165 lines
14 KiB
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165 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.727.20222" ID-GBIF-Dataset="13853cb6-522e-4f94-b633-c4a51ba63624" ID-PMC="PMC5799789" ID-Pensoft-Pub="1313-2970-727-1" ID-PubMed="29416388" ID-ZBK="12B663F7190040788E1EEF8BAC4DF81B" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-727-1" ModsDocOrigin="ZooKeys 727" ModsDocTitle="A review of the Madagascan pelican spiders of the genera Eriauchenius O. Pickard-Cambridge, 1881 and Madagascarchaea gen. n. (Araneae, Archaeidae)" checkinTime="1515696595036" checkinUser="pensoft" docAuthor="Wood, Hannah M. & Scharff, Nikolaj" docDate="2017" docId="980680473022FD9CB69384D1AE8AFA32" docLanguage="en" docName="ZooKeys 727: 1-96" docOrigin="ZooKeys 727" docSource="http://dx.doi.org/10.3897/zookeys.727.20222" docTitle="Madagascarchaea fohy Wood & Scharff, 2017, sp. n." docType="treatment" docUuid="6ACB2E06-A02F-46E2-879E-D88D40F9C556" docUuidSource="ZooBank" docVersion="5" lastPageNumber="1" masterDocId="FFEBD959ED2FFFAA9450FF8B1957B348" masterDocTitle="A review of the Madagascan pelican spiders of the genera Eriauchenius O. Pickard-Cambridge, 1881 and Madagascarchaea gen. n. (Araneae, Archaeidae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="1" updateTime="1668165276717" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of the Madagascan pelican spiders of the genera Eriauchenius O. Pickard-Cambridge, 1881 and Madagascarchaea gen. n. (Araneae, Archaeidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wood, Hannah M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Scharff, Nikolaj</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>727</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>96</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.727.20222</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.727.20222</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-727-1</mods:identifier>
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<mods:identifier type="ZBK">12B663F7190040788E1EEF8BAC4DF81B</mods:identifier>
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<mods:identifier type="ZooBank">12B663F7190040788E1EEF8BAC4DF81B</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="140245900" LSID="urn:lsid:zoobank.org:act:6ACB2E06-A02F-46E2-879E-D88D40F9C556" httpUri="http://treatment.plazi.org/id/980680473022FD9CB69384D1AE8AFA32" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/6ACB2E06-A02F-46E2-879E-D88D40F9C556" class="Arachnida" family="Archaeidae" genus="Madagascarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madagascarchaea fohy" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fohy">Madagascarchaea fohy</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. n.</taxonomicNameLabel>
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Figs 23, 33
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Male holotype: MADAGASCAR, Toamasina, Ivoloina Parque Zoologique, 12 km from Tamatave,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.056">18°03'21.6"S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.359028">49°21'32.5"E</geoCoordinate>
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, 26m, 19 Feb 2003, disturbed rainforest, general collecting night, D. Andriamalala, D. Silva, et al. (deposited in CAS; CASENT9015766).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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MADGASCAR: Female paratype, same data as holotype (CASENT9015766); 1F,1juv, Toamasina, Parc National Masoala, Ambohitsitondroina Mt., Ambanizana,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-15.572083">15°34'19.5"S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="50.006943">50°00'25"E</geoCoordinate>
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, 900-950 m, 27 Feb 2003, rainforest, general collecting, D. Andriamalala, D. Silva, et al. (CASENT9015493).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The specific name is a noun in apposition; '
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<taxonomicName genus="Palpimanoidea" lsidName="Palpimanoidea fohy" pageId="0" pageNumber="1" rank="species" species="fohy">fohy</taxonomicName>
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' means
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<normalizedToken originalValue="‘shorty’">'shorty'</normalizedToken>
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in Malagasy.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Distinguished from all other archaeids, except
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<taxonomicName lsidName="M. vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">M. vadoni</taxonomicName>
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,
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<taxonomicName lsidName="M. legendrei" pageId="0" pageNumber="1" rank="species" species="legendrei">M. legendrei</taxonomicName>
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,
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<taxonomicName lsidName="M. rabesahala" pageId="0" pageNumber="1" rank="species" species="rabesahala">M. rabesahala</taxonomicName>
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sp. n., and
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<taxonomicName lsidName="M. borimontsina" pageId="0" pageNumber="1" rank="species" species="borimontsina">M. borimontsina</taxonomicName>
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(likely, because the male is unknown) by having a retrolateral apophysis on the distal side of both the palpal femur and patella (fig. 12B from
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<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood 2008</bibRefCitation>
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), and because the FSGP is highly reduced and lacks
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<normalizedToken originalValue="“wings”">"wings"</normalizedToken>
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(Figs 23B, 24B, 25B).
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<taxonomicName lsidName="M. fohy" pageId="0" pageNumber="1" rank="species" species="fohy">M. fohy</taxonomicName>
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sp. n. is distinguished from
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<taxonomicName lsidName="M. vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">M. vadoni</taxonomicName>
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,
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<taxonomicName lsidName="M. legendrei" pageId="0" pageNumber="1" rank="species" species="legendrei">M. legendrei</taxonomicName>
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, and
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<taxonomicName lsidName="M. rabesahala" pageId="0" pageNumber="1" rank="species" species="rabesahala">M. rabesahala</taxonomicName>
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sp. n. by having a heavily sclerotized rod-shaped sclerite on the palpal bulb (Fig. 23H, J, arrow), and by the presence of a retrolateral protrusion on the cymbium that has setae (Fig. 23C, arrow).
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<taxonomicName lsidName="M. fohy" pageId="0" pageNumber="1" rank="species" species="fohy">M. fohy</taxonomicName>
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sp. n. is further distinguished from
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<taxonomicName lsidName="M. vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">M. vadoni</taxonomicName>
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by having 6 spines that are not on protrusions on the cephalic crown, opposed to the small protrusions seen in
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<taxonomicName lsidName="M. vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">M. vadoni</taxonomicName>
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(Wood, 2008: fig. 18A, C); from
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<taxonomicName lsidName="M. borimontsina" pageId="0" pageNumber="1" rank="species" species="borimontsina">M. borimontsina</taxonomicName>
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by having less than 6 true teeth on the cheliceral retromargin (
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<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood 2008</bibRefCitation>
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), by lacking a point on the AME bulge (
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<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood 2008</bibRefCitation>
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: fig. 18D), and by lacking a cheliceral swelling on the posterior-basal side of the chelicerae (
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<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood 2008</bibRefCitation>
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: fig. 18D).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Male holotype (CASENT9015766, from Ivoloina Parque Zoologique, Madagascar). Total length 1.61, carapace 0.68 long, 0.55 wide. Abdomen 0.85 long, 0.93 high. Carapace tilt angle 58.1°, tilt height (CtH) 1.29, constriction 0.33, head length 0.56, neck length 0.70. CtH divided by carapace length 1.91. Cephalon with AME on large bulge, and with 6 small post-ocular spines on the crown, not on protrusions, and 1 small spine between the LE and median eyes (on each side, for a total of 2). Chelicerae 1.40 long, and with a small spine 0.27 from base of chelicerae that projects downward (Fig. 23A). Femur I 1.96 long. Sternum 0.45 long, 0.27 wide. Carapace, chelicerae, sternum and legs reddish brown with white setae. Legs with darker annulations on tibiae and metatarsi. Abdomen mottled with dark brown areas and lighter yellowish areas, interspersed with white setae (Fig. 23A). Pedipalpal bulb of the "
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<taxonomicName genus="Palpimanoidea" lsidName="Palpimanoidea vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">vadoni</taxonomicName>
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group" form (Fig. 23
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<normalizedToken originalValue="C–L">C-L</normalizedToken>
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): pedipalpal bulbs elongated with conductor swirling around a mostly membraneous embolus; conductor base triangular (Fig. 23D, G, see basal
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<normalizedToken originalValue="“c”">"c"</normalizedToken>
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), with remainder of conductor elongate and cradling embolus, and with a curved tip; MA translucent and fans out (Fig. 23F, I). Cymbium with retrolateral protrusion that has setae (Fig. 23C, arrow). Embolus with heavily sclerotized rod-like piece (Fig. 23H, J, arrow).
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</paragraph>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 23.
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<taxonomicName class="Arachnida" family="Archaeidae" genus="Madagascarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madagascarchaea fohy" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fohy">Madagascarchaea fohy</taxonomicName>
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sp. n. A male (holotype, CASENT9015766) habitus, lateral view B female (CASENT9015766) internal genitalia, dorsal view
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<normalizedToken originalValue="C–L">C-L</normalizedToken>
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male pedipalpal bulbs (holotype, CASENT9015766)
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<normalizedToken originalValue="D–F">D-F</normalizedToken>
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right bulb, image reversed C,
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<normalizedToken originalValue="G–L">G-L</normalizedToken>
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left bulb C cymbium close-up, dorsal view, arrow showing the retrolateral cymbium protrusion that has setae D, G, J prolateral view, arrow showing rod-like piece on embolus E, H, K ventral view, arrow showing rod-like piece on embolus F, I, L retrolateral view. Scale bars: 1 mm (A); 0.25 mm (B, D).
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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Female paratype (CASENT9015766). Total length 1.97, carapace 0.75 long, 0.63 wide. Abdomen 1.10 long, 1.38 high. Carapace tilt angle 54.2°, tilt height (CtH) 1.55, constriction 0.41, head length 0.73, neck length 0.82. CtH divided by carapace length 2.07. Cephalon as in male. Chelicerae 1.55 long, and with a short spine 0.28 from base of chelicerae and projecting downward (Fig. 23A). Femur I 2.04 long. Sternum 0.50 long, 0.29 wide. Colors as in male. Genitalia of the "
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<taxonomicName genus="Palpimanoidea" lsidName="Palpimanoidea vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">vadoni</taxonomicName>
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-group" form (Fig. 23B): with one group of poreplates on each side of the bursa anterior, and FSGP highly reduced, lacking
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<normalizedToken originalValue="“wings.”">"wings."</normalizedToken>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="variation">
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<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
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<paragraph pageId="0" pageNumber="1">Total length 1.67-1.97 (females; n=2); Carapace length 0.71-0.75 (females; n=2); Left and right femur I missing in one female (CASENT9015493). Femur I 2.73 times the length of carapace in females (n=1); CtH divided by carapace length 1.92-2.07 in females (n=2).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="natural history">
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<paragraph pageId="0" pageNumber="1">Natural history.</paragraph>
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<paragraph pageId="0" pageNumber="1">Specimens were collected in disturbed and undisturbed rainforest from 26-950 m in elevation by general collecting.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">Known only from northeastern and central-eastern Madagascar (Fig. 33).</paragraph>
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</subSubSection>
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</treatment>
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</document> |