treatments-xml/data/03/B2/87/03B287D3FFC0771DFE564D65D21C6BF1.xml
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<document id="19F11C86CABD2F2AD52B143F8DADF1CD" ID-CLB-Dataset="3296" ID-DOI="10.1080/00222933.2021.1984599" ID-GBIF-Dataset="b3381a22-4859-40e6-ba5e-b966903d7fe5" ID-ISSN="1464-5262" ID-Zenodo-Dep="5727887" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637769876395" checkinUser="plazi" docAuthor="Marçal, Ingrid Costa, Páez, Fernanda Polli, Silva, Priscila Frazato Da, Souza-Shibatta, Lenice &amp; Teixeira, Gustavo Monteiro" docDate="2021" docId="03B287D3FFC0771DFE564D65D21C6BF1" docLanguage="en" docName="JNATHIST.55.33-34.2145-2171.pdf" docOrigin="Journal of Natural History 55 (33 - 34)" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Aegla Leach 1820" docType="treatment" docVersion="5" lastPageNumber="2163" masterDocId="FF8BFFABFFD0770FFFC44B5FD65A6904" masterDocTitle="Cryptic diversity among populations of Aegla Leach, 1820 (Decapoda: Anomura: Aeglidae) from Tibagi River basin, Paraná state, Brazil, with descriptions of three new species" masterLastPageNumber="2171" masterPageNumber="2145" pageNumber="2161" updateTime="1699470245045" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="38692893F416685B5AC47634F2B81F78">Cryptic diversity among populations of Aegla Leach, 1820 (Decapoda: Anomura: Aeglidae) from Tibagi River basin, Paraná state, Brazil, with descriptions of three new species</mods:title>
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<mods:namePart id="712A5CFDB091008F6934D52098D70972">Marçal, Ingrid Costa</mods:namePart>
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<mods:namePart id="192D09C958C5292DB6E81C2C62DA682F">Páez, Fernanda Polli</mods:namePart>
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<mods:namePart id="8F3C6347D0712A660909DB723C918333">Silva, Priscila Frazato Da</mods:namePart>
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<mods:namePart id="C26D936AC5CB051C6024D4F8BD42C797">Souza-Shibatta, Lenice</mods:namePart>
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<mods:namePart id="21AA802901D551CE1DC98EA6EB176B9C">Teixeira, Gustavo Monteiro</mods:namePart>
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<mods:title id="742FC6D149AADC7EA32D69469F949F2D">Journal of Natural History</mods:title>
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<mods:part id="CCAD02B080CD0031446F366A47BA5B6F">
<mods:date id="B97D9A08CE82F3BFAB75EE417DEEA2D9">2021</mods:date>
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<mods:number id="1B0D6C6550D57DA1E8CEBC7A0D07C96B">2021-11-15</mods:number>
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<mods:number id="B9A2A57ED4806BDC3647B8FF53F58DC3">55</mods:number>
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<mods:number id="3BADC2F43CA0B0028F1377B4E4FA533B">33 - 34</mods:number>
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<mods:identifier id="9DFEE0BCEB29BA9B67D57A08F13CC45E" type="DOI">10.1080/00222933.2021.1984599</mods:identifier>
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<treatment id="03B287D3FFC0771DFE564D65D21C6BF1" ID-DOI="http://doi.org/10.5281/zenodo.5727897" ID-GBIF-Taxon="190792179" ID-Zenodo-Dep="5727897" LSID="urn:lsid:plazi:treatment:03B287D3FFC0771DFE564D65D21C6BF1" httpUri="http://treatment.plazi.org/id/03B287D3FFC0771DFE564D65D21C6BF1" lastPageId="18" lastPageNumber="2163" pageId="16" pageNumber="2161">
<subSubSection id="C301654EFFC0771FFE564D65D5CB6F50" box="[402,913,1594,1620]" pageId="16" pageNumber="2161" type="nomenclature">
<paragraph id="8BA436C5FFC0771FFE564D65D5CB6F50" blockId="16.[402,913,1594,1620]" box="[402,913,1594,1620]" pageId="16" pageNumber="2161">
<heading id="D0EC81A9FFC0771FFE564D65D5CB6F50" box="[402,913,1594,1620]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="2161" reason="5">
<taxonomicName id="4C1B4D46FFC0771FFE564D65D78F6F50" ID-CoL="485fdce0-0fdd-4805-8ee5-575b3a989889" authorityName="Leach" authorityYear="1820" box="[402,469,1594,1620]" class="Malacostraca" family="Aeglidae" genus="Aegla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="2171" phylum="Arthropoda" rank="genus">Aegla</taxonomicName>
<emphasis id="B96FEAD7FFC0771FFE184D65D7BC6F50" box="[476,486,1594,1620]" italics="true" pageId="16" pageNumber="2161">s</emphasis>
antosi
<emphasis id="B96FEAD7FFC0771FFDF24D65D5CB6F50" bold="true" box="[566,913,1594,1620]" italics="true" pageId="16" pageNumber="2161">sp. nov. Marçal and Teixeira</emphasis>
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<subSubSection id="C301654EFFC0771FFDF74D39D5EC6FB0" pageId="16" pageNumber="2161" type="description">
<paragraph id="8BA436C5FFC0771FFDF74D39D4AA6F84" blockId="16.[563,752,1638,1664]" box="[563,752,1638,1664]" pageId="16" pageNumber="2161">
<emphasis id="B96FEAD7FFC0771FFDF74D39D4AA6F84" bold="true" box="[563,752,1638,1664]" italics="true" pageId="16" pageNumber="2161">
(
<figureCitation id="13202A40FFC0771FFDF84D39D4BC6F84" box="[572,742,1638,1664]" captionStart-0="Figure 11" captionStart-1="Figure 12" captionStart-2="Figure 13" captionStart-3="Figure 14" captionStartId-0="17.[159,224,1557,1580]" captionStartId-1="19.[160,225,546,569]" captionStartId-2="20.[160,225,923,946]" captionStartId-3="21.[160,225,1616,1639]" captionTargetBox-0="[263,1053,964,1521]" captionTargetBox-1="[169,1144,147,507]" captionTargetBox-2="[167,1149,148,883]" captionTargetBox-3="[169,1144,1217,1578]" captionTargetId-0="figure-252@17.[261,1055,959,1526]" captionTargetId-1="figure-322@19.[162,1152,142,514]" captionTargetId-2="figure-162@20.[163,1153,142,891]" captionTargetId-3="figure-274@21.[162,1152,1212,1584]" captionTargetPageId-0="17" captionTargetPageId-1="19" captionTargetPageId-2="20" captionTargetPageId-3="21" captionText-0="Figure 11. Aegla santosi sp. nov., male holotype, CLE 19.3 mm, Codorna Branca stream, Telêmaco Borba, Paraná, Brazil (MZUEL 511). (a) dorsal view of the cephalothorax and anterior portion of the abdomen. (b) ventral view of ischium of minor cheliped (left) showing four tubercles (arrow). (c) third thoracic sternite truncate (arrow). (d) dorsal view of second abdominal epimeron (right side) showing anterolateral angle with corneous scale (white arrow) and anterior margin slightly concave (black arrow). Scale bars: a = 5.0 mm; bd = 1.0 mm." captionText-1="Figure 12. Aegla santosi sp. nov., male holotype, CLE 19.3 mm (MZUEL 511). (a) external surface view of the anterior portion of the cephalothorax showing pronounced protogastric lobes (white arrow) and well-developed subrostral process (black arrow). (b) dorsal view of the anterior region of the cephalothorax showing anterolateral spine reaching basal margin of the cornea (black arrow) and slightly elongated epibranchial area (white arrow). Scale bars: a = 1.0 mm; b = 5.0 mm." captionText-2="Figure 13. Aegla santosi sp. nov., male holotype, CLE 19.3 mm (MZUEL 511). (a) dorsal view of the dactylus, propodus and carpus of major cheliped (right) showing disciform palmar crest (thick arrow) and high carpal ridge (thin arrow). (b) ventral view of ischium of major cheliped (right) with four tubercles (arrows). (c) dorsal view of the dactylus, propodus and carpus of minor cheliped (left) showing disciform palmar crest (thick arrow) and high carpal ridge (thin arrow). (d) ventral view of ischium of minor cheliped (left) with four tubercles (arrows). Scale bars: a, c = 5.0 mm; b, d = 1.0 mm." captionText-3="Figure 14. Aegla santosi sp. nov., male holotype, CLE 19.3 mm (MZUEL 511). (a) third (thin arrow) and fourth (thick arrow) thoracic sternites. (b) dorsal view of second abdominal epimeron (right side) showing slightly concave anterior margin (thick arrow) and anterolateral angle with corneous scales (thin arrow). Scale bars = 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5727913" figureDoi-1="http://doi.org/10.5281/zenodo.5727915" figureDoi-2="http://doi.org/10.5281/zenodo.5727917" figureDoi-3="http://doi.org/10.5281/zenodo.5727919" httpUri-0="https://zenodo.org/record/5727913/files/figure.png" httpUri-1="https://zenodo.org/record/5727915/files/figure.png" httpUri-2="https://zenodo.org/record/5727917/files/figure.png" httpUri-3="https://zenodo.org/record/5727919/files/figure.png" pageId="16" pageNumber="2161">Figures 1114</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BA436C5FFC0771FFF044DC4D5EC6FB0" blockId="16.[192,950,1691,1716]" box="[192,950,1691,1716]" pageId="16" pageNumber="2161">
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urn:lsid:zoobank.org:act:
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<typeStatus id="54A08867FFC1771EFF644BD1D75169AC" box="[160,267,142,168]" pageId="17" pageNumber="2162" type="holotype">Holotype</typeStatus>
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</paragraph>
<paragraph id="8BA436C5FFC1771EFF644BEDD5816814" blockId="17.[160,1156,142,272]" pageId="17" pageNumber="2162">
<materialsCitation id="3B733C98FFC1771EFF644BEDD5816814" ID-GBIF-Occurrence="3411865314" collectingDate="2020-03-11" collectionCode="CLE" collectorName="G. M. Teixeira &amp; I. C. Marcal &amp; P. F. da Silva" country="Brazil" county="Male" elevation="792" latitude="-24.251862" location="Paranapanema River" longLatPrecision="1" longitude="-50.529606" municipality="Telemaco Borba" pageId="17" pageNumber="2162" specimenCount="1" stateProvince="Parana" typeStatus="holotype">
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[
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<quantity id="4CE39B20FFC1771EFEDE4BEDD7D869CF" box="[282,386,178,203]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" pageId="17" pageNumber="2162" unit="mm" value="19.3">19.3 mm</quantity>
],
<collectingCountry id="F30C7655FFC1771EFE514BEDD78E69CF" box="[405,468,178,203]" name="Brazil" pageId="17" pageNumber="2162">Brazil</collectingCountry>
,
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,
<collectingMunicipality id="6BC0ACBFFFC1771EFDFF4BEDD4AC69CF" box="[571,758,178,203]" pageId="17" pageNumber="2162">Telêmaco Borba</collectingMunicipality>
,
<location id="8EC4601EFFC1771EFCC64BEDD5B069CF" LSID="urn:lsid:plazi:treatment:03B287D3FFC0771DFE564D65D21C6BF1:8EC4601EFFC1771EFCC64BEDD5B069CF" box="[770,1002,178,203]" country="Brazil" county="Male" latitude="-24.251862" longLatPrecision="1" longitude="-50.529606" municipality="Telemaco Borba" name="Paranapanema River" pageId="17" pageNumber="2162" stateProvince="Parana">Paranapanema River</location>
basin,
<location id="8EC4601EFFC1771EFBFB4BEDD68D69E9" LSID="urn:lsid:plazi:treatment:03B287D3FFC0771DFE564D65D21C6BF1:8EC4601EFFC1771EFBFB4BEDD68D69E9" country="Brazil" county="Male" latitude="-24.251862" longLatPrecision="1" longitude="-50.529606" municipality="Telemaco Borba" name="Tibagi River" pageId="17" pageNumber="2162" stateProvince="Parana">Tibagi River</location>
sub-basin,
<location id="8EC4601EFFC1771EFE934B8BD45069E9" LSID="urn:lsid:plazi:treatment:03B287D3FFC0771DFE564D65D21C6BF1:8EC4601EFFC1771EFE934B8BD45069E9" box="[343,522,212,237]" country="Brazil" county="Male" latitude="-24.251862" longLatPrecision="1" longitude="-50.529606" municipality="Telemaco Borba" name="Codorna Branca" pageId="17" pageNumber="2162" stateProvince="Parana">Codorna Branca</location>
stream,
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24°15
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06.70”S
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,
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50°31
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46.58”W
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, altitude
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,
<collectorName id="26EE5313FFC1771EFBA84B8BD7416814" pageId="17" pageNumber="2162">G. M. Teixeira</collectorName>
,
<collectorName id="26EE5313FFC1771EFEE24BA8D7C26814" box="[294,408,247,272]" pageId="17" pageNumber="2162">I.C. Marçal</collectorName>
and
<collectorName id="26EE5313FFC1771EFE164BA8D40C6814" box="[466,598,247,272]" pageId="17" pageNumber="2162">P.F. da Silva</collectorName>
coll.,
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<collectingDate id="EFE1E9EDFFC1771EFD534BA8D5646814" box="[663,830,247,272]" pageId="17" pageNumber="2162" value="2020-03-11">11 March 2020</collectingDate>
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(MZUEL 511).
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</paragraph>
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<typeStatus id="54A08867FFC1771EFF644A61D74F685C" box="[160,277,318,344]" pageId="17" pageNumber="2162" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA436C5FFC1771EFF644A3DD58968C4" blockId="17.[160,1155,318,448]" pageId="17" pageNumber="2162">
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[
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[
<specimenCode id="DBBD9EBEFFC1771EFD534A3DD4A3687F" box="[663,761,354,379]" collectionCode="CLE" pageId="17" pageNumber="2162">CLE 14.1</specimenCode>
16.0 mm], same data as holotype (MZUEL 500)
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.
<materialsCitation id="3B733C98FFC1771EFEF84ADAD58968C4" ID-GBIF-Occurrence="3411865319" collectingDate="2020-03-11" collectionCode="CLE" collectorName="G. M. Teixeira &amp; I. C. Marcal &amp; P. F. da Silva" country="Brazil" county="Male" elevation="792" isTailEnumeration="true" latitude="-24.251862" location="Paranapanema River" longLatPrecision="1" longitude="-50.529606" municipality="Telemaco Borba" pageId="17" pageNumber="2162" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Parana" typeStatus="paratype">
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[
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17.0 mm] and
<specimenCount id="9D1DFD4CFFC1771EFDA84ADAD497689A" box="[620,717,389,414]" pageId="17" pageNumber="2162" type="female">1 female</specimenCount>
[
<collectionCode id="ED0AAE00FFC1771EFD1D4ADAD55F689A" box="[729,773,389,414]" pageId="17" pageNumber="2162">CLE</collectionCode>
<quantity id="4CE39B20FFC1771EFCC84ADAD529689A" box="[780,883,389,414]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" pageId="17" pageNumber="2162" unit="mm" value="16.9">16.9 mm</quantity>
], same locality, J.F.M. da Silva, E.S. da Silva and S.T. Bennemann coll.,
<date id="FFA51005FFC1771EFD574AF8D56D68C4" box="[659,823,423,448]" pageId="17" pageNumber="2162" value="2014-08-07">7 August 2014</date>
(MZUEL 470).
</materialsCitation>
</paragraph>
<paragraph id="8BA436C5FFC1771EFF644AB1D4586B0C" blockId="17.[159,1156,494,902]" box="[160,514,494,520]" pageId="17" pageNumber="2162">
<emphasis id="B96FEAD7FFC1771EFF644AB1D4586B0C" bold="true" box="[160,514,494,520]" italics="true" pageId="17" pageNumber="2162">Additional material examined</emphasis>
</paragraph>
<paragraph id="8BA436C5FFC1771DFF64494DD78E6873" blockId="17.[159,1156,494,902]" lastBlockId="18.[159,1156,142,375]" lastPageId="18" lastPageNumber="2163" pageId="17" pageNumber="2162">
<materialsCitation id="3B733C98FFC1771EFF64494DD59B6B4A" ID-GBIF-Occurrence="3411865302" collectingDate="2020-03-11" collectionCode="CLE" collectorName="Bold Systems" country="Brazil" county="Male" elevation="792" latitude="-24.251862" location="Paranapanema River" longLatPrecision="1" longitude="-50.529606" municipality="Telemaco Borba" pageId="17" pageNumber="2162" specimenCount="10" specimenCount-female="3" specimenCount-male="7" stateProvince="Parana">
<specimenCount id="9D1DFD4CFFC1771EFF64494DD6A86B2F" box="[160,242,530,555]" pageId="17" pageNumber="2162" type="male">7 males</specimenCount>
[size not recorded] and
<specimenCount id="9D1DFD4CFFC1771EFE32494DD4076B2F" box="[502,605,530,555]" pageId="17" pageNumber="2162" type="female">3 females</specimenCount>
[size not recorded], same data as holotype (MZUEL 505, genetic vouchers:
<collectorName id="26EE5313FFC1771EFE59496AD4746B4A" box="[413,558,565,590]" pageId="17" pageNumber="2162">Bold Systems</collectorName>
access
<taxonomicName id="4C1B4D46FFC1771EFD47496AD48F6B4A" authorityName="Leach" authorityYear="1820" box="[643,725,565,590]" class="Malacostraca" family="Aeglidae" genus="Aegla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="2171" phylum="Arthropoda" rank="genus">AEGLA</taxonomicName>
005-21,
<taxonomicName id="4C1B4D46FFC1771EFCE7496AD52C6B4A" authorityName="Leach" authorityYear="1820" box="[803,886,565,590]" class="Malacostraca" family="Aeglidae" genus="Aegla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="2171" phylum="Arthropoda" rank="genus">AEGLA</taxonomicName>
006-21)
</materialsCitation>
.
<materialsCitation id="3B733C98FFC1771EFC0F496AD5CD6B97" ID-GBIF-Occurrence="3411865320" collectionCode="CLE" country="Brazil" county="Telemaco Borba" elevation="761" latitude="-24.33325" location="Tibagi River" longLatPrecision="1" longitude="-50.560333" municipality="Paranapanema River" pageId="17" pageNumber="2162" specimenCount="1" specimenCount-male="1" stateProvince="Parana">
<specimenCount id="9D1DFD4CFFC1771EFC0F496AD24C6B4A" box="[971,1046,565,590]" pageId="17" pageNumber="2162" type="male">1 male</specimenCount>
[
<collectionCode id="ED0AAE00FFC1771EFBE7496AD2146B4A" box="[1059,1102,565,590]" pageId="17" pageNumber="2162">CLE</collectionCode>
<quantity id="4CE39B20FFC1771EFB91496AD6946B74" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="17" pageNumber="2162" unit="mm" value="13.3">13.3 mm</quantity>
],
<collectingCountry id="F30C7655FFC1771EFF274908D7456B74" box="[227,287,599,624]" name="Brazil" pageId="17" pageNumber="2162">Brazil</collectingCountry>
,
<collectingRegion id="49DFF827FFC1771EFEE94908D7236B74" box="[301,377,599,624]" country="Brazil" name="Parana" pageId="17" pageNumber="2162">Paraná</collectingRegion>
,
<collectingCounty id="62C54E49FFC1771EFE4C4908D4646B74" box="[392,574,599,624]" pageId="17" pageNumber="2162">Telêmaco Borba</collectingCounty>
,
<collectingMunicipality id="6BC0ACBFFFC1771EFD894908D56B6B74" box="[589,817,599,624]" pageId="17" pageNumber="2162">Paranapanema River</collectingMunicipality>
basin,
<location id="8EC4601EFFC1771EFC424908D2506B74" LSID="urn:lsid:plazi:treatment:03B287D3FFC0771DFE564D65D21C6BF1:8EC4601EFFC1771EFC424908D2506B74" box="[902,1034,599,624]" country="Brazil" county="Telemaco Borba" latitude="-24.33325" longLatPrecision="1" longitude="-50.560333" municipality="Paranapanema River" name="Tibagi River" pageId="17" pageNumber="2162" stateProvince="Parana">Tibagi River</location>
sub-basin,
<location id="8EC4601EFFC1771EFF5B4925D6B56B97" LSID="urn:lsid:plazi:treatment:03B287D3FFC0771DFE564D65D21C6BF1:8EC4601EFFC1771EFF5B4925D6B56B97" box="[159,239,634,659]" country="Brazil" county="Telemaco Borba" latitude="-24.33325" longLatPrecision="1" longitude="-50.560333" municipality="Paranapanema River" name="Varanal" pageId="17" pageNumber="2162" stateProvince="Parana">Varanal</location>
stream,
<geoCoordinate id="EE2F5002FFC1771EFEA64925D7AD6B97" box="[354,503,634,659]" degrees="24" direction="south" minutes="19" orientation="latitude" pageId="17" pageNumber="2162" precision="1" seconds="59.70" value="-24.33325">
24°19
<emphasis id="B96FEAD7FFC1771EFE644925D7FF6B95" bold="true" box="[416,421,634,657]" italics="true" pageId="17" pageNumber="2162">ʹ</emphasis>
59.70”S
</geoCoordinate>
,
<geoCoordinate id="EE2F5002FFC1771EFDCF4925D4F36B97" box="[523,681,634,659]" degrees="50" direction="west" minutes="33" orientation="longitude" pageId="17" pageNumber="2162" precision="1" seconds="37.20" value="-50.560333">
50°33
<emphasis id="B96FEAD7FFC1771EFD8C4925D4176B95" bold="true" box="[584,589,634,657]" italics="true" pageId="17" pageNumber="2162">ʹ</emphasis>
37.20”W
</geoCoordinate>
, altitude
<quantity id="4CE39B20FFC1771EFCE64925D5296B97" box="[802,883,634,659]" metricMagnitude="2" metricUnit="m" metricValue="7.609999999999999" pageId="17" pageNumber="2162" unit="m" value="761.0">
<elevation id="0036D1F6FFC1771EFCE64925D5296B97" box="[802,883,634,659]" metricMagnitude="2" metricUnit="m" metricValue="7.609999999999999" pageId="17" pageNumber="2162" unit="m" value="761.0">761 m</elevation>
</quantity>
, S
</materialsCitation>
<materialsCitation id="3B733C98FFC1771EFC534925D5F36B97" ID-GBIF-Occurrence="3411865322" box="[919,937,634,659]" collectionCode="CLE" country="Brazil" county="Telemaco Borba" elevation="761" isTailEnumeration="true" latitude="-24.33325" location="Tibagi River" longLatPrecision="1" longitude="-50.560333" municipality="Paranapanema River" pageId="17" pageNumber="2162" specimenCount="1" stateProvince="Parana">.T</materialsCitation>
. Bennemann coll.,
<date id="FFA51005FFC1771EFF5B49C2D7716BB2" box="[159,299,669,694]" pageId="17" pageNumber="2162" value="2005-05-19">19 May 2005</date>
(MZUEL 183).
<materialsCitation id="3B733C98FFC1771EFE0D49C2D5AF6BB2" ID-GBIF-Occurrence="3411865317" box="[457,1013,669,694]" collectionCode="CLE" country="Brazil" county="Telemaco Borba" elevation="761" isTailEnumeration="true" latitude="-24.33325" location="Tibagi River" longLatPrecision="1" longitude="-50.560333" municipality="Paranapanema River" pageId="17" pageNumber="2162" specimenCount="3" specimenCount-male="3" stateProvince="Parana">
<specimenCount id="9D1DFD4CFFC1771EFE0D49C2D4476BB2" box="[457,541,669,694]" pageId="17" pageNumber="2162" type="male">3 males</specimenCount>
[
<collectionCode id="ED0AAE00FFC1771EFDED49C2D40E6BB2" box="[553,596,669,694]" pageId="17" pageNumber="2162">CLE</collectionCode>
<quantity id="4CE39B20FFC1771EFD9E49C2D4AD6BB2" box="[602,759,669,694]" metricMagnitude="-2" metricUnit="m" metricValue="1.475" metricValueMax="1.68" metricValueMin="1.27" pageId="17" pageNumber="2162" unit="mm" value="14.75" valueMax="16.8" valueMin="12.7">12.716.8 mm</quantity>
], same locality and coll
</materialsCitation>
.,
<date id="FFA51005FFC1771EFBC649C2D2D86BB2" box="[1026,1154,669,694]" pageId="17" pageNumber="2162" value="2004-03">March 2004</date>
(MZUEL 349).
<materialsCitation id="3B733C98FFC1771EFEFE49E0D5686BDC" ID-GBIF-Occurrence="3411865321" box="[314,818,703,728]" collectionCode="CLE" country="Brazil" county="Telemaco Borba" elevation="761" isTailEnumeration="true" latitude="-24.33325" location="Tibagi River" longLatPrecision="1" longitude="-50.560333" municipality="Paranapanema River" pageId="17" pageNumber="2162" specimenCount="1" specimenCount-male="1" stateProvince="Parana">
<specimenCount id="9D1DFD4CFFC1771EFEFE49E0D7DC6BDC" box="[314,390,703,728]" pageId="17" pageNumber="2162" type="male">1 male</specimenCount>
[
<collectionCode id="ED0AAE00FFC1771EFE5749E0D7E46BDC" box="[403,446,703,728]" pageId="17" pageNumber="2162">CLE</collectionCode>
<quantity id="4CE39B20FFC1771EFE0349E0D4766BDC" box="[455,556,703,728]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="17" pageNumber="2162" unit="mm" value="20.5">20.5 mm</quantity>
], same locality and coll
</materialsCitation>
.,
<geoCoordinate id="EE2F5002FFC1771EFC8649E0D58D6BDC" box="[834,983,703,728]" degrees="24" direction="south" minutes="20" orientation="latitude" pageId="17" pageNumber="2162" precision="1" seconds="12.70" value="-24.336863">
24°20
<emphasis id="B96FEAD7FFC1771EFCBB499FD5DE6BD3" bold="true" box="[895,900,704,727]" italics="true" pageId="17" pageNumber="2162">ʹ</emphasis>
12.70”S
</geoCoordinate>
,
<geoCoordinate id="EE2F5002FFC1771EFC2649E0D2DA6BDC" box="[994,1152,703,728]" degrees="50" direction="west" minutes="32" orientation="longitude" pageId="17" pageNumber="2162" precision="1" seconds="03.20" value="-50.53422">
50°32
<emphasis id="B96FEAD7FFC1771EFBDB499FD27E6BD3" bold="true" box="[1055,1060,704,727]" italics="true" pageId="17" pageNumber="2162">ʹ</emphasis>
03.20”W
</geoCoordinate>
, altitude
<quantity id="4CE39B20FFC1771EFF3C49BDD7646BFF" box="[248,318,738,763]" metricMagnitude="2" metricUnit="m" metricValue="8.02" pageId="17" pageNumber="2162" unit="m" value="802.0">802 m</quantity>
,
<date id="FFA51005FFC1771EFE8349BDD4596BFF" box="[327,515,738,763]" pageId="17" pageNumber="2162" value="2005-02-11">11 February 2005</date>
(MZUEL 332).
<materialsCitation id="3B733C98FFC1771EFD5949BDD2266BFF" ID-GBIF-Occurrence="3411865318" box="[669,1148,738,763]" collectionCode="CLE" country="Brazil" county="Telemaco Borba" elevation="761" isTailEnumeration="true" latitude="-24.33325" location="Tibagi River" longLatPrecision="1" longitude="-50.560333" municipality="Paranapanema River" pageId="17" pageNumber="2162" specimenCount="1" specimenCount-male="1" stateProvince="Parana">
<specimenCount id="9D1DFD4CFFC1771EFD5949BDD4BF6BFF" box="[669,741,738,763]" pageId="17" pageNumber="2162" type="male">1 male</specimenCount>
[
<collectionCode id="ED0AAE00FFC1771EFD3449BDD5416BFF" box="[752,795,738,763]" pageId="17" pageNumber="2162">CLE</collectionCode>
<quantity id="4CE39B20FFC1771EFCE549BDD5D96BFF" box="[801,899,738,763]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="17" pageNumber="2162" unit="mm" value="16.4">16.4 mm</quantity>
], same locality and coll
</materialsCitation>
.,
<geoCoordinate id="EE2F5002FFC1771EFF5B485AD76F6A1A" box="[159,309,773,798]" degrees="24" direction="south" minutes="20" orientation="latitude" pageId="17" pageNumber="2162" precision="1" seconds="13.40" value="-24.337057">
24°20
<emphasis id="B96FEAD7FFC1771EFF19485AD6B86A18" bold="true" box="[221,226,773,796]" italics="true" pageId="17" pageNumber="2162">ʹ</emphasis>
13.40”S
</geoCoordinate>
,
<geoCoordinate id="EE2F5002FFC1771EFEF9485AD7866A1A" box="[317,476,773,798]" degrees="50" direction="west" minutes="35" orientation="longitude" pageId="17" pageNumber="2162" precision="1" seconds="23.10" value="-50.58975">
50°35
<emphasis id="B96FEAD7FFC1771EFEBF485AD7DA6A18" bold="true" box="[379,384,773,796]" italics="true" pageId="17" pageNumber="2162">ʹ</emphasis>
23.10”W
</geoCoordinate>
, altitude
<quantity id="4CE39B20FFC1771EFDFA485AD4DF6A1A" box="[574,645,773,798]" metricMagnitude="2" metricUnit="m" metricValue="6.87" pageId="17" pageNumber="2162" unit="m" value="687.0">687 m</quantity>
,
<date id="FFA51005FFC1771EFD4B485AD5086A1A" box="[655,850,773,798]" pageId="17" pageNumber="2162" value="2005-11-09">9 November 2005</date>
(MZUEL 344).
<materialsCitation id="3B733C98FFC1771EFC2B485AD4186A67" ID-GBIF-Occurrence="3411865303" collectionCode="CLE" collectorName="Harmonia" country="Brazil" county="Telemaco Borba" location="Tibagi River" municipality="Paranapanema River" pageId="17" pageNumber="2162" specimenCount="1" specimenCount-female="1" stateProvince="Parana">
<specimenCount id="9D1DFD4CFFC1771EFC2B485AD2166A1A" box="[1007,1100,773,798]" pageId="17" pageNumber="2162" type="female">1 female</specimenCount>
[
<collectionCode id="ED0AAE00FFC1771EFB9C485AD2D96A1A" box="[1112,1155,773,798]" pageId="17" pageNumber="2162">CLE</collectionCode>
17.0 mm],
<collectingCountry id="F30C7655FFC1771EFED64878D7146A44" box="[274,334,807,832]" name="Brazil" pageId="17" pageNumber="2162">Brazil</collectingCountry>
,
<collectingRegion id="49DFF827FFC1771EFE934878D7F96A44" box="[343,419,807,832]" country="Brazil" name="Parana" pageId="17" pageNumber="2162">Paraná</collectingRegion>
,
<collectingCounty id="62C54E49FFC1771EFE684878D4076A44" box="[428,605,807,832]" pageId="17" pageNumber="2162">Telêmaco Borba</collectingCounty>
,
<collectingMunicipality id="6BC0ACBFFFC1771EFDA34878D51C6A44" box="[615,838,807,832]" pageId="17" pageNumber="2162">Paranapanema River</collectingMunicipality>
basin,
<location id="8EC4601EFFC1771EFC554878D24A6A44" LSID="urn:lsid:plazi:treatment:03B287D3FFC0771DFE564D65D21C6BF1:8EC4601EFFC1771EFC554878D24A6A44" box="[913,1040,807,832]" country="Brazil" county="Telemaco Borba" municipality="Paranapanema River" name="Tibagi River" pageId="17" pageNumber="2162" stateProvince="Parana">Tibagi River</location>
sub-basin,
<collectorName id="26EE5313FFC1771EFF5B4815D7536A67" box="[159,265,842,867]" pageId="17" pageNumber="2162">Harmonia</collectorName>
stream, coordinates and coll
</materialsCitation>
. unknown,
<date id="FFA51005FFC1771EFD784815D5116A67" box="[700,843,842,867]" pageId="17" pageNumber="2162" value="2006-06-20">20 June 2006</date>
(MZUEL 414).
<materialsCitation id="3B733C98FFC1771EFC224815D4E56A82" ID-GBIF-Occurrence="3411865310" collectionCode="CLE" collectorName="Harmonia" country="Brazil" county="Telemaco Borba" location="Tibagi River" municipality="Paranapanema River" pageId="17" pageNumber="2162" specimenCount="4" specimenCount-female="4" stateProvince="Parana">
<specimenCount id="9D1DFD4CFFC1771EFC224815D2146A67" box="[998,1102,842,867]" pageId="17" pageNumber="2162" type="female">4 females</specimenCount>
[
<collectionCode id="ED0AAE00FFC1771EFB9C4815D2D96A67" box="[1112,1155,842,867]" pageId="17" pageNumber="2162">CLE</collectionCode>
<quantity id="4CE39B20FFC1771EFF5B4832D7616A82" box="[159,315,877,902]" metricMagnitude="-2" metricUnit="m" metricValue="1.415" metricValueMax="1.58" metricValueMin="1.25" pageId="17" pageNumber="2162" unit="mm" value="14.15" valueMax="15.8" valueMin="12.5">12.515.8 mm</quantity>
], same locality, coordinates and coll
</materialsCitation>
. unknown,
<date id="FFA51005FFC1771EFCFE4832D5806A82" box="[826,986,877,902]" pageId="17" pageNumber="2162" value="2012-03-16">16 March 2012</date>
(MZUEL 415).
<materialsCitation id="3B733C98FFC1771DFBB14832D7B869E9" ID-GBIF-Occurrence="3411865306" collectionCode="CLE" collectorName="Moinho Velho" country="Brazil" county="Telemaco Borba" elevation="739" lastPageId="18" lastPageNumber="2163" latitude="-24.221762" location="Tibagi River" longLatPrecision="1" longitude="-50.62845" municipality="Paranapanema River" pageId="17" pageNumber="2162" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Parana">
<specimenCount id="9D1DFD4CFFC1771DFBB14832D68F69A3" lastPageId="18" lastPageNumber="2163" pageId="17" pageNumber="2162" type="male">1 male</specimenCount>
[
<collectionCode id="ED0AAE00FFC2771DFF2D4BD1D74E69A3" box="[233,276,142,167]" pageId="18" pageNumber="2163">CLE</collectionCode>
<quantity id="4CE39B20FFC2771DFEE04BD1D7D469A3" box="[292,398,142,167]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="18" pageNumber="2163" unit="mm" value="15.2">15.2 mm</quantity>
] and
<specimenCount id="9D1DFD4CFFC2771DFE194BD1D41E69A3" box="[477,580,142,167]" pageId="18" pageNumber="2163" type="female">1 female</specimenCount>
[
<collectionCode id="ED0AAE00FFC2771DFD9C4BD1D4D969A3" box="[600,643,142,167]" pageId="18" pageNumber="2163">CLE</collectionCode>
<quantity id="4CE39B20FFC2771DFD574BD1D4A569A3" box="[659,767,142,167]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="18" pageNumber="2163" unit="mm" value="15.4">15.4 mm</quantity>
],
<collectingCountry id="F30C7655FFC2771DFCDC4BD1D50E69A3" box="[792,852,142,167]" name="Brazil" pageId="18" pageNumber="2163">Brazil</collectingCountry>
,
<collectingRegion id="49DFF827FFC2771DFCA24BD1D5E869A3" box="[870,946,142,167]" country="Brazil" name="Parana" pageId="18" pageNumber="2163">Paraná</collectingRegion>
,
<collectingCounty id="62C54E49FFC2771DFC014BD1D2DA69A3" box="[965,1152,142,167]" pageId="18" pageNumber="2163">Telêmaco Borba</collectingCounty>
,
<collectingMunicipality id="6BC0ACBFFFC2771DFF5B4BEED72569CE" box="[159,383,177,202]" pageId="18" pageNumber="2163">Paranapanema River</collectingMunicipality>
basin,
<location id="8EC4601EFFC2771DFE094BEED41769CE" LSID="urn:lsid:plazi:treatment:03B287D3FFC0771DFE564D65D21C6BF1:8EC4601EFFC2771DFE094BEED41769CE" box="[461,589,177,202]" country="Brazil" county="Telemaco Borba" latitude="-24.221762" longLatPrecision="1" longitude="-50.62845" municipality="Paranapanema River" name="Tibagi River" pageId="18" pageNumber="2163" stateProvince="Parana">Tibagi River</location>
sub-basin,
<collectorName id="26EE5313FFC2771DFD0D4BEED53969CE" box="[713,867,177,202]" pageId="18" pageNumber="2163">Moinho Velho</collectorName>
stream,
<geoCoordinate id="EE2F5002FFC2771DFC054BEED20C69CE" box="[961,1110,177,202]" degrees="24" direction="south" minutes="13" orientation="latitude" pageId="18" pageNumber="2163" precision="1" seconds="18.34" value="-24.221762">
24°13
<emphasis id="B96FEAD7FFC2771DFC3A4BEED25969CC" bold="true" box="[1022,1027,177,200]" italics="true" pageId="18" pageNumber="2163">ʹ</emphasis>
18.34”S
</geoCoordinate>
,
<geoCoordinate id="EE2F5002FFC2771DFBA44BEED74169E9" degrees="50" direction="west" minutes="37" orientation="longitude" pageId="18" pageNumber="2163" precision="1" seconds="42.42" value="-50.62845">
50° 37
<emphasis id="B96FEAD7FFC2771DFF7E4B8BD6E569EF" bold="true" box="[186,191,212,235]" italics="true" pageId="18" pageNumber="2163">ʹ</emphasis>
42.42”W
</geoCoordinate>
, altitude
<quantity id="4CE39B20FFC2771DFE444B8BD79269E9" box="[384,456,212,237]" metricMagnitude="2" metricUnit="m" metricValue="7.39" pageId="18" pageNumber="2163" unit="m" value="739.0">
<elevation id="0036D1F6FFC2771DFE444B8BD79269E9" box="[384,456,212,237]" metricMagnitude="2" metricUnit="m" metricValue="7.39" pageId="18" pageNumber="2163" unit="m" value="739.0">739 m</elevation>
</quantity>
, S
</materialsCitation>
<materialsCitation id="3B733C98FFC2771DFE264B8BD7AE69E9" ID-GBIF-Occurrence="3411865313" box="[482,500,212,237]" collectionCode="CLE" collectorName="Moinho Velho" country="Brazil" county="Telemaco Borba" elevation="739" isTailEnumeration="true" latitude="-24.221762" location="Tibagi River" longLatPrecision="1" longitude="-50.62845" municipality="Paranapanema River" pageId="18" pageNumber="2163" specimenCount="1" stateProvince="Parana">.T</materialsCitation>
. Bennemann coll.,
<date id="FFA51005FFC2771DFD054B8BD53B69E9" box="[705,865,212,237]" pageId="18" pageNumber="2163" value="2014-08-07">7 August 2014</date>
(MZUEL 426).
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<specimenCount id="9D1DFD4CFFC2771DFBC54B8BD21169E9" box="[1025,1099,212,237]" pageId="18" pageNumber="2163" type="male">1 male</specimenCount>
[
<collectionCode id="ED0AAE00FFC2771DFB9C4B8BD2D969E9" box="[1112,1155,212,237]" pageId="18" pageNumber="2163">CLE</collectionCode>
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], same locality, J
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<materialsCitation id="3B733C98FFC2771DFE7D4BA9D796680B" ID-GBIF-Occurrence="3411865307" box="[441,460,246,271]" collectionCode="CLE" collectorName="Moinho Velho" country="Brazil" county="Telemaco Borba" elevation="739" isTailEnumeration="true" latitude="-24.221762" location="Tibagi River" longLatPrecision="1" longitude="-50.62845" municipality="Paranapanema River" pageId="18" pageNumber="2163" specimenCount="1" stateProvince="Parana">.F</materialsCitation>
.M. da Silva coll.,
<date id="FFA51005FFC2771DFD454BA9D51F680B" box="[641,837,246,271]" pageId="18" pageNumber="2163" value="2014-11-03">3 November 2014</date>
(MZUEL 427).
<materialsCitation id="3B733C98FFC2771DFC274BA9D5F26851" ID-GBIF-Occurrence="3411865315" collectionCode="CLE" country="Brazil" county="Telemaco Borba" elevation="736" latitude="-24.174385" location="Tibagi River" longLatPrecision="1" longitude="-50.62457" municipality="Paranapanema River" pageId="18" pageNumber="2163" specimenCode="CLE 14.4" specimenCount="2" specimenCount-female="2" stateProvince="Parana">
<specimenCount id="9D1DFD4CFFC2771DFC274BA9D217680B" box="[995,1101,246,271]" pageId="18" pageNumber="2163" type="female">2 females</specimenCount>
[
<specimenCode id="DBBD9EBEFFC2771DFB9C4BA9D6976836" collectionCode="CLE" pageId="18" pageNumber="2163">CLE 14.4</specimenCode>
15.0 mm],
<collectingCountry id="F30C7655FFC2771DFE884A46D7D26836" box="[332,392,281,306]" name="Brazil" pageId="18" pageNumber="2163">Brazil</collectingCountry>
,
<collectingRegion id="49DFF827FFC2771DFE564A46D7846836" box="[402,478,281,306]" country="Brazil" name="Parana" pageId="18" pageNumber="2163">Paraná</collectingRegion>
,
<collectingCounty id="62C54E49FFC2771DFE234A46D4C36836" box="[487,665,281,306]" pageId="18" pageNumber="2163">Telêmaco Borba</collectingCounty>
,
<collectingMunicipality id="6BC0ACBFFFC2771DFD674A46D5D86836" box="[675,898,281,306]" pageId="18" pageNumber="2163">Paranapanema River</collectingMunicipality>
basin,
<location id="8EC4601EFFC2771DFC0A4A46D2176836" LSID="urn:lsid:plazi:treatment:03B287D3FFC0771DFE564D65D21C6BF1:8EC4601EFFC2771DFC0A4A46D2176836" box="[974,1101,281,306]" country="Brazil" county="Telemaco Borba" latitude="-24.174385" longLatPrecision="1" longitude="-50.62457" municipality="Paranapanema River" name="Tibagi River" pageId="18" pageNumber="2163" stateProvince="Parana">Tibagi River</location>
subbasin,
<location id="8EC4601EFFC2771DFF2C4A63D7606851" LSID="urn:lsid:plazi:treatment:03B287D3FFC0771DFE564D65D21C6BF1:8EC4601EFFC2771DFF2C4A63D7606851" box="[232,314,316,341]" country="Brazil" county="Telemaco Borba" latitude="-24.174385" longLatPrecision="1" longitude="-50.62457" municipality="Paranapanema River" name="Colonia" pageId="18" pageNumber="2163" stateProvince="Parana">Colônia</location>
stream,
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24°10
<emphasis id="B96FEAD7FFC2771DFE1D4A63D7846857" bold="true" box="[473,478,316,339]" italics="true" pageId="18" pageNumber="2163">ʹ</emphasis>
27.79”S
</geoCoordinate>
,
<geoCoordinate id="EE2F5002FFC2771DFDF84A63D4806851" box="[572,730,316,341]" degrees="50" direction="west" minutes="37" orientation="longitude" pageId="18" pageNumber="2163" precision="1" seconds="28.45" value="-50.62457">
50°37
<emphasis id="B96FEAD7FFC2771DFDBD4A63D4246857" bold="true" box="[633,638,316,339]" italics="true" pageId="18" pageNumber="2163">ʹ</emphasis>
28.45”W
</geoCoordinate>
, altitude
<quantity id="4CE39B20FFC2771DFC864A63D5D66851" box="[834,908,316,341]" metricMagnitude="2" metricUnit="m" metricValue="7.359999999999999" pageId="18" pageNumber="2163" unit="m" value="736.0">
<elevation id="0036D1F6FFC2771DFC864A63D5D66851" box="[834,908,316,341]" metricMagnitude="2" metricUnit="m" metricValue="7.359999999999999" pageId="18" pageNumber="2163" unit="m" value="736.0">736 m</elevation>
</quantity>
, S
</materialsCitation>
<materialsCitation id="3B733C98FFC2771DFC6C4A63D5E06851" ID-GBIF-Occurrence="3411865324" box="[936,954,316,341]" collectionCode="CLE" country="Brazil" county="Telemaco Borba" elevation="736" isTailEnumeration="true" latitude="-24.174385" location="Tibagi River" longLatPrecision="1" longitude="-50.62457" municipality="Paranapanema River" pageId="18" pageNumber="2163" specimenCount="1" stateProvince="Parana">.T</materialsCitation>
. Bennemann coll.,
<date id="FFA51005FFC2771DFF5B4A01D7676873" box="[159,317,350,375]" pageId="18" pageNumber="2163" value="2014-08-07">7 August 2014</date>
(MZUEL 437).
</paragraph>
</subSubSection>
<caption id="DF64664DFFC1771EFF5B4D4AD42B6FBB" ID-DOI="http://doi.org/10.5281/zenodo.5727913" ID-Zenodo-Dep="5727913" httpUri="https://zenodo.org/record/5727913/files/figure.png" pageId="17" pageNumber="2162" startId="17.[159,224,1557,1580]" targetBox="[263,1053,964,1521]" targetPageId="17">
<paragraph id="8BA436C5FFC1771EFF5B4D4AD42B6FBB" blockId="17.[159,1156,1557,1727]" pageId="17" pageNumber="2162">
<emphasis id="B96FEAD7FFC1771EFF5B4D4AD7506F28" bold="true" box="[159,266,1557,1580]" pageId="17" pageNumber="2162">Figure 11.</emphasis>
<taxonomicName id="4C1B4D46FFC1771EFED64D4AD7CE6F28" box="[274,404,1557,1580]" class="Malacostraca" family="Aeglidae" genus="Aegla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="2162" phylum="Arthropoda" rank="species" species="santosi" status="sp. nov.">
<emphasis id="B96FEAD7FFC1771EFED64D4AD7CE6F28" box="[274,404,1557,1580]" italics="true" pageId="17" pageNumber="2162">Aegla santosi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C57ACFFC1771EFE594D4AD7B56F28" box="[413,495,1557,1580]" pageId="17" pageNumber="2162" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype, CLE 19.3 mm, Codorna Branca stream, Telêmaco Borba, Paraná, Brazil (MZUEL 511). (a) dorsal view of the cephalothorax and anterior portion of the abdomen. (b) ventral view of ischium of minor cheliped (left) showing four tubercles (arrow). (c) third thoracic sternite truncate (arrow). (d) dorsal view of second abdominal epimeron (right side) showing anterolateral angle with corneous scale (white arrow) and anterior margin slightly concave (black arrow). Scale bars: a = 5.0 mm; bd = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C301654EFFC2771DFF644AFCD21C6BF1" pageId="18" pageNumber="2163" type="diagnosis">
<paragraph id="8BA436C5FFC2771DFF644AFCD74E68B9" blockId="18.[160,1156,419,757]" box="[160,276,419,445]" pageId="18" pageNumber="2163">
<emphasis id="B96FEAD7FFC2771DFF644AFCD74E68B9" bold="true" box="[160,276,419,445]" italics="true" pageId="18" pageNumber="2163">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BA436C5FFC2771DFF644A98D21C6BF1" blockId="18.[160,1156,419,757]" pageId="18" pageNumber="2163">Rostrum triangular, narrow base, extending beyond distal apex of compound eyes, carinate along entire length. Subrostral process well developed, occupying proximal third of subrostral margin, anterior and posterior margins forming obtuse angle. Orbital and extra-orbital sinuses deep. Anterolateral spines reaching basal margin of cornea. Epigastric prominences pronounced. Protogastric lobes pronounced. Areola trapezoidal. Cardiac area subrectangular. Epibranchial area slightly elongated. Lobe on proximal dorsal margin of dactylus rudimentary or absent. Carpal ridge high on outer surface of carpus. Ventromesial border of cheliped ischium with 4 or 5 tubercles. Ventral angles of third and fourth abdominal epimeron with corneous scales apically. Uropods wide.</paragraph>
</subSubSection>
</treatment>
</document>