243 lines
17 KiB
XML
243 lines
17 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6213078" ID-GBIF-Dataset="e04117e5-9306-489d-a0a3-9dbd97774f74" ID-GBIF-Taxon="119613327" ID-ISSN="1175-5326" ID-Zenodo-Dep="6213078" checkinTime="1461057042134" checkinUser="plazi" docAuthor="Hong, Yong & James, Samuel W." docDate="2009" docId="03858791FFEEFFF3FF67FCEDFB13F561" docLanguage="en" docName="zt02059p045.pdf" docOrigin="Zootaxa 2059" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@46f0bf0d" docStyleName="zootaxa.2007.monograph" docTitle="Pheretima camarinensis Hong & James, 2009, sp. nov." docType="treatment" docVersion="8" lastPageId="6" lastPageNumber="39" masterDocId="FFBCFFE9FFEBFFF5FFF0FF8AFFEBF41B" masterDocTitle="New earthworms of the Pheretima urceolata species group (Oligochaeta: Megascolecidae) from southern Luzon, Philippines" masterLastPageNumber="45" masterPageNumber="33" pageId="5" pageNumber="38" updateTime="1645486389336" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>New earthworms of the Pheretima urceolata species group (Oligochaeta: Megascolecidae) from southern Luzon, Philippines</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Hong, Yong</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>James, Samuel W.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2009</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>2059</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>33</mods:start>
|
||
<mods:end>45</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.5281/zenodo.186796</mods:identifier>
|
||
<mods:identifier type="GBIF-Dataset">e04117e5-9306-489d-a0a3-9dbd97774f74</mods:identifier>
|
||
<mods:identifier type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">186796</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6213078" ID-GBIF-Taxon="119613327" ID-Zenodo-Dep="6213078" LSID="urn:lsid:plazi:treatment:03858791FFEEFFF3FF67FCEDFB13F561" httpUri="http://treatment.plazi.org/id/03858791FFEEFFF3FF67FCEDFB13F561" lastPageId="6" lastPageNumber="39" pageId="5" pageNumber="38">
|
||
<subSubSection pageId="5" pageNumber="38" type="nomenclature">
|
||
<paragraph blockId="5.[151,545,871,932]" box="[151,545,871,897]" pageId="5" pageNumber="38">
|
||
<heading bold="true" box="[151,545,871,897]" fontSize="11" level="1" pageId="5" pageNumber="38" reason="1">
|
||
<emphasis bold="true" box="[151,545,871,897]" pageId="5" pageNumber="38">
|
||
<taxonomicName box="[151,445,871,897]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="38" phylum="Annelida" rank="species" species="camarinensis" status="sp. nov.">
|
||
<emphasis bold="true" box="[151,445,871,897]" italics="true" pageId="5" pageNumber="38">Pheretima camarinensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[452,545,871,897]" pageId="5" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,545,871,932]" box="[151,288,906,932]" pageId="5" pageNumber="38">
|
||
(
|
||
<figureCitation box="[159,224,906,932]" captionStart="FIGURE 1" captionStartId="2.[151,255,1523,1547]" captionTargetBox="[218,1357,245,1468]" captionTargetId="figure@2.[208,1365,237,1489]" captionTargetPageId="2" captionText="FIGURE 1. Pheretima bicolensis sp. nov. A: ventral view; B: spermatheca. Pheretima viracensis sp. nov. C: ventral view; D: spermatheca. Pheretima camarinensis sp. nov. E: ventral view; F: spermatheca. Scale bars = 2.5 mm (ventral views), 1 mm (spermathecae)." httpUri="https://zenodo.org/record/186797/files/figure.png" pageId="5" pageNumber="38">
|
||
Fig.
|
||
<number box="[214,224,906,932]" pageId="5" pageNumber="38" value="1.0">1</number>
|
||
</figureCitation>
|
||
E–F)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="38" type="materials_examined">
|
||
<paragraph blockId="5.[151,1437,986,2018]" pageId="5" pageNumber="38">
|
||
<emphasis bold="true" box="[151,267,986,1012]" pageId="5" pageNumber="38">Material.</emphasis>
|
||
<typeStatus box="[279,393,986,1012]" pageId="5" pageNumber="38">Holotype</typeStatus>
|
||
: Clitellate (NMA 004317):
|
||
<collectingCountry box="[736,871,986,1012]" name="Philippines" pageId="5" pageNumber="38">Philippines</collectingCountry>
|
||
, Camarines Sur, Isarog summit (
|
||
<geoCoordinate box="[1278,1432,986,1012]" direction="north" orientation="latitude" pageId="5" pageNumber="38" precision="9" value="13.6635">
|
||
<number box="[1278,1308,986,1012]" pageId="5" pageNumber="38" value="13.0">13</number>
|
||
°
|
||
<number box="[1332,1403,986,1012]" pageId="5" pageNumber="38" value="39.81">39.81</number>
|
||
’N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[151,311,1026,1052]" direction="east" orientation="longitude" pageId="5" pageNumber="38" precision="9" value="123.36483">
|
||
<number box="[151,195,1026,1052]" pageId="5" pageNumber="38" value="123.0">123</number>
|
||
°
|
||
<number box="[214,287,1026,1052]" pageId="5" pageNumber="38" value="21.89">21.89</number>
|
||
’E
|
||
</geoCoordinate>
|
||
),
|
||
<quantity box="[332,423,1026,1052]" metricMagnitude="3" metricUnit="m" metricValue="1.9869999999999997" pageId="5" pageNumber="38" unit="m" value="1987.0">1987 m</quantity>
|
||
, soil,
|
||
<date box="[491,650,1026,1052]" pageId="5" pageNumber="38" value="2001-05-15">
|
||
<number box="[491,520,1026,1052]" pageId="5" pageNumber="38" value="15.0">15</number>
|
||
May
|
||
<number box="[589,650,1026,1052]" pageId="5" pageNumber="38" value="2001.0">2001</number>
|
||
</date>
|
||
, S. W. James coll.
|
||
<number box="[869,884,1026,1052]" pageId="5" pageNumber="38" value="2.0">2</number>
|
||
<typeStatus box="[891,1008,1026,1052]" pageId="5" pageNumber="38">paratypes</typeStatus>
|
||
:
|
||
<number box="[1018,1032,1026,1052]" pageId="5" pageNumber="38" value="1.0">1</number>
|
||
clitellate (NMA 004322),
|
||
<number box="[1345,1360,1026,1052]" pageId="5" pageNumber="38" value="1.0">1</number>
|
||
clitellate (
|
||
<collectionCode box="[209,322,1066,1092]" pageId="5" pageNumber="38">KUNHM</collectionCode>
|
||
003013): Same data as for
|
||
<typeStatus box="[648,755,1066,1092]" pageId="5" pageNumber="38">holotype</typeStatus>
|
||
. Other material:
|
||
<number box="[953,968,1066,1092]" pageId="5" pageNumber="38" value="2.0">2</number>
|
||
clitellates,
|
||
<number box="[1103,1118,1066,1092]" pageId="5" pageNumber="38" value="5.0">5</number>
|
||
semi-clitellates, same data as for
|
||
<typeStatus box="[224,330,1106,1132]" pageId="5" pageNumber="38">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[197,830,1146,1172]" pageId="5" pageNumber="38" type="etymology">
|
||
<paragraph blockId="5.[151,1437,986,2018]" box="[197,830,1146,1172]" pageId="5" pageNumber="38">
|
||
<emphasis bold="true" box="[197,338,1146,1172]" pageId="5" pageNumber="38">Etymology.</emphasis>
|
||
The species is named for its
|
||
<typeStatus box="[679,730,1146,1172]" pageId="5" pageNumber="38">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="38" type="diagnosis">
|
||
<paragraph blockId="5.[151,1437,986,2018]" pageId="5" pageNumber="38">
|
||
<emphasis bold="true" box="[197,325,1186,1212]" pageId="5" pageNumber="38">Diagnosis.</emphasis>
|
||
<taxonomicName box="[333,444,1187,1212]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="38" phylum="Annelida" rank="species" species="urceolata">
|
||
<emphasis box="[333,444,1187,1212]" italics="true" pageId="5" pageNumber="38">urceolata</emphasis>
|
||
</taxonomicName>
|
||
-group with intestinal origin in XV, esophageal hearts in X–XIII, and two copulatory bursal pads on the sides of the chamber.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="6" lastPageNumber="39" pageId="5" pageNumber="38" type="description">
|
||
<paragraph blockId="5.[151,1437,986,2018]" pageId="5" pageNumber="38">
|
||
<emphasis bold="true" box="[197,349,1266,1292]" pageId="5" pageNumber="38">Description.</emphasis>
|
||
Light-brown dorsal pigment, setal rings unpigmented. Dimensions
|
||
<quantity box="[1147,1274,1266,1292]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="6.1" metricValueMin="5.4" pageId="5" pageNumber="38" unit="mm" value="57.5" valueMax="61.0" valueMin="54.0">54–61 mm</quantity>
|
||
by
|
||
<number box="[1319,1408,1266,1292]" pageId="5" pageNumber="38" value="3.05" valueMax="3.2" valueMin="2.9">2.9–3.2</number>
|
||
at segment X,
|
||
<quantity box="[295,437,1306,1332]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="5" pageNumber="38" unit="mm" value="2.45" valueMax="2.5" valueMin="2.4">2.4–2.5 mm</quantity>
|
||
at XXX,
|
||
<quantity box="[555,698,1306,1332]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="5" pageNumber="38" unit="mm" value="2.55" valueMax="2.6" valueMin="2.5">2.5–2.6 mm</quantity>
|
||
at clitellum, segments
|
||
<number box="[974,1066,1306,1332]" pageId="5" pageNumber="38" value="96.5" valueMax="102.0" valueMin="91.0">91–102</number>
|
||
; body circular in cross-section. Setae numbering
|
||
<number box="[356,429,1346,1372]" pageId="5" pageNumber="38" value="46.5" valueMax="49.0" valueMin="44.0">44–49</number>
|
||
at VII,
|
||
<number box="[520,593,1346,1372]" pageId="5" pageNumber="38" value="50.0" valueMax="52.0" valueMin="48.0">48–52</number>
|
||
at XX;
|
||
<number box="[687,731,1346,1372]" pageId="5" pageNumber="38" value="7.0" valueMax="9.0" valueMin="5.0">5–9</number>
|
||
between male pores, size and distance regular between dorsal and ventral, setal formula AA:AB:YZ:ZZ=
|
||
<number box="[719,733,1386,1412]" pageId="5" pageNumber="38" value="1.0">1</number>
|
||
:
|
||
<number box="[739,753,1386,1412]" pageId="5" pageNumber="38" value="1.0">1</number>
|
||
:
|
||
<number box="[759,773,1386,1412]" pageId="5" pageNumber="38" value="1.0">1</number>
|
||
:
|
||
<number box="[780,803,1386,1412]" pageId="5" pageNumber="38" value="2.0">2</number>
|
||
at XIII. Clitellum annular XIV–XVI; setae invisible externally.
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,986,2018]" pageId="5" pageNumber="38">
|
||
First dorsal pores
|
||
<number box="[406,437,1469,1495]" pageId="5" pageNumber="38" value="12.0">12</number>
|
||
/
|
||
<number box="[447,476,1469,1495]" pageId="5" pageNumber="38" value="13.0">13</number>
|
||
. One pair of spermathecal pores deep in
|
||
<number box="[949,962,1469,1495]" pageId="5" pageNumber="38" value="5.0">5</number>
|
||
/
|
||
<number box="[972,986,1469,1495]" pageId="5" pageNumber="38" value="6.0">6</number>
|
||
at
|
||
<number box="[1021,1050,1469,1495]" pageId="5" pageNumber="38" value="10.0">10</number>
|
||
th setal lines, sub-laterally placed,
|
||
<number box="[151,268,1509,1535]" pageId="5" pageNumber="38" value="0.19" valueMax="0.2" valueMin="0.18">0.18–0.20</number>
|
||
circumference apart, distance between spermathecal pores
|
||
<number box="[968,1003,1509,1535]" pageId="5" pageNumber="38" value="1.6">1.6</number>
|
||
–2.0 mm;
|
||
<quantity box="[1125,1216,1509,1535]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="38" unit="mm" value="0.4">0.4 mm</quantity>
|
||
openings of copulatory bursae paired in XVIII at
|
||
<number box="[568,597,1552,1578]" pageId="5" pageNumber="38" value="10.0">10</number>
|
||
th setal lines, ventro-lateral,
|
||
<number box="[946,1065,1552,1578]" pageId="5" pageNumber="38" value="0.195" valueMax="0.2" valueMin="0.19">0.19–0.20</number>
|
||
circumference apart; distance between opening
|
||
<quantity box="[357,502,1592,1618]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="5" pageNumber="38" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5–1.6 mm</quantity>
|
||
. Genital markings lacking.
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,986,2018]" pageId="5" pageNumber="38">
|
||
<collectingCountry box="[197,262,1632,1658]" name="Spain" pageId="5" pageNumber="38">Septa</collectingCountry>
|
||
<number box="[272,285,1632,1658]" pageId="5" pageNumber="38" value="5.0">5</number>
|
||
/
|
||
<number box="[295,339,1632,1658]" pageId="5" pageNumber="38" value="6.5" valueMax="7.0" valueMin="6.0">6–7</number>
|
||
/
|
||
<number box="[349,362,1632,1658]" pageId="5" pageNumber="38" value="8.0">8</number>
|
||
thin,
|
||
<number box="[434,447,1632,1658]" pageId="5" pageNumber="38" value="8.0">8</number>
|
||
/
|
||
<number box="[457,471,1632,1658]" pageId="5" pageNumber="38" value="9.0">9</number>
|
||
very thin muscular fiber,
|
||
<number box="[787,801,1632,1658]" pageId="5" pageNumber="38" value="9.0">9</number>
|
||
/
|
||
<number box="[811,839,1632,1658]" pageId="5" pageNumber="38" value="10.0">10</number>
|
||
absent
|
||
<date box="[933,1081,1632,1658]" pageId="5" pageNumber="38" value="1914-10-11" valueMax="1914-10-13" valueMin="1914-10-11">
|
||
<number box="[933,962,1632,1658]" pageId="5" pageNumber="38" value="10.0">10</number>
|
||
/
|
||
<number box="[972,1042,1632,1658]" pageId="5" pageNumber="38" value="12.0" valueMax="13.0" valueMin="11.0">11–13</number>
|
||
/
|
||
<number box="[1052,1081,1632,1658]" pageId="5" pageNumber="38" value="14.0">14</number>
|
||
</date>
|
||
slightly muscular. Gizzard in VIII–X, intestine begins in XV, medium paired lymph glands from XXVII along dorsal vessel; intestinal caeca simple originating in XXVII, and extending anteriorly about to XXIV, finger-shaped sac; typhlosole very low from XXVII. Hearts X–XIII esophageal.
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,986,2018]" lastBlockId="6.[151,1436,152,378]" lastPageId="6" lastPageNumber="39" pageId="5" pageNumber="38">
|
||
Ovaries and funnels in XIII, spermathecae paired in VI, with nephridia on spermatheca ducts; spermathecae with medium broad oval-shaped ampulla, duct narrower than ampulla; diverticulum ovate to globularshaped iridescent chamber, with stalk shorter than chamber. Male sexual system holandric, testes and funnels in ventral paired sacs in X–XI. Seminal vesicles two pairs in XI–XII, small dorsal lobes on upper edge, prostates in XVII–XVIII with
|
||
<number box="[471,486,1952,1978]" pageId="5" pageNumber="38" value="2.0">2</number>
|
||
primary lobes anterior lateral to copulatory bursae, with long muscular ducts, curve around anterior half in U-shape concave posteriorly, entering center via medial part of copulatory bursae without stalked glands; copulatory bursae opening flanked anterior, posterior by sucker-like circular pads more on sides than floor; each pad with associated glandular tissue; penis a small round knob from roof.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="39" type="discussion">
|
||
<paragraph blockId="6.[151,1436,152,378]" pageId="6" pageNumber="39">
|
||
<emphasis bold="true" box="[197,318,231,257]" pageId="6" pageNumber="39">Remarks.</emphasis>
|
||
The intestinal origin in XV and presence of esophageal hearts in X–XIII is unique among the species in the present paper, and shared only with
|
||
<emphasis box="[760,847,272,297]" italics="true" pageId="6" pageNumber="39">P. s i m s i</emphasis>
|
||
. Differences between the two species are that
|
||
<taxonomicName class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="39" phylum="Annelida" rank="species" species="simsi">
|
||
<emphasis italics="true" pageId="6" pageNumber="39">P. simsi</emphasis>
|
||
</taxonomicName>
|
||
has bursal pads on the floor rather than the sides, fewer setae in all regions of the body, and closer male pore spacing.
|
||
<taxonomicName box="[314,599,352,377]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="39" phylum="Annelida" rank="species" species="camarinensis" status="sp. nov.">
|
||
<emphasis box="[314,599,352,377]" italics="true" pageId="6" pageNumber="39">Pheretima camarinensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[606,700,351,377]" pageId="6" pageNumber="39">
|
||
<taxonomicNameLabel box="[606,700,351,377]" pageId="6" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is included in
|
||
<bibRefCitation author="James" box="[872,1029,352,378]" pageId="6" pageNumber="45" refString="James, S. W. (2005) Preliminary molecular phylogeny in the Pheretima group of genera (Crassiclitellata: Megascolecidae) using Bayesian analysis. In: V. V. Pop and A. A. Pop, (eds.), Advances in Earthworm Taxonomy II. Cluj University Press, Cluj-Napoca, Romania, pp. 129 - 142." type="book chapter" year="2005">
|
||
James (
|
||
<number box="[960,1022,352,378]" pageId="6" pageNumber="39" value="2005.0">2005</number>
|
||
)
|
||
</bibRefCitation>
|
||
as
|
||
<taxonomicName box="[1068,1189,352,377]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="39" phylum="Annelida" rank="genus">
|
||
<emphasis box="[1068,1189,352,377]" italics="true" pageId="6" pageNumber="39">Pheretima</emphasis>
|
||
</taxonomicName>
|
||
061-
|
||
<number box="[1253,1269,352,378]" pageId="6" pageNumber="39" value="4.0">4</number>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |