166 lines
18 KiB
XML
166 lines
18 KiB
XML
<document id="E16F2C92EA3EE1A91A958E50836C21CF" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE313FF86FF75FEE9BB12FCBD" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena genuvela Perkins 2017, new species" docType="treatment" docVersion="9" lastPageNumber="24" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="21" updateTime="1698480684898" updateUser="ExternalLinkService">
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<mods:mods id="6B54786E6D634AA3BFC71F6A3A372919" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B0DA726F2AB4A34E4A85945BBACF874D">
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<mods:title id="880B1FB88EE716C83EA88DFFA44B33E3">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
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</mods:titleInfo>
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<mods:name id="C35925EA136DADE41CF612B79ED04E4F" type="personal">
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<mods:roleTerm id="6D3482C3DF4D15CFBB4AAB843BC8B973">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DF0E55E8D9DDB5D4EF3C5C4B39787994">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="5F7F3CA1B2BD2C55E96473A867B14FA8">text</mods:typeOfResource>
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<mods:titleInfo id="3A3D2B8BD5DBC13DBECC20B105BC758C">
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<mods:title id="6AD98892D63BD255BF521249423FCB9A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="28E43CE919AF4D222C72B084A12D46AF">
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<mods:date id="9CE976C06907F761296AFFEF2C629C66">2017</mods:date>
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<mods:detail id="EA46F6253160997CC01DA7EFB88193D3" type="pubDate">
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<mods:number id="CD057543E992912D61E45C4336DCF820">2017-11-03</mods:number>
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<mods:detail id="E4DD15D70873BB5C567444FD51CA2B1C" type="volume">
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<mods:number id="B4BD0499672CF576F456A7366FAF12AA">4342</mods:number>
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</mods:detail>
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<mods:detail id="2880208D5F0775DEACB3422EB15FCF38" type="issue">
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<mods:number id="44AD797FAE71E22BAA73EDA2C09C84AC">1</mods:number>
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<mods:start id="1C1EC770E3709ED33D0F2819DC14428A">1</mods:start>
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<mods:classification id="78932F3D30567217974C2B0982E876C5">journal article</mods:classification>
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<mods:identifier id="32BEEDCA53CCA9C149693C5E466DD3EB" type="CLB-Dataset">31639</mods:identifier>
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<mods:identifier id="5E39FEB3B196E08F7CEF2532DBC304DF" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
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<mods:identifier id="7104543E675C07607A669DFBF3AA9427" type="GBIF-Dataset">15799d2a-1968-4874-8100-452597d4e26b</mods:identifier>
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<mods:identifier id="FA3CCE5E08A069B9A4CAE970D55EB9A0" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="9656E4FA1FF3A65276525E830868595E" type="Zenodo-Dep">1041066</mods:identifier>
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<mods:identifier id="7FF8EA573E16B828986080DECC876DE4" type="ZooBank">2ACD54D2-3487-432D-9323-EEC131FE2E64</mods:identifier>
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</mods:mods>
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<treatment id="038587BBE313FF86FF75FEE9BB12FCBD" ID-DOI="http://doi.org/10.5281/zenodo.5323387" ID-GBIF-Taxon="137233442" ID-Zenodo-Dep="5323387" LSID="urn:lsid:plazi:treatment:038587BBE313FF86FF75FEE9BB12FCBD" httpUri="http://treatment.plazi.org/id/038587BBE313FF86FF75FEE9BB12FCBD" lastPageId="23" lastPageNumber="24" pageId="20" pageNumber="21">
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<subSubSection id="C3366526E313FF85FF75FEE9BB46FE02" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="8B9336ADE313FF85FF75FEE9BB1EFEE3" blockId="20.[151,634,332,391]" box="[151,546,332,358]" pageId="20" pageNumber="21">
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<heading id="D0DB81C1E313FF85FF75FEE9BB1EFEE3" bold="true" box="[151,546,332,358]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
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||
<emphasis id="B958EABFE313FF85FF75FEE9BB1EFEE3" bold="true" box="[151,546,332,358]" pageId="20" pageNumber="21">
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||
<emphasis id="B958EABFE313FF85FF75FEE9B8B7FEE3" bold="true" box="[151,395,332,358]" italics="true" pageId="20" pageNumber="21">
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<taxonomicName id="4C2C4D2EE313FF85FF75FEE9B8B8FEE3" ID-CoL="3N3L8" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[151,388,332,358]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="genuvela" status="sp. nov.">Hydraena genuvela</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A26B57C4E313FF85FE70FEE9BB1EFEE3" box="[402,546,332,358]" pageId="20" pageNumber="21" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9336ADE313FF85FF75FECBBB46FE02" blockId="20.[151,634,332,391]" box="[151,634,366,391]" pageId="20" pageNumber="21">
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<figureCitation id="13172A28E313FF85FF75FECBB9DFFE02" box="[151,227,366,391]" captionStart="FIGURE 9" captionStartId="20.[151,250,1968,1990]" captionText="FIGURE 9. Hydraena genuvela, holotype, habitus dorsal, lateral, and ventral." pageId="20" pageNumber="21">Figs. 9</figureCitation>
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(habitus),
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<figureCitation id="13172A28E313FF85FEBEFECAB844FE02" box="[348,376,367,391]" captionStart="FIGURES 12 – 13" captionStartId="22.[151,264,1876,1899]" captionText="FIGURES 12 – 13. Hydraena, aedeagi of holotypes." pageId="20" pageNumber="21">13</figureCitation>
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(aedeagus),
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<figureCitation id="13172A28E313FF85FDE4FECABB0CFE02" box="[518,560,367,391]" captionStart="FIGURES 233 – 236" captionStartId="228.[151,264,1801,1824]" captionTargetBox="[328,1258,203,1770]" captionTargetId="figure@228.[317,1269,193,1780]" captionTargetPageId="228" captionText="FIGURES 233 – 236. All locality sites for Hydraena, Hydraena (Micromadraena), Limnebius, and Madagaster." httpUri="https://zenodo.org/record/1041152/files/figure.png" pageId="20" pageNumber="21">234</figureCitation>
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(map)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E313FF85FF75FE13BBB9FD92" pageId="20" pageNumber="21" type="materials_examined">
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<paragraph id="8B9336ADE313FF85FF75FE13BBB9FD92" blockId="20.[151,1436,438,643]" pageId="20" pageNumber="21">
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<emphasis id="B958EABFE313FF85FF75FE13B887FE4A" bold="true" box="[151,443,438,463]" pageId="20" pageNumber="21">
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<typeStatus id="5497880FE313FF85FF75FE13B9EEFE4A" box="[151,210,438,463]" pageId="20" pageNumber="21">Type</typeStatus>
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Material.
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<typeStatus id="5497880FE313FF85FEACFE13B887FE4A" box="[334,443,438,463]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
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</emphasis>
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(male):
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<collectingRegion id="49E8F84FE313FF85FDF8FE13BB93FE4A" box="[538,687,438,463]" country="Madagascar" name="Antsiranana" pageId="20" pageNumber="21">
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<emphasis id="B958EABFE313FF85FDF8FE13BB93FE4A" bold="true" box="[538,687,438,463]" pageId="20" pageNumber="21">Antsiranana</emphasis>
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</collectingRegion>
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, Forêt de Binara,
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<quantity id="4CD49B48E313FF85FC9BFE12BAFBFE4A" box="[889,967,438,463]" metricMagnitude="3" metricUnit="m" metricValue="9.1" pageId="20" pageNumber="21" unit="km" value="9.1">9.1 km</quantity>
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233° SW Daraina, sifted litter (leaf mold, rotten wood) rainforest, elev.
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<quantity id="4CD49B48E313FF85FE1BFE7FBBBCFE77" box="[505,640,474,499]" metricMagnitude="2" metricUnit="m" metricValue="7.25" metricValueMax="8.0" metricValueMin="6.5" pageId="20" pageNumber="21" unit="m" value="725.0" valueMax="800.0" valueMin="650.0">650–800 m</quantity>
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,
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<geoCoordinate id="EE18506AE313FF85FD70FE7EBA39FE76" box="[658,773,474,499]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="-13.25">13° 15' S</geoCoordinate>
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,
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<geoCoordinate id="EE18506AE313FF85FCF5FE7EBAB0FE77" box="[791,908,474,499]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="49.6">49° 36' E</geoCoordinate>
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,
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<date id="FF92106DE313FF85FC7CFE7EBD23FE76" box="[926,1055,474,499]" pageId="20" pageNumber="21" value="2003-12-03">3 xii 2003</date>
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, B.L.Fisher (BLF 9656) (
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<collectionCode id="ED3DAE68E313FF85FAB6FE7EBCACFE76" box="[1364,1424,475,499]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="20" pageNumber="21">CAS</collectionCode>
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).
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<typeStatus id="5497880FE313FF85FF75FE5BB833FD92" box="[151,271,510,535]" pageId="20" pageNumber="21" type="paratype">
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<emphasis id="B958EABFE313FF85FF75FE5BB833FD92" bold="true" box="[151,271,510,535]" pageId="20" pageNumber="21">Paratypes</emphasis>
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</typeStatus>
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: Same data as
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<typeStatus id="5497880FE313FF85FE54FE5BBB24FD92" box="[438,536,510,535]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
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(2
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<collectionCode id="ED3DAE68E313FF85FDDCFE5ABB45FD92" box="[574,633,511,535]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="20" pageNumber="21">CAS</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E313FF85FF25FD87BC06FD06" pageId="20" pageNumber="21" type="diagnosis">
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<paragraph id="8B9336ADE313FF85FF25FD87BC06FD06" blockId="20.[151,1436,438,643]" pageId="20" pageNumber="21">
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<emphasis id="B958EABFE313FF85FF25FD87B8E9FDBE" bold="true" box="[199,469,546,571]" pageId="20" pageNumber="21">Differential Diagnosis.</emphasis>
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Very similar in size, habitus and plaques to
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<taxonomicName id="4C2C4D2EE313FF85FC00FD81BD61FDBE" authorityName="Perkins" authorityYear="2017" box="[994,1117,547,571]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fortipes">
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<emphasis id="B958EABFE313FF85FC00FD81BD61FDBE" box="[994,1117,547,571]" italics="true" pageId="20" pageNumber="21">H. fortipes</emphasis>
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</taxonomicName>
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. Differing therefrom in the slightly more coarsely and more densely punctate dorsum. Reliable determinations will require examining the aedeagi, which differ in shapes of the main piece and the rigid gonopore bearing distal tube (
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<figureCitation id="13172A28E313FF85FB47FDCFBC10FD06" box="[1189,1324,618,643]" captionStart="FIGURES 12 – 13" captionStartId="22.[151,264,1876,1899]" captionText="FIGURES 12 – 13. Hydraena, aedeagi of holotypes." pageId="20" pageNumber="21">Figs. 12, 13</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<caption id="DF536625E313FF85FF75F815BA8CF843" box="[151,944,1968,1990]" pageId="20" pageNumber="21">
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<paragraph id="8B9336ADE313FF85FF75F815BA8CF843" blockId="20.[151,944,1968,1990]" box="[151,944,1968,1990]" pageId="20" pageNumber="21">
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<emphasis id="B958EABFE313FF85FF75F815B82EF840" bold="true" box="[151,274,1968,1990]" pageId="20" pageNumber="21">FIGURE 9.</emphasis>
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<taxonomicName id="4C2C4D2EE313FF85FEFBF815B8E3F843" authorityName="Perkins" authorityYear="2017" box="[281,479,1968,1990]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="genuvela">
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<emphasis id="B958EABFE313FF85FEFBF815B8E3F843" box="[281,479,1968,1990]" italics="true" pageId="20" pageNumber="21">Hydraena genuvela</emphasis>
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</taxonomicName>
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, holotype, habitus dorsal, lateral, and ventral.
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</paragraph>
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</caption>
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<caption id="DF536625E312FF84FF75F88FBB9FF8C4" box="[151,675,1834,1857]" pageId="21" pageNumber="22">
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<paragraph id="8B9336ADE312FF84FF75F88FBB9FF8C4" blockId="21.[151,675,1834,1857]" box="[151,675,1834,1857]" pageId="21" pageNumber="22">
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<emphasis id="B958EABFE312FF84FF75F88FB86EF8C5" bold="true" box="[151,338,1834,1857]" pageId="21" pageNumber="22">FIGURES 10–11.</emphasis>
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<taxonomicName id="4C2C4D2EE312FF84FEBAF88EB881F8C4" box="[344,445,1835,1857]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EABFE312FF84FEBAF88EB881F8C4" box="[344,445,1835,1857]" italics="true" pageId="21" pageNumber="22">Hydraena</emphasis>
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</taxonomicName>
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, aedeagi of holotypes.
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</paragraph>
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</caption>
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<subSubSection id="C3366526E312FF86FF25F8C8BA66FD75" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22" type="description">
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<paragraph id="8B9336ADE312FF84FF25F8C8BA37F82F" blockId="21.[151,1436,1901,2034]" pageId="21" pageNumber="22">
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<emphasis id="B958EABFE312FF84FF25F8C8B86DF803" bold="true" box="[199,337,1901,1926]" pageId="21" pageNumber="22">Description</emphasis>
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. Size:
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<typeStatus id="5497880FE312FF84FE41F8C8BB39F803" box="[419,517,1901,1926]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.80/0.87; head width 0.49; pronotum 0.37/0.71, PA 0.59, PB 0.58; elytra 1.16/0.87.
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</paragraph>
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<paragraph id="8B9336ADE312FF84FF25F812BC58F877" blockId="21.[151,1436,1901,2034]" pageId="21" pageNumber="22">Dorsum dark brown, frons darkest; legs (except coxae), pronotal hypomeron, and elytral epipleura light brown; venter and coxae dark brown; maxillary palpi light brown to testaceous, distal ½ of last palpomere not darker.</paragraph>
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<caption id="DF536625E311FF87FF75F8F1BB98F8EE" box="[151,676,1876,1899]" pageId="22" pageNumber="23">
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<paragraph id="8B9336ADE311FF87FF75F8F1BB98F8EE" blockId="22.[151,676,1876,1899]" box="[151,676,1876,1899]" pageId="22" pageNumber="23">
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<emphasis id="B958EABFE311FF87FF75F8F1B86FF8EF" bold="true" box="[151,339,1876,1899]" pageId="22" pageNumber="23">FIGURES 12–13.</emphasis>
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<taxonomicName id="4C2C4D2EE311FF87FEB8F8F0B883F8EE" box="[346,447,1877,1899]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EABFE311FF87FEB8F8F0B883F8EE" box="[346,447,1877,1899]" italics="true" pageId="22" pageNumber="23">Hydraena</emphasis>
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</taxonomicName>
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, aedeagi of holotypes.
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</paragraph>
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</caption>
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<paragraph id="8B9336ADE311FF87FF25F836BCA1F876" blockId="22.[151,1437,1938,2035]" pageId="22" pageNumber="23">Frons and pronotum densely moderately coarsely punctate, clypeus much more finely and sparsely punctate than frons. Punctures of pronotum larger than those of head, each puncture with short recumbent seta; interstices varying from narrow ridges to ca. 0.5xpd, some punctures on disc confluent, interstices very weakly shining.</paragraph>
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<paragraph id="8B9336ADE310FF86FF75FF3DBA41FF50" blockId="23.[151,1437,151,824]" pageId="23" pageNumber="24">Mentum sparsely finely punctulate and moderately setose, dull; postmentum subrugulose, dull. Genae raised, weakly rugulose, dull, with posterior ridge, interrupted in midline.</paragraph>
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<paragraph id="8B9336ADE310FF86FF25FF45B9DCFEC5" blockId="23.[151,1437,151,824]" pageId="23" pageNumber="24">Pronotum widely arcuate laterally, slightly sinuate behind middle; anterior margin distinctly emarginate behind eyes, very weakly emarginate behind frons; PF1 and PF2 shallow; PF3 and PF4 deep, area between them weakly tumid.</paragraph>
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<paragraph id="8B9336ADE310FF86FF25FEE9BBB5FDB8" blockId="23.[151,1437,151,824]" pageId="23" pageNumber="24">
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Elytra arcuate laterally; summit of posterior declivity slightly before midlength; lateral explanate margins very wide, margin serrate, distance between serrations in posterior 0.5 ca. 2–
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<date id="FF92106DE310FF86FC2AFED5BADEFE0D" box="[968,994,368,392]" pageId="23" pageNumber="24">3x</date>
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those in anterior 0.5, widely serrate in posterior 0.5, row of punctures at base of explanate margin very large and very deep; serial punctures round, each with minute granule at anterior margin, separated by ca. 0.5–1xpd, slightly larger than largest pronotal punctures. Even numbered intervals 2–10 raised and granulate. Sutural interval slightly peaked over posterior declivity. Odd numbered intervals flat, ca. 1xpd, shining. Apices in dorsal aspect separately weakly rounded, in posterior aspect margins do not form angle with one another.
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</paragraph>
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<paragraph id="8B9336ADE310FF86FF25FDEDBA66FD75" blockId="23.[151,1437,151,824]" pageId="23" pageNumber="24">
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P1 and P2 very narrow, P1 slightly wider than P2. P1 median carina sinuate in profile. P2 almost parallel sided, apex pointed. Plaques very small, surface irregular, located at posterior sides of median depression. Metaventrite with very short point on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at slightly arcuate posterior margin ca.
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<date id="FF92106DE310FF86FDCDFD11BB77FD48" box="[559,587,692,717]" pageId="23" pageNumber="24">5x</date>
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P2. Legs short, stout. Male fifth abdominal ventrite with transversely raised glabrous ridge. Abdominal apex with large apicomedian notch.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E310FF86FF25FD59BB12FCBD" pageId="23" pageNumber="24" type="etymology">
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<paragraph id="8B9336ADE310FF86FF25FD59BB12FCBD" blockId="23.[151,1437,151,824]" pageId="23" pageNumber="24">
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<emphasis id="B958EABFE310FF86FF25FD59B872FC90" bold="true" box="[199,334,764,789]" pageId="23" pageNumber="24">Etymology.</emphasis>
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Named in reference to the large explanate margins of the elytra, which form shields over the "knees" of the middle and hind legs.
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |