. et comb. nov.">
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<emphasis id="B91AEA814611FFDF43B9448E3548F810" box="[955,1136,1967,1993]" italics="true" pageId="29" pageNumber="30">C. trilamellaris</emphasis>
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||
(
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<bibRefCitation id="EFFF4B624611FFDF447D448E31C1F835" author="Melvill J. C. & Ponsonby J. H." pageId="29" pageNumber="30" pagination="129 - 136" refId="ref26977" refString="Melvill J. C. & Ponsonby J. H. 1908. Descriptions of nine terrestrial Mollusca from South Africa. Annals and Magazine of Natural History, Series 8, 1 (2): 129 - 136. https: // doi. org / 10.1080 / 00222930808692373" type="journal article" year="1908">Melvill & Ponsonby, 1908</bibRefCitation>
|
||
)
|
||
</taxonomicName>
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||
<taxonomicNameLabel id="A22957FA4611FFDF410B44F230D8F834" box="[265,480,2002,2029]" pageId="29" pageNumber="30" rank="species">gen. et comb. nov.</taxonomicNameLabel>
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(royal blue square). Contour at 1000 m.
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</paragraph>
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</caption>
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<subSubSection id="C37465184612FFDC40BF422A3023FD3C" pageId="30" pageNumber="31" type="materials_examined">
|
||
<paragraph id="8BD136934612FFDC40BF422A3092FEFC" blockId="30.[189,426,267,293]" box="[189,426,267,293]" pageId="30" pageNumber="31">
|
||
<heading id="D09981FF4612FFDC40BF422A3092FEFC" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
|
||
<emphasis id="B91AEA814612FFDC40BF422A3092FEFC" bold="true" box="[189,426,267,293]" pageId="30" pageNumber="31">Material examined</emphasis>
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||
</heading>
|
||
</paragraph>
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<paragraph id="8BD136934612FFDC40E8421D3064FE8F" blockId="30.[189,1399,316,456]" box="[234,348,316,342]" pageId="30" pageNumber="31">
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||
<emphasis id="B91AEA814612FFDC40E8421D3064FE8F" bold="true" box="[234,348,316,342]" pageId="30" pageNumber="31">
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||
<typeStatus id="54D588314612FFDC40E8421D3064FE8F" box="[234,348,316,342]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
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</emphasis>
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||
</paragraph>
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<paragraph id="8BD136934612FFDC40BF424335CAFE11" blockId="30.[189,1399,316,456]" pageId="30" pageNumber="31">
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||
<materialsCitation id="3B063CCE4612FFDC40BF424335CAFE11" ID-GBIF-Occurrence="2605305695" collectedFrom="montane Podocarpus forest, in leaf-litter" collectingDate="1997-07-15" collectionCode="NMSA" country="South Africa" county="Yellow-Wood Forest" latitude="-29.3" location="Giant's Castle Game Res." longLatPrecision="73" longitude="29.467" municipality="Natal Parks Board" pageId="30" pageNumber="31" specimenCode="V5678/T4264" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="holotype">
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<collectingCountry id="F37976034612FFDC40BF424330AEFEA5" box="[189,406,354,380]" name="South Africa" pageId="30" pageNumber="31">SOUTH AFRICA</collectingCountry>
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||
•
|
||
<collectingRegion id="49AAF8714612FFDC41BF42403340FEA2" box="[445,632,353,379]" country="South Africa" name="KwaZulu-Natal" pageId="30" pageNumber="31">KwaZulu-Natal</collectingRegion>
|
||
,
|
||
<location id="8EB160484612FFDC4289424032E9FEA5" LSID="urn:lsid:plazi:treatment:03C787854611FFDC424141723450F914:8EB160484612FFDC4289424032E9FEA5" box="[651,977,353,380]" country="South Africa" county="Yellow-Wood Forest" latitude="-29.3" longLatPrecision="73" longitude="29.467" municipality="Natal Parks Board" name="Giant's Castle Game Res." pageId="30" pageNumber="31" stateProvince="KwaZulu-Natal">Giant’s Castle Game Res.</location>
|
||
,
|
||
<location id="8EB160484612FFDC43E1424035D9FEA5" LSID="urn:lsid:plazi:treatment:03C787854611FFDC424141723450F914:8EB160484612FFDC43E1424035D9FEA5" box="[995,1249,353,380]" country="South Africa" county="Yellow-Wood Forest" latitude="-29.3" longLatPrecision="73" longitude="29.467" municipality="Natal Parks Board" name="Yellow-Wood Forest" pageId="30" pageNumber="31" stateProvince="KwaZulu-Natal">Yellow-Wood Forest</location>
|
||
;
|
||
<geoCoordinate id="EE5A50544612FFDC44F642433449FEA5" box="[1268,1393,354,380]" degrees="29.300" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="55" value="-29.3">29.300° S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5A50544612FFDC40BF42A63001FE7B" box="[189,313,391,418]" degrees="29.467" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="55" value="29.467">29.467° E</geoCoordinate>
|
||
;
|
||
<date id="FFD010534612FFDC414A42A630D9FE78" box="[328,481,391,418]" pageId="30" pageNumber="31" value="1997-07-15">
|
||
<collectingDate id="EF94E9BB4612FFDC414A42A630D9FE78" box="[328,481,391,418]" pageId="30" pageNumber="31" value="1997-07-15">15 Jul. 1997</collectingDate>
|
||
</date>
|
||
;
|
||
<collectingMunicipality id="6BB5ACE94612FFDC41F242A633F6FE78" box="[496,718,391,417]" pageId="30" pageNumber="31">Natal Parks Board</collectingMunicipality>
|
||
(now
|
||
<location id="8EB160484612FFDC431C42A632ABFE78" LSID="urn:lsid:plazi:treatment:03C787854611FFDC424141723450F914:8EB160484612FFDC431C42A632ABFE78" box="[798,915,391,417]" country="South Africa" county="Yellow-Wood Forest" latitude="-29.3" longLatPrecision="73" longitude="29.467" municipality="Natal Parks Board" name="Ezemvelo" pageId="30" pageNumber="31" stateProvince="KwaZulu-Natal">Ezemvelo</location>
|
||
<collectingRegion id="49AAF8714612FFDC439C42A6356EFE78" box="[926,1110,391,417]" country="South Africa" name="KwaZulu-Natal" pageId="30" pageNumber="31">KwaZulu-Natal</collectingRegion>
|
||
<location id="8EB160484612FFDC445D42A635FAFE7B" LSID="urn:lsid:plazi:treatment:03C787854611FFDC424141723450F914:8EB160484612FFDC445D42A635FAFE7B" box="[1119,1218,391,418]" country="South Africa" county="Yellow-Wood Forest" latitude="-29.3" longLatPrecision="73" longitude="29.467" municipality="Natal Parks Board" name="Wildlife" pageId="30" pageNumber="31" stateProvince="KwaZulu-Natal">Wildlife</location>
|
||
) leg.;
|
||
<collectedFrom id="3872756E4612FFDC451342A9330BFE1E" pageId="30" pageNumber="31">
|
||
montane
|
||
<taxonomicName id="4C6E4D104612FFDC40BF428C3073FE1E" box="[189,331,429,455]" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" order="Pinales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B91AEA814612FFDC40BF428C3073FE1E" box="[189,331,429,455]" italics="true" pageId="30" pageNumber="31">Podocarpus</emphasis>
|
||
</taxonomicName>
|
||
forest, in leaf-litter
|
||
</collectedFrom>
|
||
; diameter
|
||
<quantity id="4C969B764612FFDC42AE428C3220FE1E" box="[684,792,429,456]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="30" pageNumber="31" unit="mm" value="1.65">1.65 mm</quantity>
|
||
, height
|
||
<quantity id="4C969B764612FFDC4376428C32D8FE1E" box="[884,992,429,455]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="30" pageNumber="31" unit="mm" value="0.83">0.83 mm</quantity>
|
||
;
|
||
<collectionCode id="ED7FAE564612FFDC43EE428C3507FE1E" box="[1004,1087,429,455]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="30" pageNumber="31" type="Museum">NMSA</collectionCode>
|
||
<specimenCode id="DBC89EE84612FFDC4444428C35D6FE11" box="[1094,1262,429,456]" pageId="30" pageNumber="31">V5678/T4264</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD136934612FFDC40E842D0305FFDD2" blockId="30.[189,1294,497,561]" box="[234,359,497,523]" pageId="30" pageNumber="31">
|
||
<emphasis id="B91AEA814612FFDC40E842D0305FFDD2" bold="true" box="[234,359,497,523]" pageId="30" pageNumber="31">
|
||
<typeStatus id="54D588314612FFDC40E842D0305FFDD2" box="[234,359,497,523]" pageId="30" pageNumber="31" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BD136934612FFDC40BF41363436FDE8" blockId="30.[189,1294,497,561]" box="[189,1294,534,561]" pageId="30" pageNumber="31">
|
||
<materialsCitation id="3B063CCE4612FFDC40BF41363432FDE8" ID-GBIF-Occurrence="2605305631" box="[189,1290,534,561]" collectingDate="1997-07-15" collectionCode="NMSA" country="South Africa" county="Yellow-Wood Forest" latitude="-29.3" location="SOUTH AFRICA" longLatPrecision="73" longitude="29.467" municipality="Natal Parks Board" pageId="30" pageNumber="31" specimenCode="P1018/T4265" specimenCount="23" stateProvince="KwaZulu-Natal" typeStatus="paratype">
|
||
<collectingCountry id="F37976034612FFDC40BF413630B6FDE8" box="[189,398,535,561]" name="South Africa" pageId="30" pageNumber="31">SOUTH AFRICA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D68FD1A4612FFDC41A541363372FDE9" box="[423,586,534,561]" pageId="30" pageNumber="31" type="generic">23 specimens</specimenCount>
|
||
; same collection data as for holotype;
|
||
<collectionCode id="ED7FAE564612FFDC440E41363567FDE8" box="[1036,1119,535,561]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="30" pageNumber="31" type="Museum">NMSA</collectionCode>
|
||
<specimenCode id="DBC89EE84612FFDC446441373432FDE8" box="[1126,1290,534,561]" pageId="30" pageNumber="31">P1018/T4265</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD136934612FFDC40E8417B309EFDAD" blockId="30.[189,1399,602,741]" box="[234,422,602,628]" pageId="30" pageNumber="31">
|
||
<emphasis id="B91AEA814612FFDC40E8417B309EFDAD" bold="true" box="[234,422,602,628]" pageId="30" pageNumber="31">Other material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BD136934612FFDC40BF41A13023FD3C" blockId="30.[189,1399,602,741]" pageId="30" pageNumber="31">
|
||
<materialsCitation id="3B063CCE4612FFDC40BF41A1302FFD3C" ID-GBIF-Occurrence="2605305716" collectedFrom="montane forest, in leaf-litter" collectingDate="2004-09-24" collectionCode="NMSA" collectorName="A. Moussalli & D. Stuart-Fox" country="South Africa" latitude="-29.2856" location="environs of Main Cave" longLatPrecision="6" longitude="29.5179" municipality="Giant's Castle" pageId="30" pageNumber="31" specimenCode="W3106" specimenCount="2" stateProvince="KwaZulu-Natal">
|
||
<collectingCountry id="F37976034612FFDC40BF41A130A8FD43" box="[189,400,640,666]" name="South Africa" pageId="30" pageNumber="31">SOUTH AFRICA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D68FD1A4612FFDC41A841A13307FD43" box="[426,575,640,666]" pageId="30" pageNumber="31" type="generic">2 specimens</specimenCount>
|
||
;
|
||
<collectingRegion id="49AAF8714612FFDC424E41A1323FFD43" box="[588,775,640,666]" country="South Africa" name="KwaZulu-Natal" pageId="30" pageNumber="31">KwaZulu-Natal</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BB5ACE94612FFDC431641A13285FD43" box="[788,957,640,666]" pageId="30" pageNumber="31">Giant’s Castle</collectingMunicipality>
|
||
,
|
||
<location id="8EB160484612FFDC43C841A135E7FD43" LSID="urn:lsid:plazi:treatment:03C787854611FFDC424141723450F914:8EB160484612FFDC43C841A135E7FD43" box="[970,1247,640,666]" country="South Africa" latitude="-29.2856" longLatPrecision="6" longitude="29.5179" municipality="Giant's Castle" name="environs of Main Cave" pageId="30" pageNumber="31" stateProvince="KwaZulu-Natal">environs of Main Cave</location>
|
||
;
|
||
<geoCoordinate id="EE5A50544612FFDC44E941A13449FD43" box="[1259,1393,640,667]" degrees="29.2856" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="5" value="-29.2856">29.2856° S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5A50544612FFDC40BF4184307EFD19" box="[189,326,677,704]" degrees="29.5179" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="5" value="29.5179">29.5179° E</geoCoordinate>
|
||
;
|
||
<date id="FFD010534612FFDC4157418730CEFD19" box="[341,502,678,704]" pageId="30" pageNumber="31" value="2004-09-24">
|
||
<collectingDate id="EF94E9BB4612FFDC4157418730CEFD19" box="[341,502,678,704]" pageId="30" pageNumber="31" value="2004-09-24">24 Sep. 2004</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="269B53454612FFDC4201418733A5FD19" box="[515,669,678,704]" pageId="30" pageNumber="31">A. Moussalli</collectorName>
|
||
and
|
||
<collectorName id="269B53454612FFDC42DB41873244FD19" box="[729,892,678,704]" pageId="30" pageNumber="31">D. Stuart-Fox</collectorName>
|
||
leg.;
|
||
<collectedFrom id="3872756E4612FFDC43C14187342EFD19" box="[963,1302,678,704]" pageId="30" pageNumber="31">montane forest, in leaf-litter</collectedFrom>
|
||
;
|
||
<collectionCode id="ED7FAE564612FFDC45214187344EFD19" box="[1315,1398,678,704]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="30" pageNumber="31" type="Museum">NMSA</collectionCode>
|
||
<specimenCode id="DBC89EE84612FFDC40BF41EA302FFD3C" box="[189,279,715,741]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="30" pageNumber="31" type="Museum">W3106</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37465184612FFDC40BF402E32D7FB74" pageId="30" pageNumber="31" type="description">
|
||
<paragraph id="8BD136934612FFDC40BF402E3076FCF0" blockId="30.[189,334,783,809]" box="[189,334,783,809]" pageId="30" pageNumber="31">
|
||
<heading id="D09981FF4612FFDC40BF402E3076FCF0" bold="true" box="[189,334,783,809]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
|
||
<emphasis id="B91AEA814612FFDC40BF402E3076FCF0" bold="true" box="[189,334,783,809]" pageId="30" pageNumber="31">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD136934612FFDC40BF406132D7FB74" blockId="30.[189,1399,832,1198]" pageId="30" pageNumber="31">
|
||
Shell small, diameter up to
|
||
<quantity id="4C969B764612FFDC41FE4061335FFC82" box="[508,615,832,859]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="30" pageNumber="31" unit="mm" value="1.65">1.65 mm</quantity>
|
||
, H/D ratio ±0.50; spire flat or at most slightly raised; whorls tightly coiled; last adult whorl slightly descendant; suture indented, periphery evenly convex. Protoconch comprising apical cap plus approx. 0.75 whorl; diameter ±330 μm; initially smooth, but for the most part sculptured by close-set axial riblets, with indistinct traces of irregular spiral threads. Teleoconch of up to 3.5 whorls; sculptured by distinct, close-set, compound axial riblets with 3–4 finer intermediary axial threads; intervals between riblets 1–2 times riblet width at whorl periphery; spiral sculpture of microscopic threads, strongest below suture. Umbilicus of moderate width. Aperture lunate, somewhat broader basally; parietal and columellar dentition lacking; palatal region with two relatively small denticles, one at mid-whorl, the other basal, set back approx. ⅛ whorl behind outer lip (sometimes weak). Shell translucent, corneous-brown to straw-brown when fresh.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37465184612FFDC40BF47FE32E8FA42" pageId="30" pageNumber="31" type="distribution">
|
||
<paragraph id="8BD136934612FFDC40BF47FE330CFB20" blockId="30.[189,564,1247,1273]" box="[189,564,1247,1273]" pageId="30" pageNumber="31">
|
||
<heading id="D09981FF4612FFDC40BF47FE330CFB20" bold="true" box="[189,564,1247,1273]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
|
||
<emphasis id="B91AEA814612FFDC40BF47FE330CFB20" bold="true" box="[189,564,1247,1273]" pageId="30" pageNumber="31">Distribution and conservation</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD136934612FFDC40BF463132E8FA42" blockId="30.[189,1399,1296,1435]" pageId="30" pageNumber="31">
|
||
A narrow-range endemic (
|
||
<figureCitation id="13552A164612FFDC41FB46313368FAF3" box="[505,592,1296,1322]" captionStart="Fig" captionStartId="29.[189,232,1896,1922]" captionTargetBox="[302,1285,1093,1856]" captionTargetId="figure@29.[302,1285,1093,1856]" captionTargetPageId="29" captionText="Fig. 11. Distribution of Costulodonta gen. nov. species. C. acinaces (Connolly, 1933) gen. et comb. nov. (green square), C. bidens gen. et sp. nov. (turquoise squares), C. burnupi (Connolly, 1933) gen. et comb. nov. (orange square) C. pluridens gen. et sp. nov. (pink squares), C. trilamellaris (Melvill & Ponsonby, 1908) gen. et comb. nov. (royal blue square). Contour at 1000 m." figureDoi="http://doi.org/10.5281/zenodo.3762302" httpUri="https://zenodo.org/record/3762302/files/figure.png" pageId="30" pageNumber="31">Fig. 11</figureCitation>
|
||
), known only from the Drakensberg foothills (‘Little Berg’) in the Giant’s Castle area,
|
||
<collectingRegion id="49AAF8714612FFDC41A346143364FA96" box="[417,604,1333,1359]" country="South Africa" name="KwaZulu-Natal" pageId="30" pageNumber="31">KwaZulu-Natal</collectingRegion>
|
||
, at approx.
|
||
<quantity id="4C969B764612FFDC42D946173243FA89" box="[731,891,1334,1360]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="30" pageNumber="31" unit="m" value="1750.0" valueMax="1800.0" valueMin="1700.0">1700–1800 m</quantity>
|
||
a.s.l.; in leaf-litter of northern afrotemperate forest. The area falls within the Giant’s Castle Game Reserve, which is part of the Maloti-Drakensberg World Heritage Site. It is thus afforded a high degree of protection.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37465184612FFDC40BF46E43450F914" pageId="30" pageNumber="31" type="discussion">
|
||
<paragraph id="8BD136934612FFDC40BF46E43017FA06" blockId="30.[189,303,1477,1503]" box="[189,303,1477,1503]" pageId="30" pageNumber="31">
|
||
<heading id="D09981FF4612FFDC40BF46E43017FA06" bold="true" box="[189,303,1477,1503]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
|
||
<emphasis id="B91AEA814612FFDC40BF46E43017FA06" bold="true" box="[189,303,1477,1503]" pageId="30" pageNumber="31">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD136934612FFDC40BF46D73450F914" blockId="30.[189,1399,1526,1741]" pageId="30" pageNumber="31">
|
||
Amongst its congeners,
|
||
<taxonomicName id="4C6E4D104612FFDC41DD46D733ECF9C9" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[479,724,1526,1552]" class="Gastropoda" family="Charopidae" genus="Costulodonta" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="bidens" status="gen. et sp. nov.">
|
||
<emphasis id="B91AEA814612FFDC41DD46D733ECF9C9" box="[479,724,1526,1552]" italics="true" pageId="30" pageNumber="31">Costulodonta bidens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22957FA4612FFDC42DF46D732ABF9C9" box="[733,915,1526,1552]" pageId="30" pageNumber="31" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
is rendered distinctive on account of its relatively simple apertural dentition. Superficially the shell shows considerable resemblance to that of ‘
|
||
<taxonomicName id="4C6E4D104612FFDC40C545603062F982" authorityName="Pilsbry" authorityYear="1893" box="[199,346,1601,1627]" class="Bryopsida" family="Mniaceae" genus="Trachycystis" kingdom="Plantae" order="Bryales" pageId="30" pageNumber="31" phylum="Bryophyta" rank="genus">
|
||
<emphasis id="B91AEA814612FFDC40C545603062F982" box="[199,346,1601,1627]" italics="true" pageId="30" pageNumber="31">Trachycystis</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<emphasis id="B91AEA814612FFDC416A456030CCF982" box="[360,500,1601,1627]" italics="true" pageId="30" pageNumber="31">contabulata</emphasis>
|
||
Connolly, 1932, but that species is larger (diameter up to
|
||
<quantity id="4C969B764612FFDC449F456035C1F985" box="[1181,1273,1601,1628]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="30" pageNumber="31" unit="mm" value="2.4">2.4 mm</quantity>
|
||
) and lacks apertural dentition. However, it does have a similarly sculptured protoconch and the two species may in fact be related. Though known to occur in the forests of the broader Giant’s Castle area (
|
||
<bibRefCitation id="EFFF4B624612FFDC44F945AD3067F915" author="Herbert D. & Kilburn D." pageId="30" pageNumber="31" refId="ref26755" refString="Herbert D. & Kilburn D. 2004. Field Guide to the Land Snails and Slugs of Eastern South Africa. Natal Museum, Pietermaritzburg." type="book" year="2004">Herbert & Kilburn 2004</bibRefCitation>
|
||
), ‘
|
||
<emphasis id="B91AEA814612FFDC417E459230B4F915" box="[380,396,1715,1740]" italics="true" pageId="30" pageNumber="31">T</emphasis>
|
||
’
|
||
<emphasis id="B91AEA814612FFDC419E45933310F915" box="[412,552,1714,1740]" italics="true" pageId="30" pageNumber="31">contabulata</emphasis>
|
||
has, to date, not been found to co-occur with
|
||
<taxonomicName id="4C6E4D104612FFDC444345923589F915" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" box="[1089,1201,1714,1741]" class="Gastropoda" family="Charopidae" genus="Costulodonta" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="bidens" status="gen. et sp. nov.">
|
||
<emphasis id="B91AEA814612FFDC444345923589F915" box="[1089,1201,1714,1741]" italics="true" pageId="30" pageNumber="31">C. bidens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22957FA4612FFDC44BA45923450F914" box="[1208,1384,1715,1741]" pageId="30" pageNumber="31" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |