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<document id="6EAE1C65632F838A1CF94AECC965BBD5" ID="10.11646/zootaxa.4237.1.8SLASH10304" ID-CLB-Dataset="36418" ID-DOI="10.11646/zootaxa.4237.1.8" ID-GBIF-Dataset="f69cf65a-b415-426d-beeb-b21c1bd87bcc" ID-ISSN="1175-5326" ID-Zenodo-Dep="344551" ID-ZooBank="24A17254-7E24-41E3-ACA6-BC2795F674CF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1488189657570" checkinUser="plazi" docAuthor="Porch, Nick &amp; Smith, Tessa R." docDate="2017" docId="0385878CCE33FFFDC085FA72324BFE9B" docLanguage="en" docName="zootaxa.4237.1.8SLASH10304.pdf" docOrigin="Zootaxa 4237 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pycnomerus rimatara Porch &amp; Smith, 2017, sp. nov." docType="treatment" docUuid="D7D0AAB7-B592-432E-9F27-E4BCBCB19427" docUuidSource="ZooBank" docVersion="7" lastPageNumber="161" masterDocId="FFBCFFF4CE30FFF5C012FFC4313FFFDB" masterDocTitle="New Pycnomerus Erichson (Coleoptera: Zopheridae: Pycnomerini) from Rimatara, French Polynesia" masterLastPageNumber="166" masterPageNumber="154" pageNumber="157" updateTime="1698437369583" updateUser="ExternalLinkService">
<mods:mods id="8CB711090742E884D605FF1A07A5AC88" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="14A0B0B1DE7EA20A92F9FD79DC9E6966">
<mods:title id="3093BE4B1943B34C8AD76229E7710852">New Pycnomerus Erichson (Coleoptera: Zopheridae: Pycnomerini) from Rimatara, French Polynesia</mods:title>
</mods:titleInfo>
<mods:name id="F341C89FB05C91B4B3B9B45F1ADDD527" type="personal">
<mods:role id="2639B141D88EA9F4D75FB7739E047527">
<mods:roleTerm id="3120D980D2C056DDEB4547AD293F98AB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E7073FB33653A648DA79A7CD12BDC309">Porch, Nick</mods:namePart>
</mods:name>
<mods:name id="BB51165C414FB9AEA7D6975E50ACF5FA" type="personal">
<mods:role id="DBD445F5B084E94E4F71A55DD135BAE4">
<mods:roleTerm id="F236D96A1FCA11F7164875C1AA8B19DE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DAA38C94F8EE9ED807841BDFCFC88B49">Smith, Tessa R.</mods:namePart>
</mods:name>
<mods:typeOfResource id="726193358611F1DA36E43ED52964D64E">text</mods:typeOfResource>
<mods:relatedItem id="BCE3363FD26EC67F002078592A811BBB" type="host">
<mods:titleInfo id="0266147CA2D70C43E35A41A03E5601E2">
<mods:title id="3727AAD4F55C7EE8CEE9CD4AB36C1397">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EEC447C025F2F4419F12606E1C83E965">
<mods:date id="5B69B4B20C05F05D5C861846794CA18B">2017</mods:date>
<mods:detail id="5CC0AF87234E92234AB38D0DDB508075" type="volume">
<mods:number id="6ED2C533185953BE27326E0DA5B050A0">4237</mods:number>
</mods:detail>
<mods:detail id="5D523E57DF8E6B7413F805FAF8943E39" type="issue">
<mods:number id="FBFF9560D3B5D2C18607920F537677D9">1</mods:number>
</mods:detail>
<mods:extent id="C90E5D6237411ACA3AC17985FBA7D32B" unit="page">
<mods:start id="4AD297D7D9D33E8DA61DBE693EA8116A">154</mods:start>
<mods:end id="2E1536E591CC9461846D73E03A5124F7">166</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A6C5F0E5AFDFD9A13A5FA1546EC2067F">journal article</mods:classification>
<mods:identifier id="5907993C90EE14987676BD2F00A73D6B" type="CLB-Dataset">36418</mods:identifier>
<mods:identifier id="1CBBE9D4F8AFB214172BD5093C5A3C96" type="DOI">10.11646/zootaxa.4237.1.8</mods:identifier>
<mods:identifier id="41FE07AEEDE5D530B1558C809CC3197D" type="GBIF-Dataset">f69cf65a-b415-426d-beeb-b21c1bd87bcc</mods:identifier>
<mods:identifier id="995B146F671C492A32B55716532FF16F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6685CDE23543E671049FB4C20F618566" type="Zenodo-Dep">344551</mods:identifier>
<mods:identifier id="90F093B73F888536C32EC42C919A3293" type="ZooBank">24A17254-7E24-41E3-ACA6-BC2795F674CF</mods:identifier>
</mods:mods>
<treatment id="0385878CCE33FFFDC085FA72324BFE9B" ID-DOI="http://doi.org/10.5281/zenodo.6049837" ID-GBIF-Taxon="127711297" ID-Zenodo-Dep="6049837" LSID="urn:lsid:zoobank.org:act:D7D0AAB7-B592-432E-9F27-E4BCBCB19427" httpUri="http://treatment.plazi.org/id/0385878CCE33FFFDC085FA72324BFE9B" lastPageId="8" lastPageNumber="161" pageId="3" pageNumber="157">
<subSubSection id="C3366511CE33FFF6C085FA723031FA29" pageId="3" pageNumber="157" type="nomenclature">
<paragraph id="8B93369ACE33FFF6C085FA72333DFA0B" blockId="3.[151,514,1462,1522]" box="[151,514,1462,1488]" pageId="3" pageNumber="157">
<heading id="D0DB81F6CE33FFF6C085FA72333DFA0B" bold="true" box="[151,514,1462,1488]" fontSize="11" level="1" pageId="3" pageNumber="157" reason="1">
<emphasis id="B958EA88CE33FFF6C085FA72333DFA0B" bold="true" box="[151,514,1462,1488]" pageId="3" pageNumber="157">
<taxonomicName id="4C2C4D19CE33FFF6C085FA7230A2FA0B" box="[151,413,1462,1488]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="157" phylum="Arthropoda" rank="species" species="rimatara" status="sp. nov.">
<emphasis id="B958EA88CE33FFF6C085FA7230A2FA0B" bold="true" box="[151,413,1462,1488]" italics="true" pageId="3" pageNumber="157">Pycnomerus rimatara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B57F3CE33FFF6C1B6FA72333DFA0B" box="[420,514,1462,1488]" pageId="3" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B93369ACE33FFF6C085FA1D3031FA29" blockId="3.[151,514,1462,1522]" box="[151,270,1497,1522]" pageId="3" pageNumber="157">(Fig.2,3,4)</paragraph>
</subSubSection>
<subSubSection id="C3366511CE33FFF6C085F9E53557F881" pageId="3" pageNumber="157" type="materials_examined">
<paragraph id="8B93369ACE33FFF6C085F9E5327AF959" blockId="3.[151,1437,1569,2026]" pageId="3" pageNumber="157">
<materialsCitation id="3B443CC7CE33FFF6C085F9E5327AF959" ID-GBIF-Occurrence="1437756109" collectionCode="BPBM" collectorName="Maunutu Swamp, C" country="French Polynesia" location="Austral" pageId="3" pageNumber="157" specimenCount="2" typeStatus="holotype">
<emphasis id="B958EA88CE33FFF6C085F9E530F6F9E1" bold="true" box="[151,457,1569,1594]" pageId="3" pageNumber="157">
<typeStatus id="54978838CE33FFF6C085F9E531EDF9E1" box="[151,210,1569,1594]" pageId="3" pageNumber="157">Type</typeStatus>
material.
<typeStatus id="54978838CE33FFF6C14EF9E530F6F9E1" box="[348,457,1569,1594]" pageId="3" pageNumber="157" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="F33B760ACE33FFF6C1F2F9E53392F9E1" box="[480,685,1569,1594]" name="French Polynesia" pageId="3" pageNumber="157">French Polynesia</collectingCountry>
,
<location id="8EF36041CE33FFF6C2ADF9E5322DF9E1" LSID="urn:lsid:plazi:treatment:0385878CCE33FFFDC085FA72324BFE9B:8EF36041CE33FFF6C2ADF9E5322DF9E1" box="[703,786,1569,1594]" country="French Polynesia" name="Austral" pageId="3" pageNumber="157">Austral</location>
archipelago,
<taxonomicName id="4C2C4D19CE33FFF6C3A8F9E5351CF9E1" box="[954,1059,1569,1594]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="164" phylum="Arthropoda" rank="species" species="rimatara">Rimatara</taxonomicName>
,
<collectorName id="26D9534CCE33FFF6C425F9E63416F9E1" box="[1079,1321,1570,1594]" pageId="3" pageNumber="157">Maunutu Swamp, C</collectorName>
1T2 2008 excavation, 500 ml sediment sample,
<quantity id="4CD49B7FCE33FFF6C24BF98133D5F986" box="[601,746,1605,1630]" metricMagnitude="0" metricUnit="m" metricValue="1.025" metricValueMax="1.05" metricValueMin="1.0" pageId="3" pageNumber="157" unit="cm" value="102.5" valueMax="105.0" valueMin="100.0">100–105 cm</quantity>
depth—articulated male head and prothorax, dorsally and ventrally complete except for appendages.
<collectionCode id="ED3DAE5FCE33FFF6C26AF9AE33F6F95A" box="[632,713,1642,1665]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="3" pageNumber="157">BPBM</collectionCode>
, Honolulu.
</materialsCitation>
</paragraph>
<paragraph id="8B93369ACE33FFF6C0D5F9493557F881" blockId="3.[151,1437,1569,2026]" pageId="3" pageNumber="157">
<materialsCitation id="3B443CC7CE33FFF6C0D5F94934A8F911" ID-GBIF-Occurrence="1437756111" collectorName="Austral &amp; Maunutu Swamp C" country="French Polynesia" pageId="3" pageNumber="157" specimenCount="1" typeStatus="paratype">
<emphasis id="B958EA88CE33FFF6C0D5F9493059F97D" bold="true" box="[199,358,1677,1702]" pageId="3" pageNumber="157">
<typeStatus id="54978838CE33FFF6C0D5F9493000F97D" box="[199,319,1677,1702]" pageId="3" pageNumber="157" type="paratype">Paratypes</typeStatus>
(8)
</emphasis>
:
<collectingCountry id="F33B760ACE33FFF6C166F9493307F97D" box="[372,568,1677,1702]" name="French Polynesia" pageId="3" pageNumber="157">French Polynesia</collectingCountry>
,
<collectorName id="26D9534CCE33FFF6C251F94933AAF97D" box="[579,661,1677,1702]" pageId="3" pageNumber="157">Austral</collectorName>
archipelago,
<taxonomicName id="4C2C4D19CE33FFF6C33EF94932A9F97D" box="[812,918,1677,1702]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="164" phylum="Arthropoda" rank="species" species="rimatara">Rimatara</taxonomicName>
,
<collectorName id="26D9534CCE33FFF6C3B2F94A3545F97D" box="[928,1146,1678,1702]" pageId="3" pageNumber="157">Maunutu Swamp C</collectorName>
1T2 2008 excavation, 500 ml sediment sample,
<quantity id="4CD49B7FCE33FFF6C198F97530C4F912" box="[394,507,1713,1738]" metricMagnitude="-1" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="3" pageNumber="157" unit="cm" value="52.5" valueMax="55.0" valueMin="50.0">50–55 cm</quantity>
depth, 1 prothorax, 1 pronotum only;
<emphasis id="B958EA88CE33FFF6C3A6F97532D1F912" box="[948,1006,1713,1737]" italics="true" pageId="3" pageNumber="157">ditto,</emphasis>
except
<quantity id="4CD49B7FCE33FFF6C45BF9753585F912" box="[1097,1210,1713,1738]" metricMagnitude="-1" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="3" pageNumber="157" unit="cm" value="62.5" valueMax="65.0" valueMin="60.0">60–65 cm</quantity>
depth, 1 prothorax
</materialsCitation>
;
<materialsCitation id="3B443CC7CE33FFF6C085F911334EF935" ID-GBIF-Occurrence="1437756108" box="[151,625,1749,1774]" collectorName="Austral &amp; Maunutu Swamp C" country="French Polynesia" pageId="3" pageNumber="157" specimenCount="1" typeStatus="paratype">
<emphasis id="B958EA88CE33FFF6C085F91131EDF936" box="[151,210,1749,1773]" italics="true" pageId="3" pageNumber="157">ditto,</emphasis>
except
<quantity id="4CD49B7FCE33FFF6C13BF91130A7F936" box="[297,408,1749,1774]" metricMagnitude="-1" metricUnit="m" metricValue="7.25" metricValueMax="7.5" metricValueMin="7.0" pageId="3" pageNumber="157" unit="cm" value="72.5" valueMax="75.0" valueMin="70.0">70–75 cm</quantity>
depth, 1 prothorax
</materialsCitation>
;
<materialsCitation id="3B443CC7CE33FFF6C26FF911354AF935" ID-GBIF-Occurrence="1437756112" box="[637,1141,1749,1774]" collectorName="Austral &amp; Maunutu Swamp C" country="French Polynesia" pageId="3" pageNumber="157" specimenCount="1" typeStatus="paratype">
<emphasis id="B958EA88CE33FFF6C26FF9113386F936" box="[637,697,1749,1773]" italics="true" pageId="3" pageNumber="157">ditto,</emphasis>
except
<quantity id="4CD49B7FCE33FFF6C302F91132A4F936" box="[784,923,1749,1774]" metricMagnitude="0" metricUnit="m" metricValue="1.025" metricValueMax="1.05" metricValueMin="1.0" pageId="3" pageNumber="157" unit="cm" value="102.5" valueMax="105.0" valueMin="100.0">100–105 cm</quantity>
depth, 1 prothorax
</materialsCitation>
;
<materialsCitation id="3B443CC7CE33FFF6C492F91133D5F8ED" ID-GBIF-Occurrence="1437756120" collectionCode="MATO" collectorName="Austral &amp; Maunutu Swamp C" country="French Polynesia" location="Austral" municipality="Maunutu Swamp" pageId="3" pageNumber="157" specimenCode="MATO 2008" specimenCount="1" typeStatus="paratype">
<emphasis id="B958EA88CE33FFF6C492F9113584F936" box="[1152,1211,1749,1773]" italics="true" pageId="3" pageNumber="157">ditto,</emphasis>
except
<quantity id="4CD49B7FCE33FFF6C501F91134A3F936" box="[1299,1436,1749,1774]" metricMagnitude="0" metricUnit="m" metricValue="1.125" metricValueMax="1.15" metricValueMin="1.1" pageId="3" pageNumber="157" unit="cm" value="112.5" valueMax="115.0" valueMin="110.0">110–115 cm</quantity>
depth, articulated male head and prothorax;
<collectingCountry id="F33B760ACE33FFF6C284F93D3262F8C9" box="[662,861,1785,1810]" name="French Polynesia" pageId="3" pageNumber="157">French Polynesia</collectingCountry>
,
<location id="8EF36041CE33FFF6C379F93D3282F8C9" LSID="urn:lsid:plazi:treatment:0385878CCE33FFFDC085FA72324BFE9B:8EF36041CE33FFF6C379F93D3282F8C9" box="[875,957,1785,1810]" country="French Polynesia" municipality="Maunutu Swamp" name="Austral" pageId="3" pageNumber="157">Austral</location>
archipelago,
<taxonomicName id="4C2C4D19CE33FFF6C449F93D35FBF8C9" box="[1115,1220,1785,1810]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="164" phylum="Arthropoda" rank="species" species="rimatara">Rimatara</taxonomicName>
,
<collectingMunicipality id="6BF7ACE0CE33FFF6C4C0F93E34A7F8C9" box="[1234,1432,1786,1810]" pageId="3" pageNumber="157">Maunutu Swamp</collectingMunicipality>
,
<specimenCode id="DB8A9EE1CE33FFF6C085F8DA3016F8ED" box="[151,297,1822,1846]" collectionCode="MATO" pageId="3" pageNumber="157">
<collectionCode id="ED3DAE5FCE33FFF6C085F8DA31D9F8ED" box="[151,230,1822,1846]" pageId="3" pageNumber="157">MATO</collectionCode>
2008
</specimenCode>
core,
<quantity id="4CD49B7FCE33FFF6C164F8D9333AF8EE" box="[374,517,1821,1846]" metricMagnitude="0" metricUnit="m" metricValue="3.325" metricValueMax="3.35" metricValueMin="3.3" pageId="3" pageNumber="157" unit="cm" value="332.5" valueMax="335.0" valueMin="330.0">330–335 cm</quantity>
depth, 1 prothorax
</materialsCitation>
;
<materialsCitation id="3B443CC7CE33FFF6C2EBF8D93557F881" ID-GBIF-Occurrence="1437756114" collectionCode="MATO, BPBM" collectorName="Austral &amp; Maunutu Swamp C" country="French Polynesia" location="All" municipality="Maunutu Swamp" pageId="3" pageNumber="157" specimenCode="MATO 430-435" specimenCount="1" typeStatus="paratype">
<emphasis id="B958EA88CE33FFF6C2EBF8D9320BF8EE" box="[761,820,1821,1845]" italics="true" pageId="3" pageNumber="157">ditto,</emphasis>
except
<quantity id="4CD49B7FCE33FFF6C380F8D9351EF8EE" box="[914,1057,1821,1846]" metricMagnitude="0" metricUnit="m" metricValue="4.325" metricValueMax="4.35" metricValueMin="4.3" pageId="3" pageNumber="157" unit="cm" value="432.5" valueMax="435.0" valueMin="430.0">430–435 cm</quantity>
depth, 1 prothorax.
<location id="8EF36041CE33FFF6C506F8D93407F8ED" LSID="urn:lsid:plazi:treatment:0385878CCE33FFFDC085FA72324BFE9B:8EF36041CE33FFF6C506F8D93407F8ED" box="[1300,1336,1821,1846]" name="All" pageId="3" pageNumber="157">All</location>
but two specimens (
<specimenCode id="DB8A9EE1CE33FFF6C108F88630ECF881" box="[282,467,1857,1882]" collectionCode="MATO" pageId="3" pageNumber="157">
<collectionCode id="ED3DAE5FCE33FFF6C108F8863054F881" box="[282,363,1858,1882]" pageId="3" pageNumber="157">MATO</collectionCode>
<quantity id="4CD49B7FCE33FFF6C160F88530ECF881" box="[370,467,1857,1882]" metricMagnitude="0" metricUnit="m" metricValue="4.325" metricValueMax="4.35" metricValueMin="4.3" pageId="3" pageNumber="157" unit="cm" value="432.5" valueMax="435.0" valueMin="430.0">430–435</quantity>
</specimenCode>
cm and C1T2
<quantity id="4CD49B7FCE33FFF6C268F8853238F881" box="[634,775,1857,1882]" metricMagnitude="0" metricUnit="m" metricValue="1.025" metricValueMax="1.05" metricValueMin="1.0" pageId="3" pageNumber="157" unit="cm" value="102.5" valueMax="105.0" valueMin="100.0">100–105 cm</quantity>
) in
<collectionCode id="ED3DAE5FCE33FFF6C320F88632BCF882" box="[818,899,1858,1881]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="3" pageNumber="157">BPBM</collectionCode>
, otherwise in Porch.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3366511CE33FFF0C0D5F8A132BDF91A" lastPageId="5" lastPageNumber="159" pageId="3" pageNumber="157" type="diagnosis">
<paragraph id="8B93369ACE33FFF0C0D5F8A132BDF91A" blockId="3.[151,1437,1569,2026]" lastBlockId="5.[151,1437,1561,2018]" lastPageId="5" lastPageNumber="159" pageId="3" pageNumber="157">
<emphasis id="B958EA88CE33FFF6C0D5F8A1307EF8A5" bold="true" box="[199,321,1893,1918]" pageId="3" pageNumber="157">Diagnosis.</emphasis>
This species is distinguished from the known co-occurring species,
<emphasis id="B958EA88CE33FFF6C44EF8A2341FF8A6" box="[1116,1312,1893,1917]" italics="true" pageId="3" pageNumber="157">
<taxonomicName id="4C2C4D19CE33FFF6C44EF8A235E8F8A6" box="[1116,1239,1893,1917]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="157" phylum="Arthropoda" rank="species" species="prebblei" status="sp. nov.">P. prebblei</taxonomicName>
<taxonomicNameLabel id="A26B57F3CE33FFF6C4F3F8A2341FF8A6" box="[1249,1312,1894,1917]" pageId="3" pageNumber="157" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, described herein, by differing physical dimensions, shape and microsculpture.
<taxonomicName id="4C2C4D19CE33FFF6C389F84E3523F87A" box="[923,1052,1930,1953]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="157" phylum="Arthropoda" rank="species" species="rimatara">
<emphasis id="B958EA88CE33FFF6C389F84E3523F87A" box="[923,1052,1930,1953]" italics="true" pageId="3" pageNumber="157">P. rimatara</emphasis>
</taxonomicName>
has a flattened pronotal disk with bilinear arrangement of large elongate punctures either side of the middle of the disk, in contrast to
<taxonomicName id="4C2C4D19CE33FFF6C509F86A34AAF81E" box="[1307,1429,1965,1989]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="157" phylum="Arthropoda" rank="species" species="prebblei">
<emphasis id="B958EA88CE33FFF6C509F86A34AAF81E" box="[1307,1429,1965,1989]" italics="true" pageId="3" pageNumber="157">P. prebblei</emphasis>
</taxonomicName>
, which has a convex pronotal disk with evenly distributed, evenly sized punctures. The posterior angles of the pronotum are obtuse in
<taxonomicName id="4C2C4D19CE35FFF0C1B4F9DE3318F9EA" box="[422,551,1562,1585]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="159" phylum="Arthropoda" rank="species" species="rimatara">
<emphasis id="B958EA88CE35FFF0C1B4F9DE3318F9EA" box="[422,551,1562,1585]" italics="true" pageId="5" pageNumber="159">P. rimatara</emphasis>
</taxonomicName>
and rounded in
<taxonomicName id="4C2C4D19CE35FFF0C2F0F9DE3264F9E9" box="[738,859,1562,1586]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="159" phylum="Arthropoda" rank="species" species="prebblei">
<emphasis id="B958EA88CE35FFF0C2F0F9DE3264F9E9" box="[738,859,1562,1586]" italics="true" pageId="5" pageNumber="159">P. prebblei</emphasis>
</taxonomicName>
and the sides of the pronotum distinctly anteriorly divergent in the former species and subparallel in
<taxonomicName id="4C2C4D19CE35FFF0C2DAF9FA327FF98E" box="[712,832,1597,1621]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="159" phylum="Arthropoda" rank="species" species="prebblei">
<emphasis id="B958EA88CE35FFF0C2DAF9FA327FF98E" box="[712,832,1597,1621]" italics="true" pageId="5" pageNumber="159">P. prebblei</emphasis>
</taxonomicName>
. This latter character is evident in the ratio of median width relative to maximum width of the pronotum:
<taxonomicName id="4C2C4D19CE35FFF0C2E0F9A6324BF9A2" box="[754,884,1634,1657]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="159" phylum="Arthropoda" rank="species" species="rimatara">
<emphasis id="B958EA88CE35FFF0C2E0F9A6324BF9A2" box="[754,884,1634,1657]" italics="true" pageId="5" pageNumber="159">P. rimatara</emphasis>
</taxonomicName>
&lt;0.97,
<taxonomicName id="4C2C4D19CE35FFF0C3C2F9A63574F9A1" box="[976,1099,1634,1658]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="159" phylum="Arthropoda" rank="species" species="prebblei">
<emphasis id="B958EA88CE35FFF0C3C2F9A63574F9A1" box="[976,1099,1634,1658]" italics="true" pageId="5" pageNumber="159">P. prebblei</emphasis>
</taxonomicName>
&gt;0.97. The anterior pronotal angles in
<taxonomicName id="4C2C4D19CE35FFF0C116F94230BBF946" box="[260,388,1670,1693]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="159" phylum="Arthropoda" rank="species" species="rimatara">
<emphasis id="B958EA88CE35FFF0C116F94230BBF946" box="[260,388,1670,1693]" italics="true" pageId="5" pageNumber="159">P. rimatara</emphasis>
</taxonomicName>
are not distinctively produced, whereas those of
<taxonomicName id="4C2C4D19CE35FFF0C3A0F9423515F946" box="[946,1066,1669,1693]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="159" phylum="Arthropoda" rank="species" species="prebblei">
<emphasis id="B958EA88CE35FFF0C3A0F9423515F946" box="[946,1066,1669,1693]" italics="true" pageId="5" pageNumber="159">P. prebblei</emphasis>
</taxonomicName>
are obtusely produced to almost the level of the anterior margin of the medial part of the pronotum.
</paragraph>
</subSubSection>
<caption id="DF536612CE34FFF1C085F841357EF803" httpUri="https://zenodo.org/record/344553/files/figure.png" pageId="4" pageNumber="158" targetBox="[224,1353,214,1862]" targetPageId="4">
<paragraph id="8B93369ACE34FFF1C085F841357EF803" blockId="4.[151,1436,1925,2008]" pageId="4" pageNumber="158">
<emphasis id="B958EA88CE34FFF1C085F8413029F841" bold="true" box="[151,278,1925,1947]" pageId="4" pageNumber="158">FIGURE 2.</emphasis>
Holotypes. Top:
<taxonomicName id="4C2C4D19CE34FFF1C1DCF8423390F840" box="[462,687,1926,1947]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="158" phylum="Arthropoda" rank="species" species="rimatara">
<emphasis id="B958EA88CE34FFF1C1DCF8423390F840" box="[462,687,1926,1947]" italics="true" pageId="4" pageNumber="158">Pycnomerus rimatara</emphasis>
</taxonomicName>
articulated male prothorax and head, dorsal (left) and ventral (right)— (
<taxonomicName id="4C2C4D19CE34FFF1C08FF86031C3F861" box="[157,252,1956,1978]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="164" phylum="Arthropoda" rank="species" species="rimatara">Rimatara</taxonomicName>
C1T2 excavation, 250 ml sediment sample, 100–105 cm depth). Bottom:
<taxonomicName id="4C2C4D19CE34FFF1C41BF86135DBF861" box="[1033,1252,1956,1978]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="158" phylum="Arthropoda" rank="species" species="prebblei">
<emphasis id="B958EA88CE34FFF1C41BF86135DBF861" box="[1033,1252,1956,1978]" italics="true" pageId="4" pageNumber="158">Pycnomerus prebblei</emphasis>
</taxonomicName>
prothorax, dorsal (left) and ventral (right)—(
<taxonomicName id="4C2C4D19CE34FFF1C1BAF8063336F803" box="[424,521,1986,2008]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="164" phylum="Arthropoda" rank="species" species="rimatara">Rimatara</taxonomicName>
, MATO core, 430–435 cm depth). Scale bars = 0.5 mm.
</paragraph>
</caption>
<caption id="DF536612CE35FFF0C085FA793204FA29" httpUri="https://zenodo.org/record/344554/files/figure.png" pageId="5" pageNumber="159" targetBox="[159,1433,198,1438]" targetPageId="5">
<paragraph id="8B93369ACE35FFF0C085FA793204FA29" blockId="5.[151,1436,1469,1522]" pageId="5" pageNumber="159">
<emphasis id="B958EA88CE35FFF0C085FA79302CFA08" bold="true" box="[151,275,1469,1491]" pageId="5" pageNumber="159">FIGURE 3.</emphasis>
Dorsal and ventral views of well-preserved heads and prothoraces of
<taxonomicName id="4C2C4D19CE35FFF0C3F1FA7B3569FA0F" box="[995,1110,1471,1492]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="159" phylum="Arthropoda" rank="species" species="rimatara">
<emphasis id="B958EA88CE35FFF0C3F1FA7B3569FA0F" box="[995,1110,1471,1492]" italics="true" pageId="5" pageNumber="159">P. rimatara</emphasis>
</taxonomicName>
. Holotype from C1T2 100–105 for comparison. Prothoraces are all paratypes. Scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C3366511CE35FFF2C0D5F90932C0FBA4" lastPageId="7" lastPageNumber="161" pageId="5" pageNumber="159" type="description">
<paragraph id="8B93369ACE35FFF0C0D5F909339BF93D" blockId="5.[151,1437,1561,2018]" box="[199,676,1741,1766]" pageId="5" pageNumber="159">
<emphasis id="B958EA88CE35FFF0C0D5F909308CF93D" bold="true" box="[199,435,1741,1766]" pageId="5" pageNumber="159">Description. Colour</emphasis>
: Dark piceous-black.
</paragraph>
<paragraph id="8B93369ACE35FFF3C0D5F93530A1FF0E" blockId="5.[151,1437,1561,2018]" lastBlockId="6.[151,1436,151,464]" lastPageId="6" lastPageNumber="160" pageId="5" pageNumber="159">
<emphasis id="B958EA88CE35FFF0C0D5F935307DF8D1" bold="true" box="[199,322,1777,1802]" pageId="5" pageNumber="159">Prothorax</emphasis>
approximately as wide as long, greatest width at 1/4–1/5 of the prothorax length. Lateral margins straight, weakly converging behind widest point until rounded close to base, weakly-moderately crenulate and with narrow marginal sulcus adjacent to narrow, raised, border. Anterior angles weakly rounded and slightly produced behind middle. Anterior margin straight to very weakly bisinuate and unbordered except at anterior angles where sulcus of lateral margin continues around the anterior angle sinuously, a row of small punctures medially. Posterior angles obtuse. Posterior margin weakly lobed, rounded to very obtuse angulate, completely margined by sulcus, a continuation of lateral sulcus. Pair of deep fovae in basal sulcus, either side of middle, extending just to base of pronotum and merging with punctures which crenulate it. Basal border narrow, punctate, appearing thicker adjacent to basal fovae.
</paragraph>
<paragraph id="8B93369ACE36FFF3C0D5FF1B35ACFE0B" blockId="6.[151,1436,151,464]" pageId="6" pageNumber="160">
<emphasis id="B958EA88CE36FFF3C0D5FF1B3057FF23" bold="true" box="[199,360,223,248]" pageId="6" pageNumber="160">Pronotal disk</emphasis>
moderately flat over medial third, a little declivous to anterior margin, and curving to anterior and posterior lateral margins in anterior and posterior thirds, respectively. Disk indistinctly impressed either side of middle and bearing two diffuse fields of oval punctures, either side of middle, that are separated by their own, to several times their own, length. Punctures in basal third diverging and laterally oriented, becoming convergent and medially oriented in middle third, becoming smaller in size towards anterior margin. Punctures mainly parallel but convergent in anterior third. Microsculpture of pronotal disk weak, pronotal disk between puncture appearing shiny but with a reduced longitudinally elongate mesh which gives an impression of fingerprints.
</paragraph>
<caption id="DF536612CE36FFF3C085F8C8300EF8BB" httpUri="https://zenodo.org/record/344555/files/figure.png" pageId="6" pageNumber="160" targetBox="[172,1395,527,1773]" targetPageId="6">
<paragraph id="8B93369ACE36FFF3C085F8C8300EF8BB" blockId="6.[151,1436,1804,1888]" pageId="6" pageNumber="160">
<emphasis id="B958EA88CE36FFF3C085F8C83029F8FA" bold="true" box="[151,278,1804,1826]" pageId="6" pageNumber="160">FIGURE 4.</emphasis>
Dorsal views of complete and near complete elytra considered to belong (not types) to
<taxonomicName id="4C2C4D19CE36FFF3C4AFF8C9340BF8F9" box="[1213,1332,1805,1826]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="160" phylum="Arthropoda" rank="species" species="rimatara">
<emphasis id="B958EA88CE36FFF3C4AFF8C9340BF8F9" box="[1213,1332,1805,1826]" italics="true" pageId="6" pageNumber="160">P. rimatara</emphasis>
</taxonomicName>
. A single specimen (right) from MATO 355–360 shown magnified relative to other specimens to illustrate elytral microsculpture. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B93369ACE36FFF2C0D5F84832F1FF2C" blockId="6.[151,1436,1932,2029]" lastBlockId="7.[151,1437,151,2015]" lastPageId="7" lastPageNumber="161" pageId="6" pageNumber="160">
<emphasis id="B958EA88CE36FFF3C0D5F848306AF87E" bold="true" box="[199,341,1932,1957]" pageId="6" pageNumber="160">Prosternum</emphasis>
densely punctate with punctures usually separated by about their length or less, closer on middle in front coxae and on middle and apex of prosternal process where they begin to coalesce. Punctures smaller in middle of prosternum and on prosternal process, larger on sides of prosternum. Anterior of prosternum with moderately well-developed margin that extends across much of prosternum, composed of many small punctures and a shallow punctured sulcus; with a well-developed fringe of golden setae remaining on some specimens. Apex of prosternal process truncate to slightly emarginate. Hypomera punctate.
</paragraph>
<paragraph id="8B93369ACE37FFF2C0D5FEC035D8FDE4" blockId="7.[151,1437,151,2015]" pageId="7" pageNumber="161">
<emphasis id="B958EA88CE37FFF2C0D5FEC03038FEC6" bold="true" box="[199,263,260,285]" pageId="7" pageNumber="161">Head</emphasis>
(based on
<typeStatus id="54978838CE37FFF2C190FEC030DBFEC6" box="[386,484,260,285]" pageId="7" pageNumber="161" type="holotype">holotype</typeStatus>
and a
<typeStatus id="54978838CE37FFF2C23DFEC133B0FEC6" box="[559,655,261,285]" pageId="7" pageNumber="161" type="paratype">paratype</typeStatus>
still articulated with prothorax) widened in front of eyes until widest 1/3 eye length in front of eyes. Frontal margins very weakly bisinuate, narrowed (at approx. 50o angle) to moderately wide anterior margin which is weakly bisinuate with greatest development on right side. Longitudinally elongate depressions of deep confluent punctures adjacent to each eye, from just before middle of eye to just before hind margin of eye, deeper rounded to elongate (varies across specimens) depressions on each side of frons at termination of weakly indicated frontoclypeal suture. Frontoclypeal suture extends from margin of clypeus to fovae, absent across middle of frons between fovae. Head shining with punctures separated by their diameter to several times their diameter. Eye coarsely facetted, about 7–8 ommatidia in length along lateral margin, dorsally and distinctly longer ventrally (estimated 10–11 ommatidia). Males with setose pit on submentum.
</paragraph>
<paragraph id="8B93369ACE37FFF2C0D5FD8E3301FDB8" blockId="7.[151,1437,151,2015]" box="[199,574,586,611]" pageId="7" pageNumber="161">
<emphasis id="B958EA88CE37FFF2C0D5FD8E3301FDB8" bold="true" box="[199,574,586,611]" pageId="7" pageNumber="161">Physical dimensions and Ratios</emphasis>
</paragraph>
<paragraph id="8B93369ACE37FFF2C0D5FDAA346CFD28" blockId="7.[151,1437,151,2015]" pageId="7" pageNumber="161">
<emphasis id="B958EA88CE37FFF2C0D5FDAA307DFD5C" bold="true" box="[199,322,622,647]" pageId="7" pageNumber="161">Prothorax</emphasis>
measurements based on
<specimenCount id="9D2AFD13CE37FFF2C248FDB433D8FD5C" box="[602,743,622,647]" pageId="7" pageNumber="161" type="generic">7 specimens</specimenCount>
. Measurements of
<typeStatus id="54978838CE37FFF2C3A9FDAA351CFD5C" box="[955,1059,622,647]" pageId="7" pageNumber="161" type="holotype">Holotype</typeStatus>
in parentheses. Pronotum length:
<quantity id="4CD49B7FCE37FFF2C085FD563006FD71" box="[151,313,658,683]" metricMagnitude="-4" metricUnit="m" metricValue="8.35" metricValueMax="9.5" metricValueMin="7.199999999999999" pageId="7" pageNumber="161" unit="mm" value="0.835" valueMax="0.95" valueMin="0.72">0.72–0.95 mm</quantity>
(
<quantity id="4CD49B7FCE37FFF2C15AFD57308EFD70" box="[328,433,659,683]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="7" pageNumber="161" unit="mm" value="0.87">0.87 mm</quantity>
). Pronotum width:
<quantity id="4CD49B7FCE37FFF2C295FD563215FD71" box="[647,810,658,683]" metricMagnitude="-4" metricUnit="m" metricValue="8.55" metricValueMax="9.6" metricValueMin="7.5" pageId="7" pageNumber="161" unit="mm" value="0.855" valueMax="0.96" valueMin="0.75">0.75–0.96 mm</quantity>
(
<quantity id="4CD49B7FCE37FFF2C32AFD56329EFD70" box="[824,929,658,683]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="7" pageNumber="161" unit="mm" value="0.85">0.85 mm</quantity>
). Anterior width of pronotum:
<quantity id="4CD49B7FCE37FFF2C4E8FD5634A2FD71" box="[1274,1437,658,683]" metricMagnitude="-4" metricUnit="m" metricValue="6.85" metricValueMax="7.7" metricValueMin="6.0" pageId="7" pageNumber="161" unit="mm" value="0.685" valueMax="0.77" valueMin="0.6">0.60–0.77 mm</quantity>
(
<quantity id="4CD49B7FCE37FFF2C08DFD733036FD14" box="[159,265,695,719]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="7" pageNumber="161" unit="mm" value="0.71">0.71 mm</quantity>
). Medial width of pronotum:
<quantity id="4CD49B7FCE37FFF2C24BFD7333C2FD14" box="[601,765,695,719]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" metricValueMax="9.3" metricValueMin="7.1" pageId="7" pageNumber="161" unit="mm" value="0.82" valueMax="0.93" valueMin="0.71">0.71–0.93 mm</quantity>
(
<quantity id="4CD49B7FCE37FFF2C31FFD733247FD14" box="[781,888,695,719]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="7" pageNumber="161" unit="mm" value="0.82">0.82 mm</quantity>
). Basal width of pronotum:
<quantity id="4CD49B7FCE37FFF2C4A4FD723465FD14" box="[1206,1370,694,719]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" metricValueMax="7.8" metricValueMin="6.0" pageId="7" pageNumber="161" unit="mm" value="0.69" valueMax="0.78" valueMin="0.6">0.60–0.78 mm</quantity>
(
<quantity id="4CD49B7FCE37FFF2C578FD7231F8FD28" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="7" pageNumber="161" unit="mm" value="0.64">0.64 mm</quantity>
). Lateral length of pronotum:
<quantity id="4CD49B7FCE37FFF2C205FD1E3385FD29" box="[535,698,730,755]" metricMagnitude="-4" metricUnit="m" metricValue="6.75" metricValueMax="7.8" metricValueMin="5.699999999999999" pageId="7" pageNumber="161" unit="mm" value="0.675" valueMax="0.78" valueMin="0.57">0.57–0.78 mm</quantity>
(
<quantity id="4CD49B7FCE37FFF2C2DAFD1F320DFD28" box="[712,818,731,755]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="7" pageNumber="161" unit="mm" value="0.74">0.74 mm</quantity>
). Pronotum length/width ratio: 0.96–1.04 (1.02).
</paragraph>
<paragraph id="8B93369ACE37FFF2C0D5FD3A351BFC58" blockId="7.[151,1437,151,2015]" pageId="7" pageNumber="161">
Ventral Prothorax: Prosternum length:
<quantity id="4CD49B7FCE37FFF2C26CFD3A321DFCCC" box="[638,802,766,791]" metricMagnitude="-4" metricUnit="m" metricValue="5.85" metricValueMax="6.7" metricValueMin="5.0" pageId="7" pageNumber="161" unit="mm" value="0.585" valueMax="0.67" valueMin="0.5">0.50–0.67 mm</quantity>
(
<quantity id="4CD49B7FCE37FFF2C323FD3A32A3FCCC" box="[817,924,766,791]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="7" pageNumber="161" unit="mm" value="0.62">0.62 mm</quantity>
). Length prosternum in front of coxae:
<quantity id="4CD49B7FCE37FFF2C54FFD3B31C0FCE1" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="4.2" metricValueMin="3.0" pageId="7" pageNumber="161" unit="mm" value="0.36" valueMax="0.42" valueMin="0.3">0.30– 0.42 mm</quantity>
(
<quantity id="4CD49B7FCE37FFF2C103FCE630BFFCE0" box="[273,384,802,827]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="7" pageNumber="161" unit="mm" value="0.36">0.36 mm</quantity>
). Anterior foramen:
<quantity id="4CD49B7FCE37FFF2C263FCE63227FCE1" box="[625,792,802,827]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.1" metricValueMin="4.9" pageId="7" pageNumber="161" unit="mm" value="0.55" valueMax="0.61" valueMin="0.49">0.49–0.61 mm</quantity>
(
<quantity id="4CD49B7FCE37FFF2C338FCE632A7FCE0" box="[810,920,802,827]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="7" pageNumber="161" unit="mm" value="0.57">0.57 mm</quantity>
). Basal foramen:
<quantity id="4CD49B7FCE37FFF2C47BFCE6342FFCE1" box="[1129,1296,802,827]" metricMagnitude="-4" metricUnit="m" metricValue="5.549999999999999" metricValueMax="6.4" metricValueMin="4.699999999999999" pageId="7" pageNumber="161" unit="mm" value="0.5549999999999999" valueMax="0.64" valueMin="0.47">0.47–0.64 mm</quantity>
(
<quantity id="4CD49B7FCE37FFF2C530FCE634AFFCE0" box="[1314,1424,802,827]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="7" pageNumber="161" unit="mm" value="0.56">0.56 mm</quantity>
). Prosternal process intercoxal width:
<quantity id="4CD49B7FCE37FFF2C222FC8333ECFC84" box="[560,723,839,863]" metricMagnitude="-4" metricUnit="m" metricValue="1.95" metricValueMax="2.2" metricValueMin="1.7" pageId="7" pageNumber="161" unit="mm" value="0.195" valueMax="0.22" valueMin="0.17">0.17–0.22 mm</quantity>
(
<quantity id="4CD49B7FCE37FFF2C2F3FC833274FC84" box="[737,843,839,863]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="161" unit="mm" value="0.19">0.19 mm</quantity>
). Maximum prosternal process width:
<quantity id="4CD49B7FCE37FFF2C4E8FC8234A2FC84" box="[1274,1437,838,863]" metricMagnitude="-4" metricUnit="m" metricValue="2.95" metricValueMax="3.4" metricValueMin="2.5" pageId="7" pageNumber="161" unit="mm" value="0.295" valueMax="0.34" valueMin="0.25">0.25–0.34 mm</quantity>
(
<quantity id="4CD49B7FCE37FFF2C08CFCAF3037FC58" box="[158,264,875,899]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="161" unit="mm" value="0.29">0.29 mm</quantity>
). Ratio of Prosternal length/Length in front of coxae: 1.58–1.70 (1.70).
</paragraph>
<paragraph id="8B93369ACE37FFF2C0D5FC4A32C0FBA4" blockId="7.[151,1437,151,2015]" pageId="7" pageNumber="161">
<emphasis id="B958EA88CE37FFF2C0D5FC4A3038FC7C" bold="true" box="[199,263,910,935]" pageId="7" pageNumber="161">Head</emphasis>
measurements based on
<specimenCount id="9D2AFD13CE37FFF2C230FC4A3392FC7C" box="[546,685,910,935]" pageId="7" pageNumber="161" type="generic">5 specimens</specimenCount>
including
<specimenCount id="9D2AFD13CE37FFF2C335FC4B3247FC73" box="[807,888,911,936]" pageId="7" pageNumber="161" type="generic" typeStatus="types">2 types</specimenCount>
(articulated heads and prothoraces listed above) and 3 additional specimens considered to belong to this species noted below in ‘additional material’. Some measurements not possible for all specimens; noted as ‘NA’ for
<typeStatus id="54978838CE37FFF2C3C7FC123508FC34" box="[981,1079,982,1007]" pageId="7" pageNumber="161" type="holotype">holotype</typeStatus>
measurement if unavailable. Measurements of
<typeStatus id="54978838CE37FFF2C174FC3E30F1FBC8" box="[358,462,1018,1043]" pageId="7" pageNumber="161" type="holotype">Holotype</typeStatus>
in parentheses. Clypeus width:
<quantity id="4CD49B7FCE37FFF2C355FC3E32E7FBC9" box="[839,984,1018,1043]" metricMagnitude="-2" metricUnit="m" metricValue="3.0255" metricValueMax="6.0" metricValueMin="0.051" pageId="7" pageNumber="161" unit="mm" value="30.255" valueMax="60.0" valueMin="0.51">0.51–60 mm</quantity>
(
<quantity id="4CD49B7FCE37FFF2C3F8FC3E3567FBC8" box="[1002,1112,1018,1043]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="7" pageNumber="161" unit="mm" value="0.56">0.56 mm</quantity>
). Interocular width (dorsal):
<quantity id="4CD49B7FCE37FFF2C085FBDB3004FBEC" box="[151,315,1055,1079]" metricMagnitude="-4" metricUnit="m" metricValue="4.449999999999999" metricValueMax="4.8" metricValueMin="4.1" pageId="7" pageNumber="161" unit="mm" value="0.44499999999999995" valueMax="0.48" valueMin="0.41">0.41–0.48 mm</quantity>
(
<quantity id="4CD49B7FCE37FFF2C159FBDA3089FBEC" box="[331,438,1054,1079]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="161" unit="mm" value="0.45">0.45 mm</quantity>
). Temporal width:
<quantity id="4CD49B7FCE37FFF2C29DFBDA320BFBEC" box="[655,820,1054,1079]" metricMagnitude="-4" metricUnit="m" metricValue="5.549999999999999" metricValueMax="5.8" metricValueMin="5.3" pageId="7" pageNumber="161" unit="mm" value="0.5549999999999999" valueMax="0.58" valueMin="0.53">0.53–0.58 mm</quantity>
(NA). Eye length (dorsal):
<quantity id="4CD49B7FCE37FFF2C463FBDA342AFBEC" box="[1137,1301,1054,1079]" metricMagnitude="-4" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="7" pageNumber="161" unit="mm" value="0.145" valueMax="0.16" valueMin="0.13">0.13–0.16 mm</quantity>
(
<quantity id="4CD49B7FCE37FFF2C537FBDA34AFFBEC" box="[1317,1424,1054,1079]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="161" unit="mm" value="0.15">0.15 mm</quantity>
). Eye length (ventral):
<quantity id="4CD49B7FCE37FFF2C19EFB87330EFB81" box="[396,561,1091,1115]" metricMagnitude="-4" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="7" pageNumber="161" unit="mm" value="0.195" valueMax="0.2" valueMin="0.19">0.19–0.20 mm</quantity>
(
<quantity id="4CD49B7FCE37FFF2C250FB8733A0FB80" box="[578,671,1091,1115]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="161" unit="mm" value="0.2">0.2 mm</quantity>
). Ratio Eye length dorsally/Eye length ventrally: 1.09–1.14 (1.10). Ratio interocular width dorsally/interocular width ventrally: 1.08–1.14 (1.10).
</paragraph>
</subSubSection>
<subSubSection id="C3366511CE37FFF2C0D5FB4E332CFB31" pageId="7" pageNumber="161" type="distribution">
<paragraph id="8B93369ACE37FFF2C0D5FB4E332CFB31" blockId="7.[151,1437,151,2015]" pageId="7" pageNumber="161">
<emphasis id="B958EA88CE37FFF2C0D5FB4E305FFB78" bold="true" box="[199,352,1162,1187]" pageId="7" pageNumber="161">Distribution.</emphasis>
Known only from
<taxonomicName id="4C2C4D19CE37FFF2C251FB4E3395FB78" box="[579,682,1162,1187]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="rimatara">Rimatara</taxonomicName>
and expected to be a
<taxonomicName id="4C2C4D19CE37FFF2C3BDFB4E3529FB78" box="[943,1046,1162,1187]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="rimatara">Rimatara</taxonomicName>
endemic. Likely extinct: material known from other Austral Archipelago islands (Tubuai, Raivavae and Rapa) and nearby
<collectingCountry id="F33B760ACE37FFF2C480FB6A3418FB1C" box="[1170,1319,1198,1223]" name="Cook Islands" pageId="7" pageNumber="161">Cook Islands</collectingCountry>
(Mangaia and Atiu) represent different taxa.
</paragraph>
</subSubSection>
<subSubSection id="C3366511CE37FFF2C0D5FB323445FAD4" box="[199,1402,1270,1295]" pageId="7" pageNumber="161" type="etymology">
<paragraph id="8B93369ACE37FFF2C0D5FB323445FAD4" blockId="7.[151,1437,151,2015]" box="[199,1402,1270,1295]" pageId="7" pageNumber="161">
<emphasis id="B958EA88CE37FFF2C0D5FB323071FAD4" bold="true" box="[199,334,1270,1295]" pageId="7" pageNumber="161">Etymology.</emphasis>
Named for its occurrence and apparent endemism on the island of
<taxonomicName id="4C2C4D19CE37FFF2C42EFB32359AFAD4" box="[1084,1189,1270,1295]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="rimatara">Rimatara</taxonomicName>
,
<collectingCountry id="F33B760ACE37FFF2C4A3FB323448FAD4" box="[1201,1399,1270,1295]" name="French Polynesia" pageId="7" pageNumber="161">French Polynesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3366511CE37FFFDC0D5FADE324BFE9B" lastPageId="8" lastPageNumber="162" pageId="7" pageNumber="161" type="materials_examined">
<paragraph id="8B93369ACE37FFF2C0D5FADE3069F9D0" blockId="7.[151,1437,151,2015]" pageId="7" pageNumber="161">
<emphasis id="B958EA88CE37FFF2C0D5FADE337CFAE8" bold="true" box="[199,579,1306,1331]" pageId="7" pageNumber="161">Associated (Non-type) material.</emphasis>
Although currently impossible to prove unreservedly that different sclerites are conspecific with the type material (unless articulated), we consider it likely that the abundant elytral type from the
<taxonomicName id="4C2C4D19CE37FFF2C0D1FAA63015FAA0" box="[195,298,1378,1403]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="rimatara">Rimatara</taxonomicName>
sediments matches the heads and pronota of
<taxonomicName id="4C2C4D19CE37FFF2C32EFAA73280FAA1" box="[828,959,1379,1402]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="161" phylum="Arthropoda" rank="species" species="rimatara">
<emphasis id="B958EA88CE37FFF2C32EFAA73280FAA1" box="[828,959,1379,1402]" italics="true" pageId="7" pageNumber="161">P. rimatara</emphasis>
</taxonomicName>
. For
<taxonomicName id="4C2C4D19CE37FFF2C412FAA63555FAA0" box="[1024,1130,1378,1403]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="rimatara">Rimatara</taxonomicName>
, given only one species is abundant, we are sufficiently confident to consider the sclerites conspecific but because we are not able to be 100% certain, we must exclude the elytra from the type series. Similarly, although the holotype is an articulated head and pronotum we do not include isolated heads in the type series due to the possibility that they may represent more than one species.
</paragraph>
<paragraph id="8B93369ACE37FFF2C0D5F9D230DBF894" blockId="7.[151,1437,151,2015]" pageId="7" pageNumber="161">
<materialsCitation id="3B443CC7CE37FFF2C0D5F9D23524F9AC" ID-GBIF-Occurrence="1437756122" collectorName="Maunutu Swamp C" country="French Polynesia" pageId="7" pageNumber="161" specimenCount="1">
<emphasis id="B958EA88CE37FFF2C0D5F9D230D4F9F4" bold="true" box="[199,491,1558,1583]" pageId="7" pageNumber="161">Other non-type material</emphasis>
(sclerites other than prothorax or articulated head and prothorax):
<collectingCountry id="F33B760ACE37FFF2C4C6F9D234A7F9F4" box="[1236,1432,1558,1583]" name="French Polynesia" pageId="7" pageNumber="161">French Polynesia</collectingCountry>
, Austral archipelago,
<taxonomicName id="4C2C4D19CE37FFF2C192F9FE30D5F988" box="[384,490,1594,1619]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="rimatara">Rimatara</taxonomicName>
,
<collectorName id="26D9534CCE37FFF2C1E6F9FF33F1F988" box="[500,718,1595,1619]" pageId="7" pageNumber="161">Maunutu Swamp C</collectorName>
1T2 2008 excavation, 500 ml sediment sample,
<quantity id="4CD49B7FCE37FFF2C4F0F9FE346FF989" box="[1250,1360,1594,1619]" metricMagnitude="-1" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="7" pageNumber="161" unit="cm" value="62.5" valueMax="65.0" valueMin="60.0">60–65 cm</quantity>
depth, 1 head;
<emphasis id="B958EA88CE37FFF2C0E4F99B300FF9AC" box="[246,304,1631,1655]" italics="true" pageId="7" pageNumber="161">ditto,</emphasis>
except
<quantity id="4CD49B7FCE37FFF2C19CF99A30C3F9AC" box="[398,508,1630,1655]" metricMagnitude="-1" metricUnit="m" metricValue="7.25" metricValueMax="7.5" metricValueMin="7.0" pageId="7" pageNumber="161" unit="cm" value="72.5" valueMax="75.0" valueMin="70.0">70-75 cm</quantity>
depth, 1 left elytron fragment (not illustrated)
</materialsCitation>
;
<materialsCitation id="3B443CC7CE37FFF2C438F99B30B7F940" ID-GBIF-Occurrence="1437756117" collectorName="Maunutu Swamp C" country="French Polynesia" pageId="7" pageNumber="161" specimenCount="1">
<emphasis id="B958EA88CE37FFF2C438F99B355CF9AC" box="[1066,1123,1631,1655]" italics="true" pageId="7" pageNumber="161">ditto,</emphasis>
except
<quantity id="4CD49B7FCE37FFF2C4D0F99A340BF9AC" box="[1218,1332,1630,1655]" metricMagnitude="-1" metricUnit="m" metricValue="8.25" metricValueMax="8.5" metricValueMin="8.0" pageId="7" pageNumber="161" unit="cm" value="82.5" valueMax="85.0" valueMin="80.0">80–85 cm</quantity>
depth, 1 complete left elytron
</materialsCitation>
;
<materialsCitation id="3B443CC7CE37FFF2C186F9473537F940" ID-GBIF-Occurrence="1437756113" box="[404,1032,1666,1691]" collectorName="Maunutu Swamp C" country="French Polynesia" pageId="7" pageNumber="161" specimenCount="1">
<emphasis id="B958EA88CE37FFF2C186F94730F0F940" box="[404,463,1667,1691]" italics="true" pageId="7" pageNumber="161">ditto,</emphasis>
except
<quantity id="4CD49B7FCE37FFF2C23BF94633A5F941" box="[553,666,1666,1691]" metricMagnitude="-1" metricUnit="m" metricValue="9.25" metricValueMax="9.5" metricValueMin="9.0" pageId="7" pageNumber="161" unit="cm" value="92.5" valueMax="95.0" valueMin="90.0">90–95 cm</quantity>
depth, 1 complete right elytron
</materialsCitation>
;
<materialsCitation id="3B443CC7CE37FFF2C407F9473575F964" ID-GBIF-Occurrence="1437756107" collectorName="Maunutu Swamp C" country="French Polynesia" pageId="7" pageNumber="161" specimenCount="1">
<emphasis id="B958EA88CE37FFF2C407F9473570F940" box="[1045,1103,1667,1691]" italics="true" pageId="7" pageNumber="161">ditto,</emphasis>
except
<quantity id="4CD49B7FCE37FFF2C4B8F9463408F941" box="[1194,1335,1666,1691]" metricMagnitude="0" metricUnit="m" metricValue="1.025" metricValueMax="1.05" metricValueMin="1.0" pageId="7" pageNumber="161" unit="cm" value="102.5" valueMax="105.0" valueMin="100.0">100–105 cm</quantity>
depth, 1 largely complete head, 1 pronotum, 2 complete left elytra, 2 complete right elytra
</materialsCitation>
;
<materialsCitation id="3B443CC7CE37FFF2C44AF963346BF8DC" ID-GBIF-Occurrence="1437756116" collectionCode="MATO" collectorName="Maunutu Swamp C" country="French Polynesia" location="Austral" municipality="Maunutu Swamp" pageId="7" pageNumber="161" specimenCode="MATO 2008" specimenCount="1">
<emphasis id="B958EA88CE37FFF2C44AF96335ACF964" box="[1112,1171,1703,1727]" italics="true" pageId="7" pageNumber="161">ditto,</emphasis>
except
<quantity id="4CD49B7FCE37FFF2C4FDF96234BFF964" box="[1263,1408,1702,1727]" metricMagnitude="0" metricUnit="m" metricValue="1.225" metricValueMax="1.25" metricValueMin="1.2" pageId="7" pageNumber="161" unit="cm" value="122.5" valueMax="125.0" valueMin="120.0">120–125 cm</quantity>
, 1 complete right elytron, 1 complete left elytron, 1 largely complete left elytron;
<collectingCountry id="F33B760ACE37FFF2C478F90E3408F938" box="[1130,1335,1738,1763]" name="French Polynesia" pageId="7" pageNumber="161">French Polynesia</collectingCountry>
,
<location id="8EF36041CE37FFF2C558F90E34A3F938" LSID="urn:lsid:plazi:treatment:0385878CCE33FFFDC085FA72324BFE9B:8EF36041CE37FFF2C558F90E34A3F938" box="[1354,1436,1738,1763]" country="French Polynesia" municipality="Maunutu Swamp" name="Austral" pageId="7" pageNumber="161">Austral</location>
archipelago,
<taxonomicName id="4C2C4D19CE37FFF2C138F92A30ABF8DC" box="[298,404,1774,1799]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="rimatara">Rimatara</taxonomicName>
,
<collectingMunicipality id="6BF7ACE0CE37FFF2C1B0F9343357F8DC" box="[418,616,1775,1799]" pageId="7" pageNumber="161">Maunutu Swamp</collectingMunicipality>
,
<specimenCode id="DB8A9EE1CE37FFF2C264F92B3238F8DC" box="[630,775,1775,1799]" collectionCode="MATO" pageId="7" pageNumber="161">
<collectionCode id="ED3DAE5FCE37FFF2C264F92B33FAF8DC" box="[630,709,1775,1799]" pageId="7" pageNumber="161">MATO</collectionCode>
2008
</specimenCode>
core,
<quantity id="4CD49B7FCE37FFF2C340F92A32DFF8DC" box="[850,992,1774,1799]" metricMagnitude="0" metricUnit="m" metricValue="3.175" metricValueMax="3.2" metricValueMin="3.15" pageId="7" pageNumber="161" unit="cm" value="317.5" valueMax="320.0" valueMin="315.0">315–320 cm</quantity>
depth, 1 complete right elytron
</materialsCitation>
;
<materialsCitation id="3B443CC7CE37FFF2C570F92B33C4F8F0" ID-GBIF-Occurrence="1437756118" collectionCode="MATO" collectorName="Maunutu Swamp C" country="French Polynesia" location="Austral" municipality="Maunutu Swamp" pageId="7" pageNumber="161" specimenCount="1">
<emphasis id="B958EA88CE37FFF2C570F92B34A4F8DC" box="[1378,1435,1775,1799]" italics="true" pageId="7" pageNumber="161">ditto,</emphasis>
except
<quantity id="4CD49B7FCE37FFF2C0FEF8D63043F8F1" box="[236,380,1810,1835]" metricMagnitude="0" metricUnit="m" metricValue="3.575" metricValueMax="3.6" metricValueMin="3.55" pageId="7" pageNumber="161" unit="cm" value="357.5" valueMax="360.0" valueMin="355.0">355–360 cm</quantity>
depth, 1 complete right elytron
</materialsCitation>
;
<materialsCitation id="3B443CC7CE37FFF2C318F8D730DBF894" ID-GBIF-Occurrence="1437756110" collectionCode="BPBM" collectorName="Maunutu Swamp C" country="French Polynesia" location="Austral" municipality="Maunutu Swamp" pageId="7" pageNumber="161" specimenCount="1">
<emphasis id="B958EA88CE37FFF2C318F8D7327AF8F0" box="[778,837,1811,1835]" italics="true" pageId="7" pageNumber="161">ditto,</emphasis>
except
<quantity id="4CD49B7FCE37FFF2C3B4F8D63509F8F1" box="[934,1078,1810,1835]" metricMagnitude="0" metricUnit="m" metricValue="4.475" metricValueMax="4.5" metricValueMin="4.45" pageId="7" pageNumber="161" unit="cm" value="447.5" valueMax="450.0" valueMin="445.0">445–450 cm</quantity>
depth, 1 head. Representative sclerites in
<collectionCode id="ED3DAE5FCE37FFF2C104F8FC305BF894" box="[278,356,1848,1871]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="7" pageNumber="161">BPBM</collectionCode>
and Porch.
</materialsCitation>
</paragraph>
<paragraph id="8B93369ACE37FFFDC0D5F89E324BFE9B" blockId="7.[151,1437,151,2015]" lastBlockId="8.[151,1437,151,320]" lastPageId="8" lastPageNumber="162" pageId="7" pageNumber="161">
Description of
<emphasis id="B958EA88CE37FFF2C166F89E3085F8A8" bold="true" box="[372,442,1882,1907]" pageId="7" pageNumber="161">elytra</emphasis>
considered to belong to
<taxonomicName id="4C2C4D19CE37FFF2C2C9F89F3262F8A9" box="[731,861,1883,1906]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="161" phylum="Arthropoda" rank="species" species="rimatara">
<emphasis id="B958EA88CE37FFF2C2C9F89F3262F8A9" box="[731,861,1883,1906]" italics="true" pageId="7" pageNumber="161">P. rimatara</emphasis>
</taxonomicName>
: subparallel, widest about middle, weakly sinuate and narrowing to rounded-subangular humerus, marked with one-several blunt teeth, to apex. Elytral intervals 1-6 mostly wider than striae, becoming narrower from interval 6. Intervals flat, widened at elytral base and apex where they are up to twice as wide as striae. Intervals apparently lacking microsculpture, shiny, and with a single row of very widely spaced punctures, mostly&gt;4-5 times the length of the transversely elongate punctures. Striae with distinctive 'oxhide ingot' punctures separated by about, to slightly more than, their length; an oxhide ingot is a bronze age metal ingot (usually copper or tin) with a characteristic shape that is very similar to the distinctive puncture shape of some
<taxonomicName id="4C2C4D19CE38FFFDC1B7FEC1330FFEC7" box="[421,560,261,284]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="B958EA88CE38FFFDC1B7FEC1330FFEC7" box="[421,560,261,284]" italics="true" pageId="8" pageNumber="162">Pycnomerus</emphasis>
</taxonomicName>
elytra (
<bibRefCitation id="EFBD4B6BCE38FFFDC297FEC03556FEC6" author="Pulak" box="[645,1129,260,285]" pageId="8" pageNumber="162" refString="Pulak, C. (1998) The Uluburun shipwreck: an overview. The International Journal of Nautical Archaeology, 27 (3), 188 - 224." type="journal article" year="1998">
Pulak, 1998; see
<figureCitation id="13172A1FCE38FFFDC357FEC032A1FEC7" box="[837,926,260,285]" captionStart="FIGURE 1" captionStartId="3.[151,250,1273,1295]" captionTargetBox="[157,1436,203,1242]" captionTargetId="figure@3.[151,1436,195,1251]" captionTargetPageId="3" captionText="FIGURE 1. Measurements. Not all measurements for a sclerite or set of sclerites were always measured reflecting the variable nature of the subfossil material (an articulated head and pronotum may obscure, for example, the temporal width of the head) and preservation. A. An example of an oxhide ingot with the shape of the distinctive punctures found on many Pycnomerus elytra—see text for further details." httpUri="https://zenodo.org/record/344552/files/figure.png" pageId="8" pageNumber="162">Figure 1</figureCitation>
A for an example
</bibRefCitation>
). Elytron length (
<quantity id="4CD49B7FCE38FFFDC539FEC031FAFEE4" metricMagnitude="-3" metricUnit="m" metricValue="2.18" metricValueMax="2.45" metricValueMin="1.91" pageId="8" pageNumber="162" unit="mm" value="2.18" valueMax="2.45" valueMin="1.91">1.91–2.45 mm</quantity>
, Av.
<quantity id="4CD49B7FCE38FFFDC0EFFEE3305AFE9B" box="[253,357,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="8" pageNumber="162" unit="mm" value="2.15">2.15 mm</quantity>
). Elytron width (
<quantity id="4CD49B7FCE38FFFDC233FEE333F7FEE4" box="[545,712,295,320]" metricMagnitude="-4" metricUnit="m" metricValue="5.35" metricValueMax="5.8" metricValueMin="4.9" pageId="8" pageNumber="162" unit="mm" value="0.5349999999999999" valueMax="0.58" valueMin="0.49">0.49–0.58 mm</quantity>
, Av.
<quantity id="4CD49B7FCE38FFFDC312FEE33256FE9B" box="[768,873,295,320]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="8" pageNumber="162" unit="mm" value="0.54">0.54 mm</quantity>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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