287 lines
40 KiB
XML
287 lines
40 KiB
XML
<document id="BF7139DC20E62F8F8CB92F5E914BBA8B" ID-CLB-Dataset="267614" ID-DOI="10.1163/18759866-BJA10027" ID-GBIF-Dataset="2fd2ff98-fcd5-484e-a7b8-6a60f12e7084" ID-ISSN="1875-9866" ID-Zenodo-Dep="8348969" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1694665046491" checkinUser="felipe" docAuthor="Pholyotha, Arthit, Sutcharit, Chirasak, Lin, Aung & Panha, Somsak" docDate="2022" docId="038587FD036EFFCE76BCFA439789FB3F" docLanguage="en" docName="ContribZool.91.2.97-132.pdf" docOrigin="Contributions to Zoology 91 (2)" docSource="http://dx.doi.org/10.1163/18759866-bja10027" docStyle="DocumentStyle:E1C3C5FD6CDDA08D62BFE928526826AF.2:ContribZool.2019-.journal_article" docStyleId="E1C3C5FD6CDDA08D62BFE928526826AF" docStyleName="ContribZool.2019-.journal_article" docStyleVersion="2" docTitle="Sarika costata Pholyotha & Sutcharit & Lin & Panha 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="128" masterDocId="FFBCFF850373FFD17439FF98972DFFE8" masterDocTitle="Multigene phylogeny reveals the ribbed shell morphotypes in the land snail genus Sarika (Eupulmonata: Ariophantidae), with description of two new species from Thailand and Myanmar" masterLastPageNumber="132" masterPageNumber="97" pageNumber="126" updateTime="1694783892711" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
|
||
<mods:mods id="DFC4EB5390E5967D7D20C3897733FB1B" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="DE6DDB3F60538F20ADB82CFA2B12AC1E">
|
||
<mods:title id="2993284A8A125F774F8BA4EA00B6D7DB">Multigene phylogeny reveals the ribbed shell morphotypes in the land snail genus Sarika (Eupulmonata: Ariophantidae), with description of two new species from Thailand and Myanmar</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="FEC6506B89314221BD33AC3E8F8D10AB" type="personal">
|
||
<mods:role id="51EB6D0379B3E57A4C8662ACB67B2BC9">
|
||
<mods:roleTerm id="C652A866225C218B05356CDBD7391F01">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="23073C2E5A7541D2D59AF682B936E504">Pholyotha, Arthit</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="EDDCFD2CEAC2172C05382701B454572A" type="personal">
|
||
<mods:role id="22B796E2FD13856E52AF0FECBC1D1EF3">
|
||
<mods:roleTerm id="4F0A1DBD6A2367726129B678A42A3C88">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B3A08C9BB9BE73A7D77E59E07EEF2304">Sutcharit, Chirasak</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="711E173E7555426AD9918210481E1668" type="personal">
|
||
<mods:role id="2DCEF8C1817FB6B652ADB34324494A54">
|
||
<mods:roleTerm id="15A98A308F0ED0F1DDDEDC605F853C0B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8FB3FACC58AA62E2F78DB734FD36E552">Lin, Aung</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="8613F8B64E33331B89D85AE31B36B122" type="personal">
|
||
<mods:role id="A887F5F28346DD5FD37C322DA30C8E57">
|
||
<mods:roleTerm id="A3127AFB32A924CD1F32605DC3E717BB">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="DD0F2575EC867BE3BFEA60FA78868C2A">Panha, Somsak</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="F701869B91AA3D8D4C41579DEC4C3CA3">text</mods:typeOfResource>
|
||
<mods:relatedItem id="E7C6127B923D71EE2207167EEB242983" type="host">
|
||
<mods:titleInfo id="4C18B62D829E0082D0DB5B90A7F7C343">
|
||
<mods:title id="3569787FFD5CE0C8290B25530DD12679">Contributions to Zoology</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="8790BDDDF57A31A9525449A58002B5E7">
|
||
<mods:date id="BEE5655610711F2E7C2CAA3A65CCA8E7">2022</mods:date>
|
||
<mods:detail id="CAA1FA185BA225466FA67B8B1E77FF7D" type="pubDate">
|
||
<mods:number id="81B4A1C2B52A8DB88933145F02D450C9">2022-03-21</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="1A6C45528C4BFE7966B5A53FE98213CA" type="volume">
|
||
<mods:number id="F78FAFA631DC8CF4E4B708743B965584">91</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="0FC701D4CD6A1219DBBE946C075E4B11" type="issue">
|
||
<mods:number id="2E8DEABD30106A2E6CFA7B9AA767D12B">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="4FE884AE2A1DE2B593E4CE28840FB62E" unit="page">
|
||
<mods:start id="62A698F5BD1257F1206CA3252E369EB3">97</mods:start>
|
||
<mods:end id="CEFB4EB17F9D6ACE33E747F691DE7027">132</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="3996F752CD85C73DD8CF963047D0701D">
|
||
<mods:url id="1D9BC080E9B8AA4A53FDB92F1D96B22B">http://dx.doi.org/10.1163/18759866-bja10027</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="0225906C0CF8B6725C04BE07C35A0193">journal article</mods:classification>
|
||
<mods:identifier id="422836B877C5AC1179C841AB2F832F0F" type="CLB-Dataset">267614</mods:identifier>
|
||
<mods:identifier id="97D9697E0287CAE47377725FB0178E79" type="DOI">10.1163/18759866-BJA10027</mods:identifier>
|
||
<mods:identifier id="DF09BB28532478E8C97E62CFD23F7B32" type="GBIF-Dataset">2fd2ff98-fcd5-484e-a7b8-6a60f12e7084</mods:identifier>
|
||
<mods:identifier id="CC01D30CD3744A34FB315E6D4076ED72" type="ISSN">1875-9866</mods:identifier>
|
||
<mods:identifier id="A6D1B81FC040AC787F2687E5783A85E7" type="Zenodo-Dep">8348969</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="038587FD036EFFCE76BCFA439789FB3F" ID-DOI="http://doi.org/10.5281/zenodo.8348989" ID-Zenodo-Dep="8348989" LSID="urn:lsid:plazi:treatment:038587FD036EFFCE76BCFA439789FB3F" httpUri="http://treatment.plazi.org/id/038587FD036EFFCE76BCFA439789FB3F" lastPageId="31" lastPageNumber="128" pageId="29" pageNumber="126">
|
||
<subSubSection id="C3366560036EFFCC76BCFA439329FA1E" box="[645,1028,1499,1526]" pageId="29" pageNumber="126" type="nomenclature">
|
||
<paragraph id="8B9336EB036EFFCC76BCFA439329FA1E" blockId="29.[645,1028,1499,1526]" box="[645,1028,1499,1526]" pageId="29" pageNumber="126">
|
||
<heading id="D0DB8187036EFFCC76BCFA439329FA1E" box="[645,1028,1499,1526]" fontSize="11" level="2" pageId="29" pageNumber="126" reason="2">
|
||
<emphasis id="B958EAF9036EFFCC76BCFA439329FA1E" bold="true" box="[645,1028,1499,1526]" pageId="29" pageNumber="126">
|
||
<taxonomicName id="4C2C4D68036EFFCC76BCFA439488FA1E" ID-CoL="8YNYN" authority="Pholyotha" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" box="[645,933,1499,1526]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="126" phylum="Mollusca" rank="species" species="costata" status="sp. nov.">
|
||
<emphasis id="B958EAF9036EFFCC76BCFA439405FA1E" bold="true" box="[645,808,1499,1526]" italics="true" pageId="29" pageNumber="126">Sarika costata</emphasis>
|
||
Pholyotha
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26B5782036EFFCC7789FA449329FA1E" box="[944,1028,1500,1526]" pageId="29" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3366560036EFFCE76BCF9989789FB3F" lastPageId="31" lastPageNumber="128" pageId="29" pageNumber="126" type="description">
|
||
<paragraph id="8B9336EB036EFFCC76BCF9989489F9F2" blockId="29.[645,932,1536,1562]" box="[645,932,1536,1562]" pageId="29" pageNumber="126">
|
||
(
|
||
<figureCitation id="13172A6E036EFFCC76B4F99895E8F9F2" box="[653,709,1536,1562]" captionStart="FIGURE 1" captionStartId="3.[99,166,977,998]" captionTargetBox="[193,1062,164,958]" captionTargetId="figure-233@3.[193,1062,164,958]" captionTargetPageId="3" captionText="FIGURE 1 Geographic distribution of ribbed shell Sarika spp. based on the specimens examined herein. A) Distribution of S. theodori, S.costabilis sp.nov., and S. costata sp. nov. B) Distribution of S. siamensis. Black symbols indicate the DNA extracted specimens used in the molecular analysis; white symbols indicate shells or other preserved specimens." figureDoi="http://doi.org/10.5281/zenodo.8348977" httpUri="https://zenodo.org/record/8348977/files/figure.png" pageId="29" pageNumber="126">figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="13172A6E036EFFCC76F6F99895F0F9F2" box="[719,733,1536,1562]" captionStart="FIGURE 2" captionStartId="11.[100,167,1258,1279]" captionTargetBox="[193,1061,164,1238]" captionTargetId="figure-122@11.[193,1062,164,1239]" captionTargetPageId="11" captionText="FIGURE 2 Bayesian inference tree based on a 1495 bp concatenated alignment dataset of COI + 16S + 28S genes. Numbers by the nodes are the Bayesian posterior probabilities (left) and ML bootstrap values (right); shown only for the nodes supported by BI ≥ 0.95 or ML ≥ 70%. Clade A consists of the S. resplendens-S. hainesi species groups.Clade B consists of the S.siamensis species group.Images of living snails are not to scale." figureDoi="http://doi.org/10.5281/zenodo.8348979" httpUri="https://zenodo.org/record/8348979/files/figure.png" pageId="29" pageNumber="126">2</figureCitation>
|
||
,
|
||
<figureCitation id="13172A6E036EFFCC76DEF9989409F9F2" box="[743,804,1536,1562]" captionStart="FIGURE 8" captionStartId="25.[97,164,1691,1712]" captionTargetBox="[193,1062,591,1673]" captionTargetId="figure-133@25.[193,1062,591,1673]" captionTargetPageId="25" captionText="FIGURE 8 Shells of Sarika spp.A, B) S. costabilis sp. nov.A) Holotype CUMZ 7943.B) Paratype CUMZ 7944.C, D) S. costata sp.nov.C) Holotype CUMZ 7946.D) Paratype CUMZ 7947." figureDoi="http://doi.org/10.5281/zenodo.8348999" httpUri="https://zenodo.org/record/8348999/files/figure.png" pageId="29" pageNumber="126">8C, D</figureCitation>
|
||
,
|
||
<figureCitation id="13172A6E036EFFCC7716F9989441F9F2" box="[815,876,1536,1562]" captionStart="FIGURE 9" captionStartId="27.[98,165,1350,1371]" captionTargetBox="[193,1062,164,1331]" captionTargetId="figure-97@27.[193,1062,164,1331]" captionTargetPageId="27" captionText="FIGURE 9 Genitalia of Sarika spp.A, B) S. costabilis sp. nov.paratype CUMZ 7945.A) General view of the genital system.B) Internal structure of penis.C, D) S.costata sp. nov. paratype CUMZ 7947.C) General view of the genital system.D) Internal structure of penis. White arrows indicate the end of penis." figureDoi="http://doi.org/10.5281/zenodo.8349001" httpUri="https://zenodo.org/record/8349001/files/figure.png" pageId="29" pageNumber="126">9C, D</figureCitation>
|
||
,
|
||
<figureCitation id="13172A6E036EFFCC774FF99894B0F9F2" box="[886,925,1536,1562]" captionStart="FIGURE 11" captionStartId="29.[98,165,1042,1063]" captionTargetBox="[193,1062,164,1023]" captionTargetId="figure-192@29.[193,1062,164,1023]" captionTargetPageId="29" captionText="FIGURE 11 Radula of Sarika spp. A) S.theodori specimen CUMZ 7942.B) S.siamensis specimen CUMZ 7951.C) S.costabilis sp. nov. paratype CUMZ 7945. D) S. costata sp. nov. paratype CUMZ 7947.Central tooth indicated by ‘C’ and blue colour; yellow colour indicates lateral teeth in the transition to marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.8349005" httpUri="https://zenodo.org/record/8349005/files/figure.png" pageId="29" pageNumber="126">11D</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9336EB036EFFCC76BCF9D09357F926" blockId="29.[645,1157,1608,1742]" pageId="29" pageNumber="126">
|
||
<emphasis id="B958EAF9036EFFCC76BCF9D09480F98A" box="[645,941,1608,1634]" italics="true" pageId="29" pageNumber="126">Type material examined.</emphasis>
|
||
<materialsCitation id="3B443CB6036EFFCC7786F9D093ACF96E" collectionCode="CUMZ" pageId="29" pageNumber="126" specimenCode="CUMZ 7946" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="54978849036EFFCC7786F9D09307F98A" box="[959,1066,1608,1634]" pageId="29" pageNumber="126">Holotype</typeStatus>
|
||
.
|
||
<specimenCode id="DB8A9E90036EFFCC7079F9D09594F96D" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="29" pageNumber="126">CUMZ 7946</specimenCode>
|
||
(
|
||
<figureCitation id="13172A6E036EFFCC76F0F9F39439F96E" box="[713,788,1643,1670]" captionStart="FIGURE 8" captionStartId="25.[97,164,1691,1712]" captionTargetBox="[193,1062,591,1673]" captionTargetId="figure-133@25.[193,1062,591,1673]" captionTargetPageId="25" captionText="FIGURE 8 Shells of Sarika spp.A, B) S. costabilis sp. nov.A) Holotype CUMZ 7943.B) Paratype CUMZ 7944.C, D) S. costata sp.nov.C) Holotype CUMZ 7946.D) Paratype CUMZ 7947." figureDoi="http://doi.org/10.5281/zenodo.8348999" httpUri="https://zenodo.org/record/8348999/files/figure.png" pageId="29" pageNumber="126">fig. 8C</figureCitation>
|
||
, width
|
||
<quantity id="4CD49B0E036EFFCC7753F9F494E5F96E" box="[874,968,1644,1670]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="29" pageNumber="126" unit="mm" value="10.8">10.8 mm</quantity>
|
||
, height
|
||
<quantity id="4CD49B0E036EFFCC701DF9F49354F96E" box="[1060,1145,1644,1670]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="29" pageNumber="126" unit="mm" value="7.2">7.2 mm</quantity>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B443CB6036EFFCC76BCF908935BF926" collectionCode="CUMZ" pageId="29" pageNumber="126" specimenCode="CUMZ 7947" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="54978849036EFFCC76BCF90895D9F942" box="[645,756,1680,1706]" pageId="29" pageNumber="126">Paratypes</typeStatus>
|
||
. Same locality as holotype:
|
||
<specimenCode id="DB8A9E90036EFFCC7079F9089595F925" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="29" pageNumber="126">CUMZ 7947</specimenCode>
|
||
(
|
||
<figureCitation id="13172A6E036EFFCC76FFF92B943DF925" box="[710,784,1715,1741]" captionStart="FIGURE 8" captionStartId="25.[97,164,1691,1712]" captionTargetBox="[193,1062,591,1673]" captionTargetId="figure-133@25.[193,1062,591,1673]" captionTargetPageId="25" captionText="FIGURE 8 Shells of Sarika spp.A, B) S. costabilis sp. nov.A) Holotype CUMZ 7943.B) Paratype CUMZ 7944.C, D) S. costata sp.nov.C) Holotype CUMZ 7946.D) Paratype CUMZ 7947." figureDoi="http://doi.org/10.5281/zenodo.8348999" httpUri="https://zenodo.org/record/8348999/files/figure.png" pageId="29" pageNumber="126">fig. 8D</figureCitation>
|
||
, width
|
||
<quantity id="4CD49B0E036EFFCC7758F92B9492F926" box="[865,959,1715,1742]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="29" pageNumber="126" unit="mm" value="10.9">10.9 mm</quantity>
|
||
, height 7.0 mm)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9336EB036DFFCF74A6FF33962AFED9" blockId="30.[129,640,171,1742]" pageId="30" pageNumber="127">
|
||
<emphasis id="B958EAF9036DFFCF74A6FF3396EBFF2D" box="[159,454,171,197]" italics="true" pageId="30" pageNumber="127">Other material examined.</emphasis>
|
||
<materialsCitation id="3B443CB6036DFFCF75EAFF33962EFED9" collectionCode="CUMZ" country="Myanmar" latitude="11.2295" location="Phra (Buddha) Cave" longLatPrecision="1" longitude="99.17619" municipality="Division" pageId="30" pageNumber="127" specimenCode="CUMZ 7948" specimenCount="1" stateProvince="Taninthayi">
|
||
<location id="8EF36030036DFFCF75EAFF339794FF01" LSID="urn:lsid:plazi:treatment:038587FD036EFFCE76BCFA439789FB3F:8EF36030036DFFCF75EAFF339794FF01" country="Myanmar" latitude="11.2295" longLatPrecision="1" longitude="99.17619" municipality="Division" name="Phra (Buddha) Cave" pageId="30" pageNumber="127" stateProvince="Taninthayi">Phra (Buddha) Cave</location>
|
||
,
|
||
<collectingRegion id="49E8F809036DFFCF74ECFF5796CDFF01" box="[213,480,207,233]" country="Myanmar" name="Taninthayi" pageId="30" pageNumber="127">Tanintharyi Township</collectingRegion>
|
||
, Tanintharyi
|
||
<collectingMunicipality id="6BF7AC91036DFFCF74B8FF6B97CDFEE5" box="[129,224,243,269]" pageId="30" pageNumber="127">Division</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="F33B767B036DFFCF74D7FF6C9676FEE6" box="[238,347,244,270]" name="Myanmar" pageId="30" pageNumber="127">Myanmar</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE18502C036DFFCF7550FF6B96C4FEE5" box="[361,489,243,269]" degrees="11" direction="north" minutes="13" orientation="latitude" pageId="30" pageNumber="127" precision="1" seconds="46.2" value="11.2295">11°13’46.2”N</geoCoordinate>
|
||
<geoCoordinate id="EE18502C036DFFCF75CAFF6B9551FEE5" box="[499,636,243,269]" degrees="99" direction="east" minutes="10" orientation="longitude" pageId="30" pageNumber="127" precision="1" seconds="34.3" value="99.17619">99°10’34.3”E</geoCoordinate>
|
||
,
|
||
<specimenCode id="DB8A9E90036DFFCF74B8FE8F962EFED9" box="[129,259,279,305]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="30" pageNumber="127">CUMZ 7948</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9336EB036DFFCF74A6FEA49538FE76" blockId="30.[129,640,171,1742]" pageId="30" pageNumber="127">
|
||
<materialsCitation id="3B443CB6036DFFCF74A6FEA4953CFE76" country="Myanmar" county="Division" latitude="12.43175" location="Kala Island" longLatPrecision="1" longitude="98.49897" municipality="Myeik District" pageId="30" pageNumber="127" specimenCount="1" stateProvince="Taninthayi" typeStatus="holotype">
|
||
<emphasis id="B958EAF9036DFFCF74A6FEA49603FEBD" box="[159,302,315,341]" italics="true" pageId="30" pageNumber="127">
|
||
<typeStatus id="54978849036DFFCF74A6FEA497FEFEBD" box="[159,211,316,341]" pageId="30" pageNumber="127">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<location id="8EF36030036DFFCF750CFEA39698FEBD" LSID="urn:lsid:plazi:treatment:038587FD036EFFCE76BCFA439789FB3F:8EF36030036DFFCF750CFEA39698FEBD" box="[309,437,315,341]" country="Myanmar" county="Division" latitude="12.43175" longLatPrecision="1" longitude="98.49897" municipality="Myeik District" name="Kala Island" pageId="30" pageNumber="127" stateProvince="Taninthayi">Kala Island</location>
|
||
,
|
||
<location id="8EF36030036DFFCF75F8FEA39551FEBD" LSID="urn:lsid:plazi:treatment:038587FD036EFFCE76BCFA439789FB3F:8EF36030036DFFCF75F8FEA39551FEBD" box="[449,636,315,341]" country="Myanmar" county="Division" latitude="12.43175" longLatPrecision="1" longitude="98.49897" municipality="Myeik District" name="Myeik Township" pageId="30" pageNumber="127" stateProvince="Taninthayi">Myeik Township</location>
|
||
,
|
||
<collectingMunicipality id="6BF7AC91036DFFCF74B8FEC79668FE91" box="[129,325,351,377]" pageId="30" pageNumber="127">Myeik District</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49E8F809036DFFCF7548FEC796D9FE91" box="[369,500,351,377]" country="Myanmar" name="Taninthayi" pageId="30" pageNumber="127">Tanintharyi</collectingRegion>
|
||
<collectingCounty id="62F24E67036DFFCF7625FEC79556FE91" box="[540,635,351,377]" pageId="30" pageNumber="127">Division</collectingCounty>
|
||
,
|
||
<collectingCountry id="F33B767B036DFFCF74B8FE1B97C3FE75" box="[129,238,387,413]" name="Myanmar" pageId="30" pageNumber="127">Myanmar</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE18502C036DFFCF74CEFE1B9653FE76" box="[247,382,387,414]" degrees="12" direction="north" minutes="25" orientation="latitude" pageId="30" pageNumber="127" precision="1" seconds="54.3" value="12.43175">12°25’54.3”N</geoCoordinate>
|
||
<geoCoordinate id="EE18502C036DFFCF75BDFE1B953CFE76" box="[388,529,387,414]" degrees="98" direction="east" minutes="29" orientation="longitude" pageId="30" pageNumber="127" precision="1" seconds="56.3" value="98.49897">98°29’56.3”E</geoCoordinate>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9336EB036DFFCF74A6FE3F9639FDB9" blockId="30.[129,640,171,1742]" pageId="30" pageNumber="127">
|
||
<emphasis id="B958EAF9036DFFCF74A6FE3F963AFE29" box="[159,279,423,449]" italics="true" pageId="30" pageNumber="127">Etymology.</emphasis>
|
||
The specific name “
|
||
<taxonomicName id="4C2C4D68036DFFCF7632FE309577FE29" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" box="[523,602,424,449]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="127" phylum="Mollusca" rank="species" species="costata">
|
||
<emphasis id="B958EAF9036DFFCF7632FE309577FE29" box="[523,602,424,449]" italics="true" pageId="30" pageNumber="127">costata</emphasis>
|
||
</taxonomicName>
|
||
” is from the Latin word meaning “ribbed”, which refers to the shell surface having fine radial ribs that characterises this species within the genus
|
||
<taxonomicName id="4C2C4D68036DFFCF74F1FDAE9623FDB8" authorityName="Godwin-Austen" authorityYear="1907" box="[200,270,566,592]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="127" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B958EAF9036DFFCF74F1FDAE9623FDB8" box="[200,270,566,592]" italics="true" pageId="30" pageNumber="127">Sarika</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9336EB036DFFCF74A6FDC3957BFC98" blockId="30.[129,640,171,1742]" pageId="30" pageNumber="127">
|
||
<emphasis id="B958EAF9036DFFCF74A6FDC396E5FD9C" box="[159,456,602,628]" italics="true" pageId="30" pageNumber="127">Diagnostic characteristics.</emphasis>
|
||
Shell medium, globosely depressed; body whorl well-rounded; shell ribbed with smooth radial ribs; aperture narrowly crescent-shaped. Animal pale grey body, black stripes behind long tentacles and five mantle edges. Genitalia with straight epiphallic caecum, cuboidal, and irregularly oblique-folded penial pilasters.
|
||
</paragraph>
|
||
<paragraph id="8B9336EB036DFFCF74A6FCE2966BFA3A" blockId="30.[129,640,171,1742]" pageId="30" pageNumber="127">
|
||
<emphasis id="B958EAF9036DFFCF74A6FCE29676FC7B" box="[159,347,889,915]" italics="true" pageId="30" pageNumber="127">Description.Shell</emphasis>
|
||
(
|
||
<figureCitation id="13172A6E036DFFCF755FFCE196E6FC7C" box="[358,459,889,916]" captionStart="FIGURE 8" captionStartId="25.[97,164,1691,1712]" captionTargetBox="[193,1062,591,1673]" captionTargetId="figure-133@25.[193,1062,591,1673]" captionTargetPageId="25" captionText="FIGURE 8 Shells of Sarika spp.A, B) S. costabilis sp. nov.A) Holotype CUMZ 7943.B) Paratype CUMZ 7944.C, D) S. costata sp.nov.C) Holotype CUMZ 7946.D) Paratype CUMZ 7947." figureDoi="http://doi.org/10.5281/zenodo.8348999" httpUri="https://zenodo.org/record/8348999/files/figure.png" pageId="30" pageNumber="127">fig.8C, D</figureCitation>
|
||
).Shell globosely depressed, medium size (width up to
|
||
<quantity id="4CD49B0E036DFFCF7626FC069556FC50" box="[543,635,926,952]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="30" pageNumber="127" unit="mm" value="12.4">12.4 mm</quantity>
|
||
, height up to 8.0 mm) and rather thin to slightly solid. Shell colour yellowish-brown to brownish. Whorls 6–6¼, increasing regularly. Spire much elevated; apex slightly protruding; suture impressed, slightly wider near aperture opening. Embryonic shell rather smooth; later whorl with very weak radial ridges. Body whorl large and well-rounded. Upper shell surface sculptured with regular, distinct, and smooth radial ribs, and then radial ribs disappearing below periphery. Aperture crescent-shaped, narrow, well oblique, and with simple lip. Columellar margin simple and slightly reflected near umbilicus. Umbilicus narrowly opened.
|
||
</paragraph>
|
||
<paragraph id="8B9336EB036DFFCF74A6FA44943FFDE0" blockId="30.[129,640,171,1742]" lastBlockId="30.[676,1188,171,1741]" pageId="30" pageNumber="127">
|
||
<emphasis id="B958EAF9036DFFCF74A6FA449665FA1E" box="[159,328,1500,1526]" italics="true" pageId="30" pageNumber="127">Genital organs</emphasis>
|
||
(
|
||
<figureCitation id="13172A6E036DFFCF7566FA4496F6FA1E" box="[351,475,1500,1526]" captionStart="FIGURE 9" captionStartId="27.[98,165,1350,1371]" captionTargetBox="[193,1062,164,1331]" captionTargetId="figure-97@27.[193,1062,164,1331]" captionTargetPageId="27" captionText="FIGURE 9 Genitalia of Sarika spp.A, B) S. costabilis sp. nov.paratype CUMZ 7945.A) General view of the genital system.B) Internal structure of penis.C, D) S.costata sp. nov. paratype CUMZ 7947.C) General view of the genital system.D) Internal structure of penis. White arrows indicate the end of penis." figureDoi="http://doi.org/10.5281/zenodo.8349001" httpUri="https://zenodo.org/record/8349001/files/figure.png" pageId="30" pageNumber="127">fig. 9C, D</figureCitation>
|
||
). Atrium (at) enlarged and very short. Penis (p) elongate and cylindrical. Inner surface sculptured with very finely longitudinal pilasters (pp) to nearly smooth, about one-third of penis length, covered with cuboidal pilasters in middle part, and then transformed to irregular folds arranged in oblique row about one-third of penis length. Epiphallus (e) elongate, cylindrical, and same length as penis. Epiphallic caecum (ec) straight, slightly smaller diameter than epiphallus and located at middle of epiphallus. Penial retractor muscle (prm) thin and attached at tip of epiphallic caecum. Flagellum (fl) slender, same length as epiphallus, and enlarged at the tip. Vas deferens (vd) thin tube.
|
||
</paragraph>
|
||
<paragraph id="8B9336EB036DFFCF76FBFD8A9339FCC0" blockId="30.[676,1188,171,1741]" pageId="30" pageNumber="127">Vagina (v) cylindrical and slightly shorter than penis. Dart apparatus (da) enlarged, very long, and joined to atrium at vagina and penis junction. Gametolytic sac (gs) bulbous; gametolytic duct (gd) elongate cylindrical. Free oviduct (fo) enlarged cylindrical, slightly shorter than vagina. Oviduct large lobules; prostate gland running alongside oviduct.</paragraph>
|
||
<paragraph id="8B9336EB036DFFCF76FBFCAA932DFB67" blockId="30.[676,1188,171,1741]" pageId="30" pageNumber="127">
|
||
<taxonomicName id="4C2C4D68036DFFCF76FBFCAA943CFCA4" box="[706,785,818,844]" class="Bivalvia" family="Limidae" genus="Radula" kingdom="Animalia" order="Limida" pageId="30" pageNumber="127" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B958EAF9036DFFCF76FBFCAA943CFCA4" box="[706,785,818,844]" italics="true" pageId="30" pageNumber="127">Radula</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13172A6E036DFFCF771CFCAA9450FCA4" box="[805,893,818,844]" captionStart="FIGURE 11" captionStartId="29.[98,165,1042,1063]" captionTargetBox="[193,1062,164,1023]" captionTargetId="figure-192@29.[193,1062,164,1023]" captionTargetPageId="29" captionText="FIGURE 11 Radula of Sarika spp. A) S.theodori specimen CUMZ 7942.B) S.siamensis specimen CUMZ 7951.C) S.costabilis sp. nov. paratype CUMZ 7945. D) S. costata sp. nov. paratype CUMZ 7947.Central tooth indicated by ‘C’ and blue colour; yellow colour indicates lateral teeth in the transition to marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.8349005" httpUri="https://zenodo.org/record/8349005/files/figure.png" pageId="30" pageNumber="127">fig. 11D</figureCitation>
|
||
). Teeth arranged straight with half row formula: 1–(11–12)–50. Central tooth symmetrical tricuspid with large mesocone and small ectocones. Lateral teeth asymmetrical tricuspid with large mesocone, and small endocone and ectocone. Marginal teeth started around tooth number 11 or 12; inner teeth elongate bicuspid with lanceolate endocone and small ectocone; and outermost teeth gradually reduced in size.
|
||
</paragraph>
|
||
<paragraph id="8B9336EB036DFFCF76FBFB0195D2FA1E" blockId="30.[676,1188,171,1741]" pageId="30" pageNumber="127">
|
||
<emphasis id="B958EAF9036DFFCF76FBFB019452FB5B" box="[706,895,1177,1203]" italics="true" pageId="30" pageNumber="127">External features</emphasis>
|
||
(
|
||
<figureCitation id="13172A6E036DFFCF77B4FB0194EAFB5B" box="[909,967,1177,1203]" captionStart="FIGURE 2" captionStartId="11.[100,167,1258,1279]" captionTargetBox="[193,1061,164,1238]" captionTargetId="figure-122@11.[193,1062,164,1239]" captionTargetPageId="11" captionText="FIGURE 2 Bayesian inference tree based on a 1495 bp concatenated alignment dataset of COI + 16S + 28S genes. Numbers by the nodes are the Bayesian posterior probabilities (left) and ML bootstrap values (right); shown only for the nodes supported by BI ≥ 0.95 or ML ≥ 70%. Clade A consists of the S. resplendens-S. hainesi species groups.Clade B consists of the S.siamensis species group.Images of living snails are not to scale." figureDoi="http://doi.org/10.5281/zenodo.8348979" httpUri="https://zenodo.org/record/8348979/files/figure.png" pageId="30" pageNumber="127">fig. 2</figureCitation>
|
||
). Living snails with reticulated skin, dark grey eye stalks, and very pale grey body with black stripes behind long tentacles. Five well-developed mantle lobes with same colour as body. Right shell lobe moderately elongate; left shell lobe short. Dorsal lobes large and broad; anterior and posterior left dorsal lobes smaller than right dorsal lobe. Caudal fossa and caudal horn present.
|
||
</paragraph>
|
||
<paragraph id="8B9336EB036DFFCF76FBF9989406F925" blockId="30.[676,1188,171,1741]" pageId="30" pageNumber="127">
|
||
<emphasis id="B958EAF9036DFFCF76FBF998946BF9F2" box="[706,838,1536,1562]" italics="true" pageId="30" pageNumber="127">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="13172A6E036DFFCF776EF99894A3F9F2" box="[855,910,1536,1562]" captionStart="FIGURE 1" captionStartId="3.[99,166,977,998]" captionTargetBox="[193,1062,164,958]" captionTargetId="figure-233@3.[193,1062,164,958]" captionTargetPageId="3" captionText="FIGURE 1 Geographic distribution of ribbed shell Sarika spp. based on the specimens examined herein. A) Distribution of S. theodori, S.costabilis sp.nov., and S. costata sp. nov. B) Distribution of S. siamensis. Black symbols indicate the DNA extracted specimens used in the molecular analysis; white symbols indicate shells or other preserved specimens." figureDoi="http://doi.org/10.5281/zenodo.8348977" httpUri="https://zenodo.org/record/8348977/files/figure.png" pageId="30" pageNumber="127">fig. 1</figureCitation>
|
||
).
|
||
<taxonomicName id="4C2C4D68036DFFCF779CF998936EF9F1" authority="Pholyotha & Sutcharit & Lin & Panha, 2022" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" box="[933,1091,1536,1562]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="127" phylum="Mollusca" rank="species" species="costata" status="sp. nov.">
|
||
<emphasis id="B958EAF9036DFFCF779CF998936EF9F1" box="[933,1091,1536,1562]" italics="true" pageId="30" pageNumber="127">Sarika costata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B5782036DFFCF7074F9989389F9F2" box="[1101,1188,1536,1562]" pageId="30" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is known from two localities, Kala Island and on the mainland at Phra Cave in
|
||
<collectingRegion id="49E8F809036DFFCF7019F9DF95D9F96D" country="Myanmar" name="Taninthayi" pageId="30" pageNumber="127">Tanintharyi Region</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33B767B036DFFCF76C5F9F49447F96E" box="[764,874,1644,1670]" name="Myanmar" pageId="30" pageNumber="127">Myanmar</collectingCountry>
|
||
. Its distribution range needs further studies. We could not find this species in
|
||
<collectingCountry id="F33B767B036DFFCF76F9F92B940AF925" box="[704,807,1715,1741]" name="Thailand" pageId="30" pageNumber="127">Thailand</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9336EB036CFFCE74B8FF3396C7FD9C" blockId="31.[98,610,171,1239]" pageId="31" pageNumber="128">
|
||
<emphasis id="B958EAF9036CFFCE74B8FF3397CAFF2D" box="[129,231,171,197]" italics="true" pageId="31" pageNumber="128">Remarks.</emphasis>
|
||
Regardless of the genitalia and mantle characters,
|
||
<taxonomicName id="4C2C4D68036CFFCE7502FF5796FAFF01" authority="Pholyotha & Sutcharit & Lin & Panha, 2022" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" box="[315,471,207,233]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="128" phylum="Mollusca" rank="species" species="costata" status="sp. nov.">
|
||
<emphasis id="B958EAF9036CFFCE7502FF5796FAFF01" box="[315,471,207,233]" italics="true" pageId="31" pageNumber="128">Sarika costata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B5782036CFFCE75D9FF489519FF02" box="[480,564,208,234]" pageId="31" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C2C4D68036CFFCE74F5FF6C966DFEE5" authorityName=": Godwin-Austen" authorityYear="1907" box="[204,320,243,270]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="128" phylum="Mollusca" rank="species" species="theodori">
|
||
<emphasis id="B958EAF9036CFFCE74F5FF6C966DFEE5" box="[204,320,243,270]" italics="true" pageId="31" pageNumber="128">S. theodori</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2C4D68036CFFCE7572FF6C96E0FEE5" box="[331,461,244,270]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="128" phylum="Mollusca" rank="species" species="siamensis">
|
||
<emphasis id="B958EAF9036CFFCE7572FF6C96E0FEE5" box="[331,461,244,270]" italics="true" pageId="31" pageNumber="128">S. siamensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C2C4D68036CFFCE7630FF6C97A2FED9" authority="Pholyotha & Sutcharit & Lin & Panha, 2022" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="128" phylum="Mollusca" rank="species" species="costabilis" status="sp. nov.">
|
||
<emphasis id="B958EAF9036CFFCE7630FF6C97A2FED9" italics="true" pageId="31" pageNumber="128">S. costabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B5782036CFFCE74AAFE8097C9FEDA" box="[147,228,280,306]" pageId="31" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by having small and smooth radial ribs, while the three latter species have relatively larger shell with nodules or dash-shaped nodules arranged on radial ribs. Moreover, this new species has five mantle edges and the inner wall of the penis is sculptured with cuboidal and folded penial pilasters, while the other three species have four mantle edges, and the inner wall of the penis is sculptured with only cuboidal penial pilasters.
|
||
</paragraph>
|
||
<paragraph id="8B9336EB036CFFCE74B8FDE69789FB3F" blockId="31.[98,610,171,1239]" pageId="31" pageNumber="128">
|
||
Compared with the medium-sized species with costulate striation on the shell surface from
|
||
<collectingCountry id="F33B767B036CFFCE74E5FD5E966DFD08" box="[220,320,710,736]" name="Thailand" pageId="31" pageNumber="128">Thailand</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33B767B036CFFCE75B8FD5F96C3FD09" box="[385,494,711,737]" name="Myanmar" pageId="31" pageNumber="128">Myanmar</collectingCountry>
|
||
, the shell morphology of
|
||
<taxonomicName id="4C2C4D68036CFFCE7523FD7296AAFCEC" authority="Pholyotha & Sutcharit & Lin & Panha, 2022" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" box="[282,391,746,772]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="128" phylum="Mollusca" rank="species" species="costata" status="sp. nov.">
|
||
<emphasis id="B958EAF9036CFFCE7523FD7296AAFCEC" box="[282,391,746,772]" italics="true" pageId="31" pageNumber="128">S. costata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B5782036CFFCE75AAFD7296C6FCEC" box="[403,491,746,772]" pageId="31" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is similar to
|
||
<taxonomicName id="4C2C4D68036CFFCE74BDFC9696C0FCC1" authority="(Benson, 1856)" baseAuthorityName="Benson" baseAuthorityYear="1856" box="[132,493,782,809]" class="Gastropoda" family="Ariophantidae" genus="Khasiella" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="128" phylum="Mollusca" rank="species" species="pansa">
|
||
<emphasis id="B958EAF9036CFFCE74BDFC96961AFCC0" box="[132,311,782,808]" italics="true" pageId="31" pageNumber="128">Khasiella pansa</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBD4B1A036CFFCE7573FC9696CBFCC1" author="Benson, W. H." box="[330,486,782,809]" pageId="31" pageNumber="128" pagination="249 - 254" refId="ref14962" refString="Benson, W. H. (1856) Descriptions of one Indian and nine new Burmese Helices; and notes on two Burmese Cyclostomacea. Ann. Mag. Nat. Hist., 18, 249 - 254." type="journal article" year="1856">Benson, 1856</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C2C4D68036CFFCE75C2FC969788FCA4" baseAuthorityName="Benson" baseAuthorityYear="1856" class="Gastropoda" family="Ariophantidae" genus="Khasiella" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="128" phylum="Mollusca" rank="species" species="pansa">
|
||
<emphasis id="B958EAF9036CFFCE75C2FC969788FCA4" italics="true" pageId="31" pageNumber="128">Khasiella pansa</emphasis>
|
||
</taxonomicName>
|
||
was described from the
|
||
<collectingRegion id="49E8F809036CFFCE7590FCAA9536FCA4" box="[425,539,818,844]" country="Myanmar" name="Ayeyawady" pageId="31" pageNumber="128">Irrawaddy</collectingRegion>
|
||
Valley and recorded from the Mergui Archipelago (
|
||
<bibRefCitation id="EFBD4B1A036CFFCE7453FCE2962BFC7C" author="Benson, W. H." box="[106,262,889,916]" pageId="31" pageNumber="128" pagination="249 - 254" refId="ref14962" refString="Benson, W. H. (1856) Descriptions of one Indian and nine new Burmese Helices; and notes on two Burmese Cyclostomacea. Ann. Mag. Nat. Hist., 18, 249 - 254." type="journal article" year="1856">Benson, 1856</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBD4B1A036CFFCE7520FCE197B5FC50" author="Blanford, W. T. & Godwin-Austen, H. H." pageId="31" pageNumber="128" refId="ref15040" refString="Blanford, W. T. & Godwin-Austen, H. H. (1908) The fauna of British India. Mollusca. Testacellidae and Zonitidae. In: C. T. Bingham (Ed) The Fauna of British India including Ceylon and Burma. Taylor and Francis, London. https: // www. biodiversitylibrary. org / item / 47068 # page / 5 / mode / 1 up." type="url" year="1908">Blanford & Godwin-Austen, 1908</bibRefCitation>
|
||
); however, the anatomical data of
|
||
<taxonomicName id="4C2C4D68036CFFCE7672FC069788FC33" baseAuthorityName="Benson" baseAuthorityYear="1856" class="Gastropoda" family="Ariophantidae" genus="Khasiella" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="128" phylum="Mollusca" rank="species" species="pansa">
|
||
<emphasis id="B958EAF9036CFFCE7672FC069788FC33" italics="true" pageId="31" pageNumber="128">K. pansa</emphasis>
|
||
</taxonomicName>
|
||
needs further studies. In comparison, this new species has a smaller shell size, well-rounded body whorl, and narrowly crescent-shaped aperture, while
|
||
<taxonomicName id="4C2C4D68036CFFCE75FBFBB69507FBAF" baseAuthorityName="Benson" baseAuthorityYear="1856" box="[450,554,1070,1095]" class="Gastropoda" family="Ariophantidae" genus="Khasiella" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="128" phylum="Mollusca" rank="species" species="pansa">
|
||
<emphasis id="B958EAF9036CFFCE75FBFBB69507FBAF" box="[450,554,1070,1095]" italics="true" pageId="31" pageNumber="128">K. pansa</emphasis>
|
||
</taxonomicName>
|
||
has a slightly larger, subangulate body whorl, and wider aperture (
|
||
<bibRefCitation id="EFBD4B1A036CFFCE7568FBED96CAFB78" author="Benson, W. H." box="[337,487,1141,1168]" pageId="31" pageNumber="128" pagination="249 - 254" refId="ref14962" refString="Benson, W. H. (1856) Descriptions of one Indian and nine new Burmese Helices; and notes on two Burmese Cyclostomacea. Ann. Mag. Nat. Hist., 18, 249 - 254." type="journal article" year="1856">Benson, 1856</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBD4B1A036CFFCE75CDFBED963DFB5B" author="Hanley, S. & Theobald, W." pageId="31" pageNumber="128" refId="ref15679" refString="Hanley, S. & Theobald, W. (1876 [1870 - 1876]) Conchologica Indica: Illustrations of the Land and Freshwater Shells of British India. L. Reeve & Co, Covent Garden, London." type="book" year="1876">Hanley & Theobald, 1876</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBD4B1A036CFFCE7526FB0197B5FB3F" author="Blanford, W. T. & Godwin-Austen, H. H." pageId="31" pageNumber="128" refId="ref15040" refString="Blanford, W. T. & Godwin-Austen, H. H. (1908) The fauna of British India. Mollusca. Testacellidae and Zonitidae. In: C. T. Bingham (Ed) The Fauna of British India including Ceylon and Burma. Taylor and Francis, London. https: // www. biodiversitylibrary. org / item / 47068 # page / 5 / mode / 1 up." type="url" year="1908">Blanford & Godwin-Austen, 1908</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |