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<document id="C79121E3576D660612892566714F83CD" ID="10.11646/zootaxa.4270.1.1SLASH11140" ID-CLB-Dataset="32975" ID-DOI="10.5281/zenodo.583183" ID-GBIF-Dataset="6c479acc-8b18-4f0b-a6e5-85bcd6d7b6b7" ID-ISSN="1175-5326" ID-Zenodo-Dep="583183" ID-ZooBank="2D00AFF5-4FE2-4EC1-A328-C8670CFB8D6D" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1495696370209" checkinUser="plazi" docAuthor="Jong, Rienk De" docDate="2017" docId="03AA87D32860FFF7F7F0FC7FFB82B64D" docLanguage="en" docName="zootaxa.4270.1.1SLASH11140.pdf" docOrigin="Zootaxa 4270 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Riodinella nympha" docType="treatment" docVersion="9" lastPageNumber="33" masterDocId="FF93FFAB2840FFD7F767FF87FFA3B03E" masterDocTitle="Fossil butterflies, calibration points and the molecular clock (Lepidoptera: Papilionoidea)" masterLastPageNumber="63" masterPageNumber="1" pageNumber="33" updateTime="1698439901205" updateUser="ExternalLinkService">
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<mods:title id="44AAB6D991291E98F505986E800FE078">Fossil butterflies, calibration points and the molecular clock (Lepidoptera: Papilionoidea)</mods:title>
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<mods:namePart id="D65DCDFE1C2DCDB5FC2896A28719D1A3">Jong, Rienk De</mods:namePart>
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<mods:date id="51C07322774CFBCEFB7B726D00266A4A">2017</mods:date>
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<mods:classification id="84296A055D7B4CDE797202A1E00CF032">journal article</mods:classification>
<mods:identifier id="7CBB2808C5A8C47ED384F9B06FD734A6" type="CLB-Dataset">32975</mods:identifier>
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<treatment id="03AA87D32860FFF7F7F0FC7FFB82B64D" ID-DOI="http://doi.org/10.5281/zenodo.6047021" ID-GBIF-Taxon="130844863" ID-Zenodo-Dep="6047021" LSID="urn:lsid:plazi:treatment:03AA87D32860FFF7F7F0FC7FFB82B64D" httpUri="http://treatment.plazi.org/id/03AA87D32860FFF7F7F0FC7FFB82B64D" lastPageNumber="33" pageId="32" pageNumber="33">
<subSubSection id="C319654E2860FFF7F7F0FC7FFE9BB469" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="8BBC36C52860FFF7F7F0FC7FFCA1B42C" blockId="32.[151,770,1016,1042]" box="[151,770,1016,1042]" pageId="32" pageNumber="33">
<heading id="D0F481A92860FFF7F7F0FC7FFCA1B42C" bold="true" box="[151,770,1016,1042]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
<emphasis id="B977EAD72860FFF7F7F0FC7FFCA1B42C" bold="true" box="[151,770,1016,1042]" pageId="32" pageNumber="33">
<emphasis id="B977EAD72860FFF7F7F0FC7FFEA3B42C" bold="true" box="[151,256,1016,1042]" italics="true" pageId="32" pageNumber="33">
<taxonomicName id="4C034D462860FFF7F7F0FC7FFF58B42C" ID-CoL="4T4KH" box="[151,251,1016,1042]" class="Insecta" family="Riodinidae" genus="Riodinella" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="nympha">nympha</taxonomicName>
.
</emphasis>
<taxonomicName id="4C034D462860FFF7F660FC7FFCA1B42C" authority="Durden &amp; Rose, 1978" authorityName="Durden &amp; Rose" authorityYear="1978" box="[263,770,1016,1042]" class="Insecta" family="Riodinidae" genus="Riodinella" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="nympha">
<emphasis id="B977EAD72860FFF7F660FC7FFE4DB42C" bold="true" box="[263,494,1016,1042]" italics="true" pageId="32" pageNumber="33">Riodinella nympha</emphasis>
<bibRefCitation id="EF924B342860FFF7F692FC7FFCA1B42C" author="Durden" box="[501,770,1016,1042]" pageId="32" pageNumber="33" refString="Durden, C. J. &amp; Rose, H. (1978) Butterflies from the Middle Miocene: the earliest occurrence of fossil Papilionoidea (Lepidoptera). Pearce-Sellards Series, 29, 1 - 25." type="journal article" year="1978">Durden &amp; Rose, 1978</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBC36C52860FFF7F7F0FBB8FE9BB469" blockId="32.[151,1437,1086,1651]" box="[151,312,1086,1111]" pageId="32" pageNumber="33">Incertae sedis.</paragraph>
</subSubSection>
<subSubSection id="C319654E2860FFF7F7A0FBE4FB82B64D" pageId="32" pageNumber="33" type="description">
<paragraph id="8BBC36C52860FFF7F7A0FBE4FBBCB444" blockId="32.[151,1437,1086,1651]" box="[199,1055,1122,1147]" pageId="32" pageNumber="33">
<materialsCitation id="3B6B3C982860FFF7F7A0FBE4FBBCB444" ID-GBIF-Occurrence="1503739260" box="[199,1055,1122,1147]" country="United States of America" county="Rio Blanco Co." location="Rio Blanco Co." pageId="32" pageNumber="33" specimenCount="1" stateProvince="Colorado">
<collectingCountry id="F31476552860FFF7F7A0FBE4FEA1B445" box="[199,258,1123,1147]" name="United States of America" pageId="32" pageNumber="33">USA</collectingCountry>
,
<taxonomicName id="4C034D462860FFF7F66AFBE5FEDBB445" box="[269,376,1122,1147]" class="Insecta" family="Papilionidae" genus="Praepapilio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="63" phylum="Arthropoda" rank="species" species="colorado">
<collectingRegion id="49C7F8272860FFF7F66AFBE5FEDBB445" box="[269,376,1122,1147]" country="United States of America" name="Colorado" pageId="32" pageNumber="33">Colorado</collectingRegion>
</taxonomicName>
,
<collectingCounty id="62DD4E492860FFF7F6E4FBE5FD90B445" box="[387,563,1122,1147]" pageId="32" pageNumber="33">Rio Blanco Co.</collectingCounty>
, Ray Dome; early Lutetian, middle Eocene.
</materialsCitation>
</paragraph>
<paragraph id="8BBC36C52860FFF7F7A0FB01FC1DB4A1" blockId="32.[151,1437,1086,1651]" box="[199,958,1158,1183]" pageId="32" pageNumber="33">
Depositories: PLME (
<typeStatus id="54B888672860FFF7F6DAFB01FD80B4A1" box="[445,547,1158,1183]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
, no. 3a), BHM (counterpart, no. 3b).
</paragraph>
<paragraph id="8BBC36C52860FFF7F7A0FB2DFBDEB4FD" blockId="32.[151,1437,1086,1651]" box="[199,1149,1194,1219]" pageId="32" pageNumber="33">
Published figures:
<bibRefCitation id="EF924B342860FFF7F6FEFB2DFCFBB4FD" author="Durden" box="[409,856,1194,1219]" pageId="32" pageNumber="33" refString="Durden, C. J. &amp; Rose, H. (1978) Butterflies from the Middle Miocene: the earliest occurrence of fossil Papilionoidea (Lepidoptera). Pearce-Sellards Series, 29, 1 - 25." type="journal article" year="1978">
Durden &amp; Rose (1978:
<figureCitation id="13382A402860FFF7F5C7FB2DFD44B4FD" box="[672,743,1194,1219]" captionStart="FIGURES 3 – 4" captionStartId="10.[151,264,1801,1823]" captionTargetBox="[165,1429,840,1774]" captionTargetId="figure@10.[154,1438,828,1788]" captionTargetPageId="10" captionText="FIGURES 3 – 4. Wing venation of (A) Pareronia Bingham, 1907 (Pieridae), and (B) Hasora Moore, 1881 (Hesperiidae). See text for explanation." httpUri="https://zenodo.org/record/583186/files/figure.png" pageId="32" pageNumber="33">Figs 4</figureCitation>
,
<figureCitation id="13382A402860FFF7F594FB2DFCA1B4FD" box="[755,770,1194,1219]" captionStart="FIGURE 5" captionStartId="16.[151,250,1784,1806]" captionTargetBox="[238,1332,193,1743]" captionTargetId="figure@16.[238,1332,193,1754]" captionTargetPageId="16" captionText="FIGURE 5. Ƒanessa amerindica Miller &amp; Brown, 1989, holotype, collections of Invertebrate Paleontology, University of Florida. Photo by Roger Portell. Fig. 6. Pseudoneorina coulleti Nel &amp; Descimon, 1984. France, Dép. Alpes-de-Haute- Provence, Dauphin, specimen in coll. M. Henrotay; photo by courtesy of M. Henrotay. Fig. 7. Archaeolycorea ferreirai Martins-Neto, 1989. Fig. 4 A of original description. Fig. 8. Oligodonta florissantensis Brown, 1976, holotype, collections of Invertebrate Paleontology, University of Florida. Photo by courtesy of Akito Kawahara. Fig. 9. Neorinella garciae Martins- Neto, Kucera-Santos, de Moraes Vieira &amp; de Campos Fragoso, 1993. Fig. 3 A of original description. Fig. 10. Protocoeliades kristenseni de Jong, 2016, holotype, Zoologisk Museum, Copenhagen, Denmark. Photo by Department of Paleobiology, National Museum of Natural History, Washington DC, USA." httpUri="https://zenodo.org/record/583187/files/figure.png" pageId="32" pageNumber="33">5</figureCitation>
, 6E–F)
</bibRefCitation>
;
<bibRefCitation id="EF924B342860FFF7F403FB2CFBDAB4FD" author="Murata" box="[868,1145,1194,1219]" pageId="32" pageNumber="33" refString="Murata, Y. (1998) Notes on the evolutionary process of butterflies and butterfly fossils. Butterflies, 20 (part 1 &amp; 2), 4 - 17 &amp; 27 - 40. [in Japanese with English summary]" type="journal article" year="1998">
Murata (1998:
<figureCitation id="13382A402860FFF7F36BFB2DFBF0B4FD" box="[1036,1107,1194,1219]" captionStart="FIGURES 3 – 4" captionStartId="10.[151,264,1801,1823]" captionTargetBox="[165,1429,840,1774]" captionTargetId="figure@10.[154,1438,828,1788]" captionTargetPageId="10" captionText="FIGURES 3 – 4. Wing venation of (A) Pareronia Bingham, 1907 (Pieridae), and (B) Hasora Moore, 1881 (Hesperiidae). See text for explanation." httpUri="https://zenodo.org/record/583186/files/figure.png" pageId="32" pageNumber="33">Figs 4</figureCitation>
,
<figureCitation id="13382A402860FFF7F338FB2DFBCCB4FD" box="[1119,1135,1194,1219]" captionStart="FIGURE 5" captionStartId="16.[151,250,1784,1806]" captionTargetBox="[238,1332,193,1743]" captionTargetId="figure@16.[238,1332,193,1754]" captionTargetPageId="16" captionText="FIGURE 5. Ƒanessa amerindica Miller &amp; Brown, 1989, holotype, collections of Invertebrate Paleontology, University of Florida. Photo by Roger Portell. Fig. 6. Pseudoneorina coulleti Nel &amp; Descimon, 1984. France, Dép. Alpes-de-Haute- Provence, Dauphin, specimen in coll. M. Henrotay; photo by courtesy of M. Henrotay. Fig. 7. Archaeolycorea ferreirai Martins-Neto, 1989. Fig. 4 A of original description. Fig. 8. Oligodonta florissantensis Brown, 1976, holotype, collections of Invertebrate Paleontology, University of Florida. Photo by courtesy of Akito Kawahara. Fig. 9. Neorinella garciae Martins- Neto, Kucera-Santos, de Moraes Vieira &amp; de Campos Fragoso, 1993. Fig. 3 A of original description. Fig. 10. Protocoeliades kristenseni de Jong, 2016, holotype, Zoologisk Museum, Copenhagen, Denmark. Photo by Department of Paleobiology, National Museum of Natural History, Washington DC, USA." httpUri="https://zenodo.org/record/583187/files/figure.png" pageId="32" pageNumber="33">5</figureCitation>
)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BBC36C52860FFF7F7A0FB49FD11B581" blockId="32.[151,1437,1086,1651]" pageId="32" pageNumber="33">
Described as belonging to
<taxonomicName id="4C034D462860FFF7F691FB49FDD6B4D9" box="[502,629,1230,1255]" class="Insecta" family="Lycaenidae" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="family">Lycaenidae</taxonomicName>
(at the time including
<taxonomicName id="4C034D462860FFF7F41EFB49FC55B4D9" box="[889,1014,1230,1255]" class="Insecta" family="Riodinidae" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="family">Riodinidae</taxonomicName>
) and compared, among others, to the extant genus
<taxonomicName id="4C034D462860FFF7F64CFB75FE27B535" box="[299,388,1266,1291]" class="Insecta" family="Riodinidae" genus="Riodina" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B977EAD72860FFF7F64CFB75FE27B535" box="[299,388,1266,1291]" italics="true" pageId="32" pageNumber="33">Riodina</emphasis>
</taxonomicName>
. In addition to photos of the
<typeStatus id="54B888672860FFF7F5AEFB75FC8DB535" box="[713,814,1266,1291]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
, a reconstruction of the wing venation and an extensive description is given. In the reconstruction of the hindwing a long tail at the end of M3 and CuA1 has been reproduced, or rather produced, since it is not present in the fossil.
<bibRefCitation id="EF924B342860FFF7F4F9FABDFBC5B56D" box="[926,1126,1338,1363]" pageId="32" pageNumber="33" refString="Hall, J. P. W., Robbins, R. K. &amp; Harvey, D. J. (2004) Extinction and biogeography in the Caribbean: new evidence from a fossil riodinid butterfly in Dominican amber. Proceedings of the Royal Society of London B, 271, 797 - 801. https: // doi. org / 10.1098 / rspb. 2004.2691" type="journal article">
Hall
<emphasis id="B977EAD72860FFF7F4B0FABBFBA8B56D" box="[983,1035,1338,1363]" italics="true" pageId="32" pageNumber="33">et al</emphasis>
. (2004)
</bibRefCitation>
extensively explain that in wing venation there is nothing to link the fossil exclusively to the
<taxonomicName id="4C034D462860FFF7F4E3FAD9FBA0B549" box="[900,1027,1374,1399]" class="Insecta" family="Riodinidae" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="family">Riodinidae</taxonomicName>
; several of the venational characters also are found in
<taxonomicName id="4C034D462860FFF7F630FA05FE45B5A5" box="[343,486,1410,1435]" class="Insecta" family="Papilionidae" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="family">Papilionidae</taxonomicName>
,
<taxonomicName id="4C034D462860FFF7F696FA05FDEEB5A5" box="[497,589,1410,1435]" class="Insecta" family="Pieridae" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="family">Pieridae</taxonomicName>
and
<taxonomicName id="4C034D462860FFF7F5E5FA05FCBEB5A5" box="[642,797,1410,1435]" class="Insecta" family="Nymphalidae" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="family">Nymphalidae</taxonomicName>
.
<bibRefCitation id="EF924B342860FFF7F440FA05FC45B5A5" box="[807,998,1410,1435]" pageId="32" pageNumber="33" refString="Hall, J. P. W., Robbins, R. K. &amp; Harvey, D. J. (2004) Extinction and biogeography in the Caribbean: new evidence from a fossil riodinid butterfly in Dominican amber. Proceedings of the Royal Society of London B, 271, 797 - 801. https: // doi. org / 10.1098 / rspb. 2004.2691" type="journal article">
Hall
<emphasis id="B977EAD72860FFF7F43AFA03FC2DB5A5" box="[861,910,1410,1435]" italics="true" pageId="32" pageNumber="33">et al</emphasis>
. (2004)
</bibRefCitation>
conclude that it should be treated as an unplaced butterfly, with which I can fully agree.
</paragraph>
<paragraph id="8BBC36C52860FFF7F7A0FA4DFB82B64D" blockId="32.[151,1437,1086,1651]" pageId="32" pageNumber="33">
According to the original description “the male genitalia are definitely comparable to modern members of this subtribe”, but the genitalia are not described except for a structure &quot;in the anterior third of the capsule&quot;. This structure is interpreted as representing a Y-shaped furca, an interpretation that is considered unreliable. Apparently the specimen is supposed to be a male, but under “Characters agreeing with the family” [
<taxonomicName id="4C034D462860FFF7F3CBF9B1FA91B671" box="[1196,1330,1590,1615]" class="Insecta" family="Lycaenidae" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="family">Lycaenidae</taxonomicName>
] is listed “Prothoracic femur 0.6 of mesothoracic femur in female” (
<bibRefCitation id="EF924B342860FFF7F440F9DDFBBAB64D" author="Durden" box="[807,1049,1626,1651]" pageId="32" pageNumber="33" refString="Durden, C. J. &amp; Rose, H. (1978) Butterflies from the Middle Miocene: the earliest occurrence of fossil Papilionoidea (Lepidoptera). Pearce-Sellards Series, 29, 1 - 25." type="journal article" year="1978">Durden &amp; Rose 1978</bibRefCitation>
)
</paragraph>
</subSubSection>
</treatment>
</document>
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