320 lines
49 KiB
XML
320 lines
49 KiB
XML
<document id="98947786E95CBD6F67F404BAD3C5CED7" ID-DOI="10.5281/zenodo.5353780" ID-ISSN="2345-7600" ID-Zenodo-Dep="5353780" ID-ZooBank="8493E540-DB48-4CA6-8A73-9D232D5546E1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630212606084" checkinUser="tatiana" docAuthor="Grinang, Jongkar & Ng, Peter K. L." docDate="2014" docId="03AE760B7024FFBCFC79F936C1A349B5" docLanguage="en" docName="62rbz389-395.pdf" docOrigin="Raffles Bulletin of Zoology 62" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Thelphusula pueh Grinang & Ng 2014, new species" docType="treatment" docUuid="F6BD9559-7E88-4785-9413-7811F16D1862" docUuidSource="ZooBank" docVersion="4" lastPageNumber="394" masterDocId="FF970E737025FFB9FFC5FFC8C56C4C7D" masterDocTitle="Taxonomy of the freshwater crab Coccusa cristicervix Ng & Grinang, 2004 (Decapoda: Brachyura: Gecarcinucidae), with description of a new species of Thelphusula from Sarawak, Malaysia, Borneo" masterLastPageNumber="395" masterPageNumber="389" pageNumber="390" updateTime="1699187930052" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="EA262D71AC8CB2D4AFBE31CF1DB1A9E4">Taxonomy of the freshwater crab Coccusa cristicervix Ng & Grinang, 2004 (Decapoda: Brachyura: Gecarcinucidae), with description of a new species of Thelphusula from Sarawak, Malaysia, Borneo</mods:title>
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<mods:namePart id="22AD449A502BB3AC53A56F5BCE1129CE">Grinang, Jongkar</mods:namePart>
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<mods:namePart id="9C8F9822ED83E8F40C30CD6DB13A32ED">Ng, Peter K. L.</mods:namePart>
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<mods:title id="66FCB69AC3E120A7DBAB209B59123300">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="D1079DD02C9CBC30E7F591A747E68802">
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<mods:date id="8FD7B2E283DDCA072C05F132676B8E10">2014</mods:date>
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<mods:number id="B5B9B54678B9C9B29B15486EC67FF991">2014-06-27</mods:number>
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<mods:number id="8462C2F6CEE905DC5081E00BA9796273">62</mods:number>
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<heading id="D0F070717024FFB8FC79F936C0704B68" bold="true" box="[956,1308,1790,1813]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="390" reason="2">
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<taxonomicName id="4C07BC9E7024FFB8FC79F936C1EB4B68" ID-CoL="7C6BQ" authority="Grinang & Ng, 2014" authorityName="Grinang & Ng" authorityYear="2014" box="[956,1159,1790,1813]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="pueh" status="sp. nov.">
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<emphasis id="B9731B0F7024FFB8FC79F936C1EB4B68" bold="true" box="[956,1159,1790,1813]" italics="true" pageId="1" pageNumber="390">Thelphusula pueh</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A240A6747024FFB8FB53F936C0704B68" box="[1174,1308,1790,1813]" pageId="1" pageNumber="390" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C31D94967024FFB8FBCBF8D6C1A64B48" box="[1038,1226,1822,1845]" pageId="1" pageNumber="390" type="description">
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<paragraph id="8BB8C71D7024FFB8FBCBF8D6C1A64B48" blockId="1.[956,1308,1790,1845]" box="[1038,1226,1822,1845]" pageId="1" pageNumber="390">
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(
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<figureCitation id="133CDB987024FFB8FBD3F8D6C10F4B48" box="[1046,1123,1822,1845]" captionStart="Fig" captionStartId="3.[113,148,2034,2055]" captionTargetBox="[120,1445,189,2010]" captionTargetId="figure-73@3.[120,1445,189,2010]" captionTargetPageId="3" captionText="Fig. 2. Thelphusula pueh, new species. Holotype male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353784" httpUri="https://zenodo.org/record/5353784/files/figure.png" pageId="1" pageNumber="390">Figs. 2</figureCitation>
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,
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<figureCitation id="133CDB987024FFB8FBB4F8D6C1CB4B48" box="[1137,1191,1822,1845]" captionStart="Fig" captionStartId="4.[136,171,2004,2025]" captionTargetBox="[136,1485,266,1950]" captionTargetId="figure-7@4.[108,1500,238,1953]" captionTargetPageId="4" captionText="Fig. 3. A–E, Thelphusula cristicervix (Ng & Grinang, 2004), male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak; F–J, Thelphusula pueh, new species, male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, F, ventral view of left G1; B, G, ventral view of left G2; C, H, dorsal view of left G1; D, I, ventral view of terminal segment of left G1; E, J, dorsal view of terminal segment of left G1. Scale bars: A–C, F–H = 1.0 mm; D, E, I, J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5353786" httpUri="https://zenodo.org/record/5353786/files/figure.png" pageId="1" pageNumber="390">3F–J</figureCitation>
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,
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<figureCitation id="133CDB987024FFB8FB71F8D6C1AE4B48" box="[1204,1218,1822,1845]" captionStart="Fig" captionStartId="5.[113,148,2006,2027]" captionTargetBox="[115,753,1453,1949]" captionTargetId="figure-96@5.[114,756,1450,1982]" captionTargetPageId="5" captionText="Fig. 5. Colours in life, Thelphusula pueh, new species. A, paratype female (16.3 × 12.9 mm) (SBC); B, paratype female (13.0 × 10.1 mm) (SBC). Both specimens from Kampung Sebako, Pueh, Lundu District, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5353790" httpUri="https://zenodo.org/record/5353790/files/figure.png" pageId="1" pageNumber="390">5</figureCitation>
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)
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</paragraph>
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<emphasis id="B9731B0F7024FFB8FCE8F896C1744B08" bold="true" box="[813,1048,1886,1909]" pageId="1" pageNumber="390">Material examined.</emphasis>
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<materialsCitation id="3B6FCD407024FFB8FBE6F896C1D74BA8" collectingDate="2013-11-29" collectionCode="ZRC" collectorName="R. Sujang" country="Malaysia" elevation="26" location="Lundu District" pageId="1" pageNumber="390" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54BC79BF7024FFB8FBE6F896C1E34B08" box="[1059,1167,1886,1909]" pageId="1" pageNumber="390" type="holotype">Holotype</typeStatus>
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: male (16.2
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<emphasis id="B9731B0F7024FFB8FAE7F896C05D4B08" bold="true" box="[1314,1329,1886,1909]" pageId="1" pageNumber="390">×</emphasis>
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<quantity id="4CFF6AF87024FFB8FAF9F896C0C84B08" box="[1340,1444,1886,1909]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="1" pageNumber="390" unit="mm" value="12.1">12.1 mm</quantity>
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) (
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<collectionCode id="ED165FD87024FFB8FCF1F8B6C6004BE8" box="[820,876,1918,1941]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="390">ZRC</collectionCode>
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), edge of rice field with swampy vegetation,
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<elevation id="002A202E7024FFB8FAF1F8B6C6E64BC8" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="390" unit="m" value="26.0">
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<quantity id="4CFF6AF87024FFB8FAF1F8B6C0044BE8" box="[1332,1384,1918,1941]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="390" unit="m" value="26.0">26 m</quantity>
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above sea level
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</elevation>
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, Kampung Sebako, Pueh,
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<location id="8ED891C67024FFB8FB64F856C0534BC8" LSID="urn:lsid:plazi:treatment:03AE760B7024FFBCFC79F936C1A349B5:8ED891C67024FFB8FB64F856C0534BC8" box="[1185,1343,1950,1973]" country="Malaysia" name="Lundu District" pageId="1" pageNumber="390">Lundu District</location>
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,
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<collectingCountry id="F310878D7024FFB8FA8CF856C0C44BC8" box="[1353,1448,1950,1973]" name="Malaysia" pageId="1" pageNumber="390">Sarawak</collectingCountry>
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, coll.
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<collectorName id="26F2A2CB7024FFB8FCA0F876C6B94BA8" box="[869,981,1982,2005]" pageId="1" pageNumber="390">R. Sujang</collectorName>
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,
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<date id="FFB9E1DD7024FFB8FC26F876C1D74BA8" box="[995,1211,1982,2005]" pageId="1" pageNumber="390" value="2013-11-29">
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<collectingDate id="EFFD18357024FFB8FC26F876C1D74BA8" box="[995,1211,1982,2005]" pageId="1" pageNumber="390" value="2013-11-29">29 November 2013</collectingDate>
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</date>
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</materialsCitation>
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.
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<materialsCitation id="3B6FCD407024FFB8FB0DF876C1AB4B88" collectionCode="ZRC" country="Malaysia" pageId="1" pageNumber="390" specimenCount="6" specimenCount-female="6" typeStatus="paratype">
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<typeStatus id="54BC79BF7024FFB8FB0DF876C0584BA8" box="[1224,1332,1982,2005]" pageId="1" pageNumber="390" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9D010C947024FFB8FA84F876C0C74BA8" box="[1345,1451,1982,2005]" pageId="1" pageNumber="390" type="female">6 females</specimenCount>
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(
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<quantity id="4CFF6AF87024FFB8FCF1F816C6A14B88" box="[820,973,2014,2037]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" metricValueMax="1.98" metricValueMin="1.26" pageId="1" pageNumber="390" unit="mm" value="16.2" valueMax="19.8" valueMin="12.6">12.6–19.8 mm</quantity>
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<emphasis id="B9731B0F7024FFB8FC17F816C68D4B88" bold="true" box="[978,993,2014,2037]" pageId="1" pageNumber="390">×</emphasis>
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<quantity id="4CFF6AF87024FFB8FC23F816C11F4B88" box="[998,1139,2014,2037]" metricMagnitude="-2" metricUnit="m" metricValue="1.275" metricValueMax="1.56" metricValueMin="0.99" pageId="1" pageNumber="390" unit="mm" value="12.75" valueMax="15.6" valueMin="9.9">9.9–15.6 mm</quantity>
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) (
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<collectionCode id="ED165FD87024FFB8FB42F816C1D34B88" box="[1159,1215,2014,2037]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="390">ZRC</collectionCode>
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)
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</materialsCitation>
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,
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||
<materialsCitation id="3B6FCD407024FFB8FB15F816C0244468" collectingDate="2013-11-29" collectionCode="SBC" collectorName="R. Sujang" country="Malaysia" elevation="26" location="Lundu District" pageId="1" pageNumber="390" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
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<specimenCount id="9D010C947024FFB8FB15F816C0584B88" box="[1232,1332,2014,2037]" pageId="1" pageNumber="390" type="female">4 females</specimenCount>
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(
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<quantity id="4CFF6AF87024FFB8FA84F816C63A4468" metricMagnitude="-2" metricUnit="m" metricValue="1.76" metricValueMax="1.94" metricValueMin="1.58" pageId="1" pageNumber="390" unit="mm" value="17.6" valueMax="19.4" valueMin="15.8">15.8–19.4 mm</quantity>
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<emphasis id="B9731B0F7024FFB8FC9EF836C6064468" bold="true" box="[859,874,2046,2069]" pageId="1" pageNumber="390">×</emphasis>
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<quantity id="4CFF6AF87024FFB8FCAAF836C16B4468" box="[879,1031,2046,2069]" metricMagnitude="-2" metricUnit="m" metricValue="1.295" metricValueMax="1.45" metricValueMin="1.14" pageId="1" pageNumber="390" unit="mm" value="12.95" valueMax="14.5" valueMin="11.4">11.4–14.5 mm</quantity>
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) (
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<collectionCode id="ED165FD87024FFB8FBDEF836C13D4468" box="[1051,1105,2046,2069]" country="USA" lsid="urn:lsid:biocol.org:col:12519" name="University of California" pageId="1" pageNumber="390" type="Herbarium">SBC</collectionCode>
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), same data as holotype
|
||
</materialsCitation>
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;
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||
<materialsCitation id="3B6FCD407024FFBCFA94F836C7384C88" collectingDate="2013-11-10" collectionCode="ZRC" collectorName="L. Noun" country="Malaysia" elevation="31" lastPageId="5" lastPageNumber="394" location="Bau District" pageId="1" pageNumber="390" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<specimenCount id="9D010C947024FFB8FA94F836C0C04468" box="[1361,1452,2046,2069]" pageId="1" pageNumber="390" type="female">1 female</specimenCount>
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(24.1 ×
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<quantity id="4CFF6AF87024FFB8FC45F7D6C68C4448" box="[896,992,2078,2101]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="1" pageNumber="390" unit="mm" value="14.8">14.8 mm</quantity>
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) (
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<collectionCode id="ED165FD87024FFB8FC30F7D6C1424448" box="[1013,1070,2078,2101]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="390">ZRC</collectionCode>
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), in rice field,
|
||
<elevation id="002A202E7024FFB8FB03F7D6C0CB4448" box="[1222,1447,2078,2101]" metricMagnitude="1" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="390" unit="m" value="31.0">
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<quantity id="4CFF6AF87024FFB8FB03F7D6C1904448" box="[1222,1276,2078,2101]" metricMagnitude="1" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="390" unit="m" value="31.0">31 m</quantity>
|
||
above sea level
|
||
</elevation>
|
||
, Kampung Peninjau Lama, Gunung Serambu,
|
||
<location id="8ED891C67020FFBCFDA0FF76C7804CA8" LSID="urn:lsid:plazi:treatment:03AE760B7024FFBCFC79F936C1A349B5:8ED891C67020FFBCFDA0FF76C7804CA8" box="[613,748,190,213]" country="Malaysia" name="Bau District" pageId="5" pageNumber="394">Bau District</location>
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,
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||
<collectingCountry id="F310878D7020FFBCFFB4FF16C5BC4C88" box="[113,208,222,245]" name="Malaysia" pageId="5" pageNumber="394">Sarawak</collectingCountry>
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, coll.
|
||
<collectorName id="26F2A2CB7020FFBCFED1FF16C41D4C88" box="[276,369,222,245]" pageId="5" pageNumber="394">L. Noun</collectorName>
|
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,
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||
<date id="FFB9E1DD7020FFBCFEBBFF16C7384C88" box="[382,596,222,245]" pageId="5" pageNumber="394" value="2013-11-10">
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<collectingDate id="EFFD18357020FFBCFEBBFF16C7384C88" box="[382,596,222,245]" pageId="5" pageNumber="394" value="2013-11-10">10 November 2013</collectingDate>
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</date>
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||
</materialsCitation>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
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<caption id="DF7897957027FFBBFF4DF83AC1D6443C" ID-DOI="http://doi.org/10.5281/zenodo.5353782" ID-Zenodo-Dep="5353782" httpUri="https://zenodo.org/record/5353782/files/figure.png" pageId="2" pageNumber="391" startId="2.[136,171,2034,2055]" targetBox="[142,1467,188,2010]" targetPageId="2">
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<paragraph id="8BB8C71D7027FFBBFF4DF83AC1D6443C" blockId="2.[136,1474,2034,2113]" pageId="2" pageNumber="391">
|
||
Fig. 1.
|
||
<taxonomicName id="4C07BC9E7027FFBBFF17F83AC7C4447A" authority="(Ng & Grinang, 2004)" authorityName="Grinang & Ng" authorityYear="2014" baseAuthorityName="Ng & Grinang" baseAuthorityYear="2004" box="[210,680,2034,2055]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="cristicervix">
|
||
<emphasis id="B9731B0F7027FFBBFF17F83AC4D1447A" box="[210,445,2034,2055]" italics="true" pageId="2" pageNumber="391">Thelphusula cristicervix</emphasis>
|
||
(
|
||
<bibRefCitation id="EF96BAEC7027FFBBFE08F83AC7CD447A" author="Ng PKL & Grinang J" box="[461,673,2034,2055]" pageId="2" pageNumber="391" pagination="299 - 325" refId="ref4094" refString="Ng PKL & Grinang J (2004) Decapod crustaceans with descriptions of three new species. In: Yong HS Ng FSP & Yen EEL (eds.) Sarawak Bau Limestone Biodiversity. Sarawak Museum Journal, 59, No. 80 (New Series), Special Issue No. 6: 299 - 325." type="book chapter" year="2004">Ng & Grinang, 2004</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7897957026FFBAFFB4F83AC681443C" ID-DOI="http://doi.org/10.5281/zenodo.5353784" ID-Zenodo-Dep="5353784" httpUri="https://zenodo.org/record/5353784/files/figure.png" pageId="3" pageNumber="392" startId="3.[113,148,2034,2055]" targetBox="[120,1445,189,2010]" targetPageId="3">
|
||
<paragraph id="8BB8C71D7026FFBAFFB4F83AC681443C" blockId="3.[113,1452,2034,2113]" pageId="3" pageNumber="392">
|
||
Fig. 2.
|
||
<taxonomicName id="4C07BC9E7026FFBAFF7EF83AC405447A" authority="Grinang & Ng, 2014" authorityName="Grinang & Ng" authorityYear="2014" box="[187,361,2034,2055]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="pueh" status="sp. nov.">
|
||
<emphasis id="B9731B0F7026FFBAFF7EF83AC405447A" box="[187,361,2034,2055]" italics="true" pageId="3" pageNumber="392">Thelphusula pueh</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A240A6747026FFBAFEB2F83AC49C447A" box="[375,496,2034,2055]" pageId="3" pageNumber="392" rank="species">new species</taxonomicNameLabel>
|
||
. Holotype male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7897957021FFBDFF4DF81CC106443D" ID-DOI="http://doi.org/10.5281/zenodo.5353786" ID-Zenodo-Dep="5353786" httpUri="https://zenodo.org/record/5353786/files/figure.png" pageId="4" pageNumber="393" startId="4.[136,171,2004,2025]" targetBox="[136,1485,266,1950]" targetPageId="4">
|
||
<paragraph id="8BB8C71D7021FFBDFF4DF81CC106443D" blockId="4.[136,1474,2004,2112]" pageId="4" pageNumber="393">
|
||
Fig. 3. A–E,
|
||
<taxonomicName id="4C07BC9E7021FFBDFEC6F81CC7A74B94" authority="(Ng & Grinang, 2004)" authorityName="Grinang & Ng" authorityYear="2014" baseAuthorityName="Ng & Grinang" baseAuthorityYear="2004" box="[259,715,2004,2025]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="cristicervix">
|
||
<emphasis id="B9731B0F7021FFBDFEC6F81CC4804B94" box="[259,492,2004,2025]" italics="true" pageId="4" pageNumber="393">Thelphusula cristicervix</emphasis>
|
||
(
|
||
<bibRefCitation id="EF96BAEC7021FFBDFE3DF81CC7A94B94" author="Ng PKL & Grinang J" box="[504,709,2004,2025]" pageId="4" pageNumber="393" pagination="299 - 325" refId="ref4094" refString="Ng PKL & Grinang J (2004) Decapod crustaceans with descriptions of three new species. In: Yong HS Ng FSP & Yen EEL (eds.) Sarawak Bau Limestone Biodiversity. Sarawak Museum Journal, 59, No. 80 (New Series), Special Issue No. 6: 299 - 325." type="book chapter" year="2004">Ng & Grinang, 2004</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak; F–J,
|
||
<taxonomicName id="4C07BC9E7021FFBDFEDEF839C4A5447B" authority="Grinang & Ng, 2014" authorityName="Grinang & Ng" authorityYear="2014" box="[283,457,2033,2054]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="pueh" status="sp. nov.">
|
||
<emphasis id="B9731B0F7021FFBDFEDEF839C4A5447B" box="[283,457,2033,2054]" italics="true" pageId="4" pageNumber="393">Thelphusula pueh</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A240A6747021FFBDFE12F839C73C447B" box="[471,592,2033,2054]" pageId="4" pageNumber="393" rank="species">new species</taxonomicNameLabel>
|
||
, male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, F, ventral view of left G1; B, G, ventral view of left G2; C, H, dorsal view of left G1; D, I, ventral view of terminal segment of left G1; E, J, dorsal view of terminal segment of left G1. Scale bars: A–C, F–H = 1.0 mm; D, E, I, J = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31D94967020FFBCFFB4FED6C1CB4E5F" pageId="5" pageNumber="394" type="diagnosis">
|
||
<paragraph id="8BB8C71D7020FFBCFFB4FED6C0C04DE0" blockId="5.[113,753,286,469]" lastBlockId="5.[813,1452,190,413]" pageId="5" pageNumber="394">
|
||
<emphasis id="B9731B0F7020FFBCFFB4FED6C58A4D48" bold="true" box="[113,230,286,309]" pageId="5" pageNumber="394">Diagnosis.</emphasis>
|
||
Carapace gently ovate, wider than long, dorsal surface rugose, anterolateral margin convex, epibranchial tooth absent, cervical groove relatively shallow, shallow crista; postorbital cristae weak; H-shaped depression shallow (
|
||
<figureCitation id="133CDB987020FFBCFFBCFE56C5854DC8" box="[121,233,414,437]" captionStart="Fig" captionStartId="3.[113,148,2034,2055]" captionTargetBox="[120,1445,189,2010]" captionTargetId="figure-73@3.[120,1445,189,2010]" captionTargetPageId="3" captionText="Fig. 2. Thelphusula pueh, new species. Holotype male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353784" httpUri="https://zenodo.org/record/5353784/files/figure.png" pageId="5" pageNumber="394">Fig. 2A, B</figureCitation>
|
||
). Frontal margin deflexed; frontal median triangle absent (
|
||
<figureCitation id="133CDB987020FFBCFF08FE76C4464DA8" box="[205,298,446,469]" captionStart="Fig" captionStartId="3.[113,148,2034,2055]" captionTargetBox="[120,1445,189,2010]" captionTargetId="figure-73@3.[120,1445,189,2010]" captionTargetPageId="3" captionText="Fig. 2. Thelphusula pueh, new species. Holotype male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353784" httpUri="https://zenodo.org/record/5353784/files/figure.png" pageId="5" pageNumber="394">Fig. 2C</figureCitation>
|
||
). Median tooth on posterior margin of epistome subtruncate (
|
||
<figureCitation id="133CDB987020FFBCFBD1FF76C1094CA8" box="[1044,1125,190,213]" captionStart="Fig" captionStartId="3.[113,148,2034,2055]" captionTargetBox="[120,1445,189,2010]" captionTargetId="figure-73@3.[120,1445,189,2010]" captionTargetPageId="3" captionText="Fig. 2. Thelphusula pueh, new species. Holotype male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353784" httpUri="https://zenodo.org/record/5353784/files/figure.png" pageId="5" pageNumber="394">Fig. 2C</figureCitation>
|
||
).Ambulatory legs not distinctly elongated (
|
||
<figureCitation id="133CDB987020FFBCFC6DFF17C16E4C8B" box="[936,1026,223,246]" captionStart="Fig" captionStartId="3.[113,148,2034,2055]" captionTargetBox="[120,1445,189,2010]" captionTargetId="figure-73@3.[120,1445,189,2010]" captionTargetPageId="3" captionText="Fig. 2. Thelphusula pueh, new species. Holotype male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353784" httpUri="https://zenodo.org/record/5353784/files/figure.png" pageId="5" pageNumber="394">Fig. 2A</figureCitation>
|
||
). Major chela with large gap between fingers when closed (
|
||
<figureCitation id="133CDB987020FFBCFBCAFEC9C10D4D65" box="[1039,1121,257,280]" captionStart="Fig" captionStartId="3.[113,148,2034,2055]" captionTargetBox="[120,1445,189,2010]" captionTargetId="figure-73@3.[120,1445,189,2010]" captionTargetPageId="3" captionText="Fig. 2. Thelphusula pueh, new species. Holotype male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353784" httpUri="https://zenodo.org/record/5353784/files/figure.png" pageId="5" pageNumber="394">Fig. 2F</figureCitation>
|
||
). Male abdomen with somite 6 relatively short, lateral margins distinctly concave (
|
||
<figureCitation id="133CDB987020FFBCFA8FFEEAC0CC4D44" box="[1354,1440,290,313]" captionStart="Fig" captionStartId="3.[113,148,2034,2055]" captionTargetBox="[120,1445,189,2010]" captionTargetId="figure-73@3.[120,1445,189,2010]" captionTargetPageId="3" captionText="Fig. 2. Thelphusula pueh, new species. Holotype male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353784" httpUri="https://zenodo.org/record/5353784/files/figure.png" pageId="5" pageNumber="394">Fig. 2D</figureCitation>
|
||
). G1 terminal segment relatively cylindrical, straight, slightly more than 2 times length of subterminal segment (
|
||
<figureCitation id="133CDB987020FFBCFA92FEADC6304DE0" captionStart="Fig" captionStartId="4.[136,171,2004,2025]" captionTargetBox="[136,1485,266,1950]" captionTargetId="figure-7@4.[108,1500,238,1953]" captionTargetPageId="4" captionText="Fig. 3. A–E, Thelphusula cristicervix (Ng & Grinang, 2004), male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak; F–J, Thelphusula pueh, new species, male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, F, ventral view of left G1; B, G, ventral view of left G2; C, H, dorsal view of left G1; D, I, ventral view of terminal segment of left G1; E, J, dorsal view of terminal segment of left G1. Scale bars: A–C, F–H = 1.0 mm; D, E, I, J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5353786" httpUri="https://zenodo.org/record/5353786/files/figure.png" pageId="5" pageNumber="394">Fig. 3F, H–J</figureCitation>
|
||
). G2 distal segment short, curved, tip rounded (
|
||
<figureCitation id="133CDB987020FFBCFA8EFE4EC0F34DE0" box="[1355,1439,390,413]" captionStart="Fig" captionStartId="4.[136,171,2004,2025]" captionTargetBox="[136,1485,266,1950]" captionTargetId="figure-7@4.[108,1500,238,1953]" captionTargetPageId="4" captionText="Fig. 3. A–E, Thelphusula cristicervix (Ng & Grinang, 2004), male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak; F–J, Thelphusula pueh, new species, male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, F, ventral view of left G1; B, G, ventral view of left G2; C, H, dorsal view of left G1; D, I, ventral view of terminal segment of left G1; E, J, dorsal view of terminal segment of left G1. Scale bars: A–C, F–H = 1.0 mm; D, E, I, J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5353786" httpUri="https://zenodo.org/record/5353786/files/figure.png" pageId="5" pageNumber="394">Fig. 3G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB8C71D7020FFBCFCE8FE01C1CB4E5F" blockId="5.[813,1452,457,546]" pageId="5" pageNumber="394">
|
||
<emphasis id="B9731B0F7020FFBCFCE8FE01C6ED4D9D" bold="true" box="[813,897,457,480]" pageId="5" pageNumber="394">Colour.</emphasis>
|
||
In life, the species is dark brown overall on all the dorsal surfaces, with the palm of the chela and sometimes the legs more orange (
|
||
<figureCitation id="133CDB987020FFBCFBE7FDC3C1F64E5F" box="[1058,1178,523,546]" captionStart="Fig" captionStartId="5.[113,148,2006,2027]" captionTargetBox="[115,753,1453,1949]" captionTargetId="figure-96@5.[114,756,1450,1982]" captionTargetPageId="5" captionText="Fig. 5. Colours in life, Thelphusula pueh, new species. A, paratype female (16.3 × 12.9 mm) (SBC); B, paratype female (13.0 × 10.1 mm) (SBC). Both specimens from Kampung Sebako, Pueh, Lundu District, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5353790" httpUri="https://zenodo.org/record/5353790/files/figure.png" pageId="5" pageNumber="394">Fig. 5A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D94967020FFBCFCE8FD86C0C04EFB" pageId="5" pageNumber="394" type="etymology">
|
||
<paragraph id="8BB8C71D7020FFBCFCE8FD86C0C04EFB" blockId="5.[813,1452,590,646]" pageId="5" pageNumber="394">
|
||
<emphasis id="B9731B0F7020FFBCFCE8FD86C6C14E18" bold="true" box="[813,941,590,613]" pageId="5" pageNumber="394">Etymology.</emphasis>
|
||
The species is named after Pueh, the area where it was collected. The name is used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D94967020FFBCFCE8FD7AC1A349B5" pageId="5" pageNumber="394" type="discussion">
|
||
<paragraph id="8BB8C71D7020FFBCFCE8FD7AC1C4493F" blockId="5.[813,1453,690,1346]" pageId="5" pageNumber="394">
|
||
<emphasis id="B9731B0F7020FFBCFCE8FD7AC6F74EB4" bold="true" box="[813,923,690,713]" pageId="5" pageNumber="394">Remarks.</emphasis>
|
||
<taxonomicName id="4C07BC9E7020FFBCFC5AFD7AC1F34EB4" authorityName="Grinang & Ng" authorityYear="2014" baseAuthorityName="Ng & Grinang" baseAuthorityYear="2004" box="[927,1183,690,713]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="cristicervix">
|
||
<emphasis id="B9731B0F7020FFBCFC5AFD7AC1F34EB4" box="[927,1183,690,713]" italics="true" pageId="5" pageNumber="394">Thelphusula cristicervix</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C07BC9E7020FFBCFB0AFD7AC0774EB4" authority="Grinang & Ng, 2014" authorityName="Grinang & Ng" authorityYear="2014" box="[1231,1307,690,713]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="pueh" status="sp. nov.">
|
||
<emphasis id="B9731B0F7020FFBCFB0AFD7AC0774EB4" box="[1231,1307,690,713]" italics="true" pageId="5" pageNumber="394">T. pueh</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A240A6747020FFBCFAE2FD7AC0CB4EB4" box="[1319,1447,690,713]" pageId="5" pageNumber="394" rank="species">new species</taxonomicNameLabel>
|
||
, are the only two species of
|
||
<taxonomicName id="4C07BC9E7020FFBCFB8DFD1BC1A04E97" authorityName="Bott" authorityYear="1969" box="[1096,1228,723,746]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9731B0F7020FFBCFB8DFD1BC1A04E97" box="[1096,1228,723,746]" italics="true" pageId="5" pageNumber="394">Thelphusula</emphasis>
|
||
</taxonomicName>
|
||
known from western
|
||
<collectingCountry id="F310878D7020FFBCFCE8FD3DC6E04F71" box="[813,908,757,780]" name="Malaysia" pageId="5" pageNumber="394">Sarawak</collectingCountry>
|
||
. They are easily differentiated from congeners in having a relatively more ovate carapace, a more convex anterolateral margin and more strongly deflexed frontal margin. Their G1s are also superficially similar in shape.
|
||
<taxonomicName id="4C07BC9E7020FFBCFCE8FCB2C69A4FEC" authorityName="Grinang & Ng" authorityYear="2014" box="[813,1014,890,913]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="pueh">
|
||
<emphasis id="B9731B0F7020FFBCFCE8FCB2C69A4FEC" box="[813,1014,890,913]" italics="true" pageId="5" pageNumber="394">Thelphusula pueh</emphasis>
|
||
</taxonomicName>
|
||
can most easily be separated from
|
||
<taxonomicName id="4C07BC9E7020FFBCFA52FCB2C6C94FCF" authorityName="Grinang & Ng" authorityYear="2014" baseAuthorityName="Ng & Grinang" baseAuthorityYear="2004" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="cristicervix">
|
||
<emphasis id="B9731B0F7020FFBCFA52FCB2C6C94FCF" italics="true" pageId="5" pageNumber="394">T. cristicervix</emphasis>
|
||
</taxonomicName>
|
||
by its live colours; with the carapace and legs of
|
||
<taxonomicName id="4C07BC9E7020FFBCFC8FFC75C6B14FA9" authorityName="Grinang & Ng" authorityYear="2014" baseAuthorityName="Ng & Grinang" baseAuthorityYear="2004" box="[842,989,957,980]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="cristicervix">
|
||
<emphasis id="B9731B0F7020FFBCFC8FFC75C6B14FA9" box="[842,989,957,980]" italics="true" pageId="5" pageNumber="394">T. cristicervix</emphasis>
|
||
</taxonomicName>
|
||
red (
|
||
<figureCitation id="133CDB987020FFBCFBD0FC75C1E64FA9" box="[1045,1162,957,980]" captionStart="Fig" captionStartId="5.[113,148,1245,1266]" captionTargetBox="[115,752,520,1221]" captionTargetId="figure-13@5.[113,752,518,1221]" captionTargetPageId="5" captionText="Fig. 4. Colours in life, Thelphusula cristicervix (Ng & Grinang, 2004). A, male (ca. 19.0 mm carapace width), in situ in forest, specimen not collected; B, female (19.3 × 14.2 mm) (ZRC) [Photograph by: Tan HH]. Both specimens from Sama Jaya Nature Reserve, Kuching Division, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5353788" httpUri="https://zenodo.org/record/5353788/files/figure.png" pageId="5" pageNumber="394">Fig. 4A, B</figureCitation>
|
||
), even in small specimens, whereas
|
||
<taxonomicName id="4C07BC9E7020FFBCFC4BFC16C68C4F88" authorityName="Grinang & Ng" authorityYear="2014" box="[910,992,990,1013]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="pueh">
|
||
<emphasis id="B9731B0F7020FFBCFC4BFC16C68C4F88" box="[910,992,990,1013]" italics="true" pageId="5" pageNumber="394">T. pueh</emphasis>
|
||
</taxonomicName>
|
||
is dark brown (
|
||
<figureCitation id="133CDB987020FFBCFB50FC16C07C4F88" box="[1173,1296,990,1013]" captionStart="Fig" captionStartId="5.[113,148,2006,2027]" captionTargetBox="[115,753,1453,1949]" captionTargetId="figure-96@5.[114,756,1450,1982]" captionTargetPageId="5" captionText="Fig. 5. Colours in life, Thelphusula pueh, new species. A, paratype female (16.3 × 12.9 mm) (SBC); B, paratype female (13.0 × 10.1 mm) (SBC). Both specimens from Kampung Sebako, Pueh, Lundu District, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5353790" httpUri="https://zenodo.org/record/5353790/files/figure.png" pageId="5" pageNumber="394">Fig. 5A, B</figureCitation>
|
||
).
|
||
<taxonomicName id="4C07BC9E7020FFBCFAE2FC16C60D486B" authorityName="Grinang & Ng" authorityYear="2014" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="pueh">
|
||
<emphasis id="B9731B0F7020FFBCFAE2FC16C60D486B" italics="true" pageId="5" pageNumber="394">Thelphusula pueh</emphasis>
|
||
</taxonomicName>
|
||
also has proportionately shorter ambulatory legs (
|
||
<figureCitation id="133CDB987020FFBCFA45FC37C6234845" captionStart="Fig" captionStartId="3.[113,148,2034,2055]" captionTargetBox="[120,1445,189,2010]" captionTargetId="figure-73@3.[120,1445,189,2010]" captionTargetPageId="3" captionText="Fig. 2. Thelphusula pueh, new species. Holotype male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353784" httpUri="https://zenodo.org/record/5353784/files/figure.png" pageId="5" pageNumber="394">Fig. 2A</figureCitation>
|
||
) (versus longer in
|
||
<taxonomicName id="4C07BC9E7020FFBCFBECFBE9C1AE4845" authorityName="Grinang & Ng" authorityYear="2014" baseAuthorityName="Ng & Grinang" baseAuthorityYear="2004" box="[1065,1218,1057,1080]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="cristicervix">
|
||
<emphasis id="B9731B0F7020FFBCFBECFBE9C1AE4845" box="[1065,1218,1057,1080]" italics="true" pageId="5" pageNumber="394">T. cristicervix</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="133CDB987020FFBCFB16FBE9C0404845" box="[1235,1324,1057,1080]" captionStart="Fig" captionStartId="2.[136,171,2034,2055]" captionTargetBox="[142,1467,188,2010]" captionTargetId="figure-69@2.[142,1467,188,2010]" captionTargetPageId="2" captionText="Fig. 1. Thelphusula cristicervix (Ng & Grinang, 2004). Male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353782" httpUri="https://zenodo.org/record/5353782/files/figure.png" pageId="5" pageNumber="394">Fig. 1A</figureCitation>
|
||
); relatively shallower cervical grooves (
|
||
<figureCitation id="133CDB987020FFBCFBB2FB8AC1BD4824" box="[1143,1233,1090,1113]" captionStart="Fig" captionStartId="3.[113,148,2034,2055]" captionTargetBox="[120,1445,189,2010]" captionTargetId="figure-73@3.[120,1445,189,2010]" captionTargetPageId="3" captionText="Fig. 2. Thelphusula pueh, new species. Holotype male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353784" httpUri="https://zenodo.org/record/5353784/files/figure.png" pageId="5" pageNumber="394">Fig. 2B</figureCitation>
|
||
) (versus distinctly deeper in
|
||
<taxonomicName id="4C07BC9E7020FFBCFC59FBABC15E4807" authorityName="Grinang & Ng" authorityYear="2014" baseAuthorityName="Ng & Grinang" baseAuthorityYear="2004" box="[924,1074,1123,1146]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="cristicervix">
|
||
<emphasis id="B9731B0F7020FFBCFC59FBABC15E4807" box="[924,1074,1123,1146]" italics="true" pageId="5" pageNumber="394">T. cristicervix</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="133CDB987020FFBCFB87FBABC1F54807" box="[1090,1177,1123,1146]" captionStart="Fig" captionStartId="2.[136,171,2034,2055]" captionTargetBox="[142,1467,188,2010]" captionTargetId="figure-69@2.[142,1467,188,2010]" captionTargetPageId="2" captionText="Fig. 1. Thelphusula cristicervix (Ng & Grinang, 2004). Male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353782" httpUri="https://zenodo.org/record/5353782/files/figure.png" pageId="5" pageNumber="394">Fig. 1A</figureCitation>
|
||
); smooth lateral margins of abdomen (
|
||
<figureCitation id="133CDB987020FFBCFC7EFB4DC16248E1" box="[955,1038,1157,1180]" captionStart="Fig" captionStartId="3.[113,148,2034,2055]" captionTargetBox="[120,1445,189,2010]" captionTargetId="figure-73@3.[120,1445,189,2010]" captionTargetPageId="3" captionText="Fig. 2. Thelphusula pueh, new species. Holotype male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353784" httpUri="https://zenodo.org/record/5353784/files/figure.png" pageId="5" pageNumber="394">Fig. 2D</figureCitation>
|
||
) (versus uneven in
|
||
<taxonomicName id="4C07BC9E7020FFBCFB24FB4DC01848E1" authorityName="Grinang & Ng" authorityYear="2014" baseAuthorityName="Ng & Grinang" baseAuthorityYear="2004" box="[1249,1396,1157,1180]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="cristicervix">
|
||
<emphasis id="B9731B0F7020FFBCFB24FB4DC01848E1" box="[1249,1396,1157,1180]" italics="true" pageId="5" pageNumber="394">T. cristicervix</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="133CDB987020FFBCFA44FB4DC62348C0" captionStart="Fig" captionStartId="2.[136,171,2034,2055]" captionTargetBox="[142,1467,188,2010]" captionTargetId="figure-69@2.[142,1467,188,2010]" captionTargetPageId="2" captionText="Fig. 1. Thelphusula cristicervix (Ng & Grinang, 2004). Male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353782" httpUri="https://zenodo.org/record/5353782/files/figure.png" pageId="5" pageNumber="394">Fig. 1D</figureCitation>
|
||
); and the outer margin of the G1 subterminal segment is almost straight with the terminal segment more strongly bent (
|
||
<figureCitation id="133CDB987020FFBCFCA8FB21C68E497D" box="[877,994,1257,1280]" captionStart="Fig" captionStartId="4.[136,171,2004,2025]" captionTargetBox="[136,1485,266,1950]" captionTargetId="figure-7@4.[108,1500,238,1953]" captionTargetPageId="4" captionText="Fig. 3. A–E, Thelphusula cristicervix (Ng & Grinang, 2004), male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak; F–J, Thelphusula pueh, new species, male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, F, ventral view of left G1; B, G, ventral view of left G2; C, H, dorsal view of left G1; D, I, ventral view of terminal segment of left G1; E, J, dorsal view of terminal segment of left G1. Scale bars: A–C, F–H = 1.0 mm; D, E, I, J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5353786" httpUri="https://zenodo.org/record/5353786/files/figure.png" pageId="5" pageNumber="394">Fig. 3F, H</figureCitation>
|
||
) (versus outer margin of G1 subterminal segment gradually curved with the terminal segment gently bent in
|
||
<taxonomicName id="4C07BC9E7020FFBCFC44FAE3C179493F" authorityName="Grinang & Ng" authorityYear="2014" baseAuthorityName="Ng & Grinang" baseAuthorityYear="2004" box="[897,1045,1323,1346]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="cristicervix">
|
||
<emphasis id="B9731B0F7020FFBCFC44FAE3C179493F" box="[897,1045,1323,1346]" italics="true" pageId="5" pageNumber="394">T. cristicervix</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="133CDB987020FFBCFBE1FAE3C1F0493F" box="[1060,1180,1323,1346]" captionStart="Fig" captionStartId="4.[136,171,2004,2025]" captionTargetBox="[136,1485,266,1950]" captionTargetId="figure-7@4.[108,1500,238,1953]" captionTargetPageId="4" captionText="Fig. 3. A–E, Thelphusula cristicervix (Ng & Grinang, 2004), male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak; F–J, Thelphusula pueh, new species, male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, F, ventral view of left G1; B, G, ventral view of left G2; C, H, dorsal view of left G1; D, I, ventral view of terminal segment of left G1; E, J, dorsal view of terminal segment of left G1. Scale bars: A–C, F–H = 1.0 mm; D, E, I, J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5353786" httpUri="https://zenodo.org/record/5353786/files/figure.png" pageId="5" pageNumber="394">Fig. 3A, C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF7897957020FFBCFFB4FB15C48D4918" ID-DOI="http://doi.org/10.5281/zenodo.5353788" ID-Zenodo-Dep="5353788" httpUri="https://zenodo.org/record/5353788/files/figure.png" pageId="5" pageNumber="394" startId="5.[113,148,1245,1266]" targetBox="[115,752,520,1221]" targetPageId="5">
|
||
<paragraph id="8BB8C71D7020FFBCFFB4FB15C48D4918" blockId="5.[113,752,1245,1381]" pageId="5" pageNumber="394">
|
||
Fig. 4. Colours in life,
|
||
<taxonomicName id="4C07BC9E7020FFBCFE9EFB15C5C14973" authority="(Ng & Grinang, 2004)" authorityName="Grinang & Ng" authorityYear="2014" baseAuthorityName="Ng & Grinang" baseAuthorityYear="2004" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="cristicervix">
|
||
<emphasis id="B9731B0F7020FFBCFE9EFB15C72B488F" box="[347,583,1245,1266]" italics="true" pageId="5" pageNumber="394">Thelphusula cristicervix</emphasis>
|
||
(
|
||
<bibRefCitation id="EF96BAEC7020FFBCFD92FB15C5CA4973" author="Ng PKL & Grinang J" pageId="5" pageNumber="394" pagination="299 - 325" refId="ref4094" refString="Ng PKL & Grinang J (2004) Decapod crustaceans with descriptions of three new species. In: Yong HS Ng FSP & Yen EEL (eds.) Sarawak Bau Limestone Biodiversity. Sarawak Museum Journal, 59, No. 80 (New Series), Special Issue No. 6: 299 - 325." type="book chapter" year="2004">Ng & Grinang, 2004</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. A, male (ca. 19.0 mm carapace width), in situ in forest, specimen not collected; B, female (19.3 × 14.2 mm) (ZRC) [Photograph by: Tan HH]. Both specimens from Sama Jaya Nature Reserve, Kuching Division, Sarawak.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB8C71D7020FFBCFCE8FAA6C1A349B5" blockId="5.[813,1452,1390,1480]" pageId="5" pageNumber="394">
|
||
Like
|
||
<taxonomicName id="4C07BC9E7020FFBCFCA0FAA6C69449F8" authorityName="Grinang & Ng" authorityYear="2014" baseAuthorityName="Ng & Grinang" baseAuthorityYear="2004" box="[869,1016,1390,1413]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="cristicervix">
|
||
<emphasis id="B9731B0F7020FFBCFCA0FAA6C69449F8" box="[869,1016,1390,1413]" italics="true" pageId="5" pageNumber="394">T. cristicervix</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C07BC9E7020FFBCFBC3FAA6C13849F8" authorityName="Grinang & Ng" authorityYear="2014" box="[1030,1108,1390,1413]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="pueh">
|
||
<emphasis id="B9731B0F7020FFBCFBC3FAA6C13849F8" box="[1030,1108,1390,1413]" italics="true" pageId="5" pageNumber="394">T. pueh</emphasis>
|
||
</taxonomicName>
|
||
has a preference for damp soils and constructs shallow burrows. Its presence in a more open habitat (ricefields) may not be natural.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |