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525 lines
69 KiB
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<document ID-CLB-Dataset="89308" ID-DOI="http://dx.doi.org/10.3897/zookeys.1110.85399" ID-GBIF-Dataset="04ac90a6-24ce-43c8-a06c-b1f1680b3b8e" ID-PMC="PMC9848723" ID-Pensoft-Pub="1313-2970-1110-39" ID-Pensoft-UUID="766F9A94793D5793A9BE1E70F575798B" ID-PubMed="36761454" ID-ZooBank="4681CC6DE5F347C6B1D052DEA78BE7C3" ModsDocID="1313-2970-1110-39" checkinTime="1657006630715" checkinUser="pensoft" docAuthor="Sian Man, Nem, Siriboon, Thanit, Lin, Aung, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="A9B3BC261C09572FA717DFAC7ACCC058" docLanguage="en" docName="ZooKeys 1110: 39-102" docOrigin="ZooKeys 1110" docPubDate="2022-07-04" docSource="http://dx.doi.org/10.3897/zookeys.1110.85399" docTitle="Carinartemis sankeyi" docType="treatment" docVersion="5" id="766F9A94793D5793A9BE1E70F575798B" lastPageNumber="39" masterDocId="766F9A94793D5793A9BE1E70F575798B" masterDocTitle="Revision of the carnivorous land snail family Streptaxidae (Stylommatophora, Achatinina) in Myanmar, with description of four new species" masterLastPageNumber="102" masterPageNumber="39" pageNumber="39" updateTime="1732936926474" updateUser="ExternalLinkService">
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<mods:title id="39B4F5FB14BDFEF9774D8CDD633D8DCE">Revision of the carnivorous land snail family Streptaxidae (Stylommatophora, Achatinina) in Myanmar, with description of four new species</mods:title>
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<mods:namePart id="FEC1F437E056983ED90464EE3E8A19B1">Sian Man, Nem</mods:namePart>
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<mods:affiliation id="26B805CBA188B9960D49B7C538D38163">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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<mods:namePart id="3378EAB2DA26A9FC41661C804C6EA4D2">Siriboon, Thanit</mods:namePart>
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<mods:affiliation id="DDBC7491F33FFDB3820DA0290D7CE70D">Department of Biology, Faculty of Science, Srinakharinwirot University, Bangkok, 10110, Thailand</mods:affiliation>
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<mods:namePart id="0392E51E14C44D5067B6B03DE4A567CD">Lin, Aung</mods:namePart>
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<mods:affiliation id="D8F144DF9AEBB9D7659CBA6DC93CB0B1">Fauna and Flora International, No. 35, 3</mods:affiliation>
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<mods:namePart id="B4C6305C30A828CD69EDF56EDE489004">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="1FE2BFA17E8AB207A4A00A59E896D274">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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<mods:namePart id="87AE2C4AA19591EF8B465409D39AC17C">Panha, Somsak</mods:namePart>
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<mods:affiliation id="9939D2DC38F3F078B2754D1DD3AC7E03">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="4CD78C8282D5D9181706784DFF2141B6" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:title id="874FFC377AE663022DF7C1610611A063">ZooKeys</mods:title>
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<mods:date id="3827C3840EBEAD93B6256552166E88BF">2022</mods:date>
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<mods:number id="E9CA46FE4F97E782E7073EB432C6352A">1110</mods:number>
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<mods:identifier id="9B172F1C0E893F6768AA5C60EBE84BC6" type="DOI">http://dx.doi.org/10.3897/zookeys.1110.85399</mods:identifier>
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<subSection id="CF3DD558DB68DD2FEF6D0E58B8E2DDAF" pageId="0" pageNumber="39" type="systematics">
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<treatment id="A9B3BC261C09572FA717DFAC7ACCC058" ID-GBIF-Taxon="196387993" LSID="urn:lsid:plazi:treatment:A9B3BC261C09572FA717DFAC7ACCC058" httpUri="http://treatment.plazi.org/id/A9B3BC261C09572FA717DFAC7ACCC058" lastPageNumber="39" pageId="0" pageNumber="39" scope_family="Streptaxidae" scope_order="Stylommatophora" scope_subOrder="Achatinina">
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<subSubSection id="83F3D18F6090C4787F8AE019C537AF69" pageId="0" pageNumber="39" type="nomenclature">
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<paragraph id="C43396EA0F0CEDDDBA773308E333C47E" pageId="0" pageNumber="39">
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<taxonomicName id="620F94D591B830CEDD3CA59264F1AAFC" ID-CoL="9FW8W" LSID="A9B3BC26-1C09-572F-A717-DFAC7ACCC058" authority="(Benson, 1859)" baseAuthorityName="Benson" baseAuthorityYear="1859" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carinartemis sankeyi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="sankeyi">Carinartemis sankeyi (Benson, 1859)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="F91B5122DCEFA1B0E1484ABE5ABD25D2" pageId="0" pageNumber="39" type="description">
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<paragraph id="ED6DC744A6AA504FD18AD8D4F20E18B7" pageId="0" pageNumber="39">
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<figureCitation id="68976B9667B7AD28E1DD0169AFE16CF9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Approximate sampling locations used in this study. The location name and coordinates are listed in Table 1. Historical type localities of a broad scale, such as Tavoy, Burma, Moulmein, Mergui, and Tenasserim, are not indicated on this map." figureDoi="10.3897/zookeys.1110.85399.figure1" httpUri="https://binary.pensoft.net/fig/710466" pageId="0" pageNumber="39">Figs 1</figureCitation>
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<figureCitation id="F63C4B1009DBC01F46C16E5CD9130E50" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Living snails A Discartemon tonywhitteni from near Phra (Buddha) Cave, Tanintharyi Region (shell width ~ 10 mm) B Discartemon paurodeviatus sp. nov. from the type locality (shell width ~ 11 mm) C Haploptychius heliakosus sp. nov. from the type locality (shell height ~ 9 mm) D Carinartemis exacutus from Lun Nga Mountain, Kayin State (shell height ~ 14 mm) E, F Carinartemis sankeyi (E) from Kayon Cave, Mon State (shell height ~ 10 mm) and (F) from Saddan Cave, Mawlamyine, Mon State (shell height ~ 10 mm)." figureDoi="10.3897/zookeys.1110.85399.figure2" httpUri="https://binary.pensoft.net/fig/710467" pageId="0" pageNumber="39">, 2E, F</figureCitation>
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<figureCitation id="C342C0C218CCE1D5D3ED81FC8CAE6963" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Carinartemis sankeyi A syntype UMZC I. 102740 from Moulmein B specimen NHMUK 1888.12.4.792 - 794 from Moulmein C specimen NHMUK 1906.2.2.341 from Moulmein with apertural dentition D specimen NHMUK 1871.9.23.61 from Moulmein E specimen NHMUK ex. Cuming collection from Mergui." figureDoi="10.3897/zookeys.1110.85399.figure19" httpUri="https://binary.pensoft.net/fig/710484" pageId="0" pageNumber="39">, 19</figureCitation>
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<figureCitation id="D21C08A418BFB2DED02FFCCC0295183F" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Carinartemis sankeyi A specimen NHMUK 1903.7.1.4001 from Moulmein B specimen NHMUK 1888.12.4.781 - 783 from Kwengan Hill C specimen CUMZ 13031 from Dhammatat Cave, Mawlamyine, Mon State with apertural dentition D specimen CUMZ 13029 from Kayon Cave, Mawlamyine, Mon State with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure20" httpUri="https://binary.pensoft.net/fig/710485" pageId="0" pageNumber="39">, 20</figureCitation>
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<figureCitation id="F5FD757F4093851901B15857AE364061" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Genital anatomy of A, B Carinartemis exacutus, specimen CUMZ 13021 A reproductive system and B insertion of vas deferens into penial sheath C, D Carinartemis sankeyi, specimen CUMZ 13029 C reproductive system and D insertion of vas deferens into penial sheath." figureDoi="10.3897/zookeys.1110.85399.figure21" httpUri="https://binary.pensoft.net/fig/710486" pageId="0" pageNumber="39">, 21C, D</figureCitation>
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<figureCitation id="F21700252E31CE5FC9CC0D7E68BFC9A5" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Internal sculpture of genitalia of Carinartemis sankeyi, specimen CUMZ 13029 A atrium surface B arrangement of penial hooks on proximal part of penis C lateral view of exposed proximal penial hooks D penial hooks on middle part of penis E arrangement of penial hooks on distal part of penis F uncovered distal penial hooks with pointed tip G distal penial hooks embedded in penial papillae H vaginal surface with vaginal pores." figureDoi="10.3897/zookeys.1110.85399.figure23" httpUri="https://binary.pensoft.net/fig/710488" pageId="0" pageNumber="39">, 23</figureCitation>
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<figureCitation id="68670B7AE5DC7240FA29C95BFD2C4FFC" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Radula of specimens A Discartemon tonywhitteni, paratype CUMZ 13001 B Discartemon paurodeviatus sp. nov., paratype CUMZ 13003 C Haploptychius heliakosus sp. nov., paratype CUMZ 13014 D Carinartemis exacutus, specimen CUMZ 13021 E, F Carinartemis sankeyi, specimen CUMZ 13029 E overview of radula and F high magnification of central teeth." figureDoi="10.3897/zookeys.1110.85399.figure24" httpUri="https://binary.pensoft.net/fig/710489" pageId="0" pageNumber="39">, 24E, F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="AE05882D375DFA1697314F30CC2DB3F0" pageId="0" pageNumber="39" type="reference_group">
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<paragraph id="994064336C0F962B5E4E5B254A0DE544" pageId="0" pageNumber="39">
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<taxonomicName id="E3FADB62B11FADDB9081D6C96D10E23B" class="Gastropoda" family="Streptaxidae" genus="Streptaxis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Streptaxis sankeyi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="sankeyi">Streptaxis sankeyi</taxonomicName>
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Benson, 1859a: 472. Type locality: Moulmein [Mawlamyine Township, Mon State, Myanmar].
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<bibRefCitation id="6FB19F8FD7605A9DBFA22FCABB122B63" author="Pfeiffer, L" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="39" refId="B63" refString="Pfeiffer, L, 1868. Monographia Heliceorum Viventium (Volume 5). Backhaus FA, Lipsiae, 565." title="Monographia Heliceorum Viventium (Volume 5). Backhaus FA, Lipsiae, 565." year="1868">Pfeiffer 1868</bibRefCitation>
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: 442.
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<bibRefCitation id="6F9D12FCFD58B07044CF76492CD1F544" author="Hanley, SCT" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="39" refId="B33" refString="Hanley, SCT, Theobald, W, 1870. Conchologia Indica: Illustrations of the Land and Freshwater Shells of British India. London[i-xviii+] 1-65 pp.[160 pls.] [pp 1-18 pls 1-40(1870) ; pp 19-28, pls 41-60 (1872); pp 29-34, pls 61-80 (1873); pp 35-48, pls 81-120 (1874); pp 49-56, pls 121-140 (1875); pp 57-65, pls 141-160 (1876)] [Published in parts, dates after Prashad (1927)]." title="Conchologia Indica: Illustrations of the Land and Freshwater Shells of British India. London [i-xviii +] 1 - 65 pp. [160 pls.] [pp 1 - 18 pls 1 - 40 (1870); pp 19 - 28, pls 41 - 60 (1872); pp 29 - 34, pls 61 - 80 (1873); pp 35 - 48, pls 81 - 120 (1874); pp 49 - 56, pls 121 - 140 (1875); pp 57 - 65, pls 141 - 160 (1876)] [Published in parts, dates after Prashad (1927)]." year="1870">Hanley and Theobald 1870</bibRefCitation>
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: 4, pl. 8, fig. 7.
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<bibRefCitation id="42BFD98C53087FFBDF2498108058A981" DOI="https://doi.org/10.5962/bhl.title.6534" author="Tryon Jr, GW" journalOrPublisher="Gustav Fischer, Jena" pageId="0" pageNumber="39" refId="B90" refString="Tryon Jr, GW, 1885. Manual of Conchology, Structure and Systematic, with Illustrations of the Species. Volume 1. The Academy of Natural Science of Philadelphia, 364 pp. https://doi.org/10.5962/bhl.title.6534" title="Manual of Conchology, Structure and Systematic, with Illustrations of the Species. Volume 1. The Academy of Natural Science of Philadelphia, 364 pp." url="https://doi.org/10.5962/bhl.title.6534" year="1885">Tryon 1885</bibRefCitation>
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: 72, pl. 14, figs 2, 3.
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||
<bibRefCitation id="1DF5AB42F0CABFC2D0AA969E4A675503" DOI="https://doi.org/10.5962/bhl.title.13103" author="Blanford, WT" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="39" refId="B10" refString="Blanford, WT, Godwin-Austen, HH, 1908. The Fauna of British India, Including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae. Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.13103" title="The Fauna of British India, Including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae." url="https://doi.org/10.5962/bhl.title.13103" year="1908">Blanford and Godwin-Austen 1908</bibRefCitation>
|
||
: 8, fig. 7.
|
||
</paragraph>
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<paragraph id="45B4E9B16BC54C8A31E102605D1757E2" pageId="0" pageNumber="39">
|
||
<taxonomicName id="0B126042C75B9C1B1439E6782FF27B3F" authorityName="Stoliczka" authorityYear="1871" class="Gastropoda" family="Streptaxidae" genus="Streptaxis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Streptaxis sankeyanus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="sankeyanus">Streptaxis sankeyanus</taxonomicName>
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Stoliczka, 1871: 167, 168, pl. 7, fig. 14 (unjustified emendation).
|
||
<bibRefCitation id="0F407A4F8FD8A1D144223C21B48575CF" author="Hanley, SCT" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="39" refId="B33" refString="Hanley, SCT, Theobald, W, 1870. Conchologia Indica: Illustrations of the Land and Freshwater Shells of British India. London[i-xviii+] 1-65 pp.[160 pls.] [pp 1-18 pls 1-40(1870) ; pp 19-28, pls 41-60 (1872); pp 29-34, pls 61-80 (1873); pp 35-48, pls 81-120 (1874); pp 49-56, pls 121-140 (1875); pp 57-65, pls 141-160 (1876)] [Published in parts, dates after Prashad (1927)]." title="Conchologia Indica: Illustrations of the Land and Freshwater Shells of British India. London [i-xviii +] 1 - 65 pp. [160 pls.] [pp 1 - 18 pls 1 - 40 (1870); pp 19 - 28, pls 41 - 60 (1872); pp 29 - 34, pls 61 - 80 (1873); pp 35 - 48, pls 81 - 120 (1874); pp 49 - 56, pls 121 - 140 (1875); pp 57 - 65, pls 141 - 160 (1876)] [Published in parts, dates after Prashad (1927)]." year="1870">Hanley and Theobald 1870</bibRefCitation>
|
||
: 4, pl. 8, fig. 7.
|
||
<bibRefCitation id="45876C40A6FC3A1F60B7CC74A046DFC4" DOI="https://doi.org/10.5962/bhl.title.11957" author="Nevill, G" journalOrPublisher="Indian Museum, Calcutta" pageId="0" pageNumber="39" refId="B48" refString="Nevill, G, 1878. Hand list of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia. Indian Museum, Calcutta, DOI: https://doi.org/10.5962/bhl.title.11957" title="Hand list of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia." url="https://doi.org/10.5962/bhl.title.11957" year="1878">Nevill 1878</bibRefCitation>
|
||
: 3.
|
||
<bibRefCitation id="EA30F15E7A730F95038B7C6132F85A94" DOI="https://doi.org/10.5962/bhl.title.6534" author="Tryon Jr, GW" journalOrPublisher="Gustav Fischer, Jena" pageId="0" pageNumber="39" refId="B90" refString="Tryon Jr, GW, 1885. Manual of Conchology, Structure and Systematic, with Illustrations of the Species. Volume 1. The Academy of Natural Science of Philadelphia, 364 pp. https://doi.org/10.5962/bhl.title.6534" title="Manual of Conchology, Structure and Systematic, with Illustrations of the Species. Volume 1. The Academy of Natural Science of Philadelphia, 364 pp." url="https://doi.org/10.5962/bhl.title.6534" year="1885">Tryon 1885</bibRefCitation>
|
||
: 72, pl. 14, figs 9, 10.
|
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</paragraph>
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<paragraph id="65BF82B9D1CD4814246F417C0F1B0C63" pageId="0" pageNumber="39">
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<taxonomicName id="C960CF14B9E345024004BF5D93D1B3B3" authorityName="Stoliczka" authorityYear="1871" class="Gastropoda" family="Streptaxidae" genus="Streptaxis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Streptaxis hanleyanus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="hanleyanus">Streptaxis hanleyanus</taxonomicName>
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Stoliczka, 1871: 168, 169, pl. 7, fig. 15. Type locality: Prope Moulmein, ad flumen Attaran [Attaran River, Mawlamyine, Mon State, Myanmar].
|
||
<bibRefCitation id="3DF5D8D8AE4DB91FFADAC40B537D7270" DOI="https://doi.org/10.5962/bhl.title.13103" author="Blanford, WT" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="39" refId="B10" refString="Blanford, WT, Godwin-Austen, HH, 1908. The Fauna of British India, Including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae. Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.13103" title="The Fauna of British India, Including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae." url="https://doi.org/10.5962/bhl.title.13103" year="1908">Blanford and Godwin-Austen 1908</bibRefCitation>
|
||
: 8, 9.
|
||
</paragraph>
|
||
<paragraph id="44B9B0F8932E8B47DA52A87AE1EC4E15" pageId="0" pageNumber="39">
|
||
<taxonomicName id="394071B8E791A8AA536E0FDE243E7BF6" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius sankeyi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="sankeyi">Haploptychius sankeyi</taxonomicName>
|
||
-
|
||
<bibRefCitation id="5907EE9B2C56274CDCA013EA81B48709" author="Kobelt, W" journalOrPublisher="Journal of the Society for Bibliography of Natural History" pageId="0" pageNumber="39" refId="B37" refString="Kobelt, W, 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung: Streptaxidae und Daudebardiidae. Systematisches Conchylien-Cabinet von Martini und Chemnitz 1(12b) (2): 1-211 pls42-71 [pp. 1-96 pls 42-59 (1905) ; 97-211, pls 60-71 (1906)]." title="Die Raublungenschnecken (Agnatha). Zweite Abtheilung: Streptaxidae und Daudebardiidae. Systematisches Conchylien-Cabinet von Martini und Chemnitz 1 (12 b) (2): 1 - 211 pls 42 - 71 [pp. 1 - 96 pls 42 - 59 (1905); 97 - 211, pls 60 - 71 (1906)]." year="1905">Kobelt 1906</bibRefCitation>
|
||
: 147, pl. 57, figs 11, 12, pl. 62, figs 6, 7.
|
||
<bibRefCitation id="893DE347FD0C3A9C59DA50CEA9149661" author="Richardson, L" journalOrPublisher="Tryonia" pageId="0" pageNumber="39" pagination="1 - 326" refId="B70" refString="Richardson, L, 1988. Streptaxacea: Catalog of species, Part I, Streptaxidae. Tryonia 16: 1 - 326" title="Streptaxacea: Catalog of species, Part I, Streptaxidae." volume="16" year="1988">Richardson 1988</bibRefCitation>
|
||
: 220.
|
||
<bibRefCitation id="8A533AF3C0539D3A53306BD111B9350C" author="Siriboon, T" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="39" pagination="161 - 174" refId="B77" refString="Siriboon, T, Sutcharit, C, Naggs, F, Rowson, B, Panha, S, 2014b. Revision of the carnivorous snail genus Indoartemon Forcart, 1946 and a new genus Carinartemis from Thailand (Pulmonata: Streptaxidae). The Raffles Bulletin of Zoology 62: 161 - 174" title="Revision of the carnivorous snail genus Indoartemon Forcart, 1946 and a new genus Carinartemis from Thailand (Pulmonata: Streptaxidae)." volume="62" year="2014 b">Siriboon et al. 2014b</bibRefCitation>
|
||
: 169.
|
||
</paragraph>
|
||
<paragraph id="052DF95C5B8383456B636253C3E08918" pageId="0" pageNumber="39">
|
||
<taxonomicName id="01A24B9EC299D92E47F06925DA692AB5" class="Gastropoda" family="Streptaxidae" genus="Oophana" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Oophana hanleyana" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="hanleyana">Oophana hanleyana</taxonomicName>
|
||
-
|
||
<bibRefCitation id="E582E3048FB36E7C304365499FBC8196" author="Richardson, L" journalOrPublisher="Tryonia" pageId="0" pageNumber="39" pagination="1 - 326" refId="B70" refString="Richardson, L, 1988. Streptaxacea: Catalog of species, Part I, Streptaxidae. Tryonia 16: 1 - 326" title="Streptaxacea: Catalog of species, Part I, Streptaxidae." volume="16" year="1988">Richardson 1988</bibRefCitation>
|
||
: 235.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E30124FD962C5B5F7056AAA5811B20E1" pageId="0" pageNumber="39" type="materials_examined">
|
||
<paragraph id="6132AB4DFD8AEE3081ED2C301B39ADDD" pageId="0" pageNumber="39">Material examined.</paragraph>
|
||
<paragraph id="80F75AA4A6781EED00F5E74543060672" pageId="0" pageNumber="39">
|
||
<materialsCitation id="4ADFA9CE0DECD500019D2EEA66E0EE4F" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Moulmein. Moulmein, Hawkins, Blanford, Godwin-Austen, Acc. No., Oldham, Paboung Toung, Hungerford, Mergui, Cuming, Kwengan Hill" country="Myanmar" county="Mawlamyine Township" elevation="600" latitude="16.50639" location="Saddan Cave" longLatPrecision="1" longitude="97.81009" municipality="Dhammatat Cave" specimenCount="1738" stateProvince="Mon State" typeStatus="Syntype">
|
||
<emphasis id="F0494C5F3BA4AB31C0CAFFE99113DC70" bold="true" italics="true" pageId="0" pageNumber="39">
|
||
<typeStatus id="EBEF6685016B21E7F4BA8B4001D3EA25">Syntype</typeStatus>
|
||
</emphasis>
|
||
UMZC I.102740 (6 shells; Fig.
|
||
<figureCitation id="42019D66608DF75CEB61852184AC4B87" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Carinartemis sankeyi A syntype UMZC I. 102740 from Moulmein B specimen NHMUK 1888.12.4.792 - 794 from Moulmein C specimen NHMUK 1906.2.2.341 from Moulmein with apertural dentition D specimen NHMUK 1871.9.23.61 from Moulmein E specimen NHMUK ex. Cuming collection from Mergui." figureDoi="10.3897/zookeys.1110.85399.figure19" httpUri="https://binary.pensoft.net/fig/710484" pageId="0" pageNumber="39">19A</figureCitation>
|
||
) from
|
||
<collectorName id="AD410234A4A7BBB6C0D3F16704FC8912">Moulmein. Moulmein</collectorName>
|
||
: NHMUK 1954.6.3.556 (1 shell) ex.
|
||
<collectorName id="72446ADA91EB06B7219DF4E258A14436">Hawkins</collectorName>
|
||
collection. NHMUK 1906.2.2.341 (4 shells; Fig.
|
||
<figureCitation id="F7F350AF1DB4FFD392BDBC1BDA25D699" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Carinartemis sankeyi A syntype UMZC I. 102740 from Moulmein B specimen NHMUK 1888.12.4.792 - 794 from Moulmein C specimen NHMUK 1906.2.2.341 from Moulmein with apertural dentition D specimen NHMUK 1871.9.23.61 from Moulmein E specimen NHMUK ex. Cuming collection from Mergui." figureDoi="10.3897/zookeys.1110.85399.figure19" httpUri="https://binary.pensoft.net/fig/710484" pageId="0" pageNumber="39">19C</figureCitation>
|
||
) ex.
|
||
<collectorName id="F0C85BFC5EA75109C9313D97F3CF355A">Blanford</collectorName>
|
||
collection. NHMUK 1872.9.3.15 (5 shells) ex.
|
||
<collectorName id="3E77AAB306A4E73F650ABC7C493EABE5">Godwin-Austen</collectorName>
|
||
collection. NHMUK 1903.7.1.4001 (1 shell; Fig.
|
||
<figureCitation id="D52F291FF741DF6D2504D6D96ED4E7FF" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Carinartemis sankeyi A specimen NHMUK 1903.7.1.4001 from Moulmein B specimen NHMUK 1888.12.4.781 - 783 from Kwengan Hill C specimen CUMZ 13031 from Dhammatat Cave, Mawlamyine, Mon State with apertural dentition D specimen CUMZ 13029 from Kayon Cave, Mawlamyine, Mon State with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure20" httpUri="https://binary.pensoft.net/fig/710485" pageId="0" pageNumber="39">20A</figureCitation>
|
||
) ex.
|
||
<collectorName id="E20FC843EE97CD7E53A1EA535C40985A">Godwin-Austen</collectorName>
|
||
collection. NHMUK
|
||
<collectorName id="C3B5702CB26C3F10D696A92F20359DE0">Acc. No.</collectorName>
|
||
<specimenCount id="50AFBFA4E9B9097ADD98D265EEC929C4" type="generic">1733 ex.</specimenCount>
|
||
<collectorName id="669E48DEBCFB140AFB959D80F4E37DD9">Oldham</collectorName>
|
||
collection (2 shells). NHMUK 1888.12.4.792-794 (3 shells; Fig.
|
||
<figureCitation id="2F7E8CA981D2D6A95CC40636D188F22A" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Carinartemis sankeyi A syntype UMZC I. 102740 from Moulmein B specimen NHMUK 1888.12.4.792 - 794 from Moulmein C specimen NHMUK 1906.2.2.341 from Moulmein with apertural dentition D specimen NHMUK 1871.9.23.61 from Moulmein E specimen NHMUK ex. Cuming collection from Mergui." figureDoi="10.3897/zookeys.1110.85399.figure19" httpUri="https://binary.pensoft.net/fig/710484" pageId="0" pageNumber="39">19B</figureCitation>
|
||
). NHMUK 1871.9.23.61 (1 shell; Fig.
|
||
<figureCitation id="04C682227396D36CC8CB1ECD408F07B3" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Carinartemis sankeyi A syntype UMZC I. 102740 from Moulmein B specimen NHMUK 1888.12.4.792 - 794 from Moulmein C specimen NHMUK 1906.2.2.341 from Moulmein with apertural dentition D specimen NHMUK 1871.9.23.61 from Moulmein E specimen NHMUK ex. Cuming collection from Mergui." figureDoi="10.3897/zookeys.1110.85399.figure19" httpUri="https://binary.pensoft.net/fig/710484" pageId="0" pageNumber="39">19D</figureCitation>
|
||
).
|
||
<collectorName id="235ECA2F673526C3E54D295D8CEDA7CA">Paboung Toung</collectorName>
|
||
, Burma: NHMUK 1891.3.17.577-578 (2 shells) ex.
|
||
<collectorName id="005A45DD4D209DF2282612133FDBF723">Hungerford</collectorName>
|
||
collection.
|
||
<collectorName id="34165FCD3F2FF3AC00971D26C5578FA6">Mergui</collectorName>
|
||
: NHMUK 20210051 (1 shell; Fig.
|
||
<figureCitation id="47D4C1CB63195713A2D884FE81A93B6F" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Carinartemis sankeyi A syntype UMZC I. 102740 from Moulmein B specimen NHMUK 1888.12.4.792 - 794 from Moulmein C specimen NHMUK 1906.2.2.341 from Moulmein with apertural dentition D specimen NHMUK 1871.9.23.61 from Moulmein E specimen NHMUK ex. Cuming collection from Mergui." figureDoi="10.3897/zookeys.1110.85399.figure19" httpUri="https://binary.pensoft.net/fig/710484" pageId="0" pageNumber="39">19E</figureCitation>
|
||
) ex.
|
||
<collectorName id="53799CD1B6D0AC9E43359F3D9BAE5886">Cuming</collectorName>
|
||
collection.
|
||
<collectorName id="AC28F00EFD3CF64208EC0CC3E7A2CD7F">Kwengan Hill</collectorName>
|
||
: NHMUK 1888.12.4.781-783 (3 shells; Fig.
|
||
<figureCitation id="8689888CDF74929C6CF4607866E0AA83" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Carinartemis sankeyi A specimen NHMUK 1903.7.1.4001 from Moulmein B specimen NHMUK 1888.12.4.781 - 783 from Kwengan Hill C specimen CUMZ 13031 from Dhammatat Cave, Mawlamyine, Mon State with apertural dentition D specimen CUMZ 13029 from Kayon Cave, Mawlamyine, Mon State with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure20" httpUri="https://binary.pensoft.net/fig/710485" pageId="0" pageNumber="39">20B</figureCitation>
|
||
) ex.
|
||
<collectorName id="519B586D43D6CDDA43F519217020BC71">Blanford</collectorName>
|
||
collection.
|
||
<location id="F193B099C5BAE57F9C73D6E274760D4E" LSID="urn:lsid:plazi:treatment:A9B3BC261C09572FA717DFAC7ACCC058:F193B099C5BAE57F9C73D6E274760D4E" country="Myanmar" county="Mawlamyine Township" latitude="16.50639" longLatPrecision="1" longitude="97.81009" municipality="Dhammatat Cave" name="Saddan Cave" stateProvince="Mon State">Saddan Cave</location>
|
||
situated on same karsts ~
|
||
<quantity id="0D69FBC7B1689EB76EEE7DEE5E6C8A0D" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
|
||
<elevation id="FD3B7BD5694FCB2787C7AA87F34E6B11" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</elevation>
|
||
</quantity>
|
||
south of
|
||
<location id="9B05CD7AAA3E378E2DDB98268008E159" LSID="urn:lsid:plazi:treatment:A9B3BC261C09572FA717DFAC7ACCC058:9B05CD7AAA3E378E2DDB98268008E159" country="Myanmar" county="Mawlamyine Township" latitude="16.50639" longLatPrecision="1" longitude="97.81009" municipality="Dhammatat Cave" name="Kayon Cave" stateProvince="Mon State">Kayon Cave</location>
|
||
,
|
||
<location id="40B4E835C8811C31007E04290AB02E12" LSID="urn:lsid:plazi:treatment:A9B3BC261C09572FA717DFAC7ACCC058:40B4E835C8811C31007E04290AB02E12" country="Myanmar" county="Mawlamyine Township" latitude="16.50639" longLatPrecision="1" longitude="97.81009" municipality="Dhammatat Cave" name="Mawlamyine Township" stateProvince="Mon State">Mawlamyine Township</location>
|
||
,
|
||
<location id="E6FF1A1C9C5ABFE964EAF3DDA9367744" LSID="urn:lsid:plazi:treatment:A9B3BC261C09572FA717DFAC7ACCC058:E6FF1A1C9C5ABFE964EAF3DDA9367744" country="Myanmar" county="Mawlamyine Township" latitude="16.50639" longLatPrecision="1" longitude="97.81009" municipality="Dhammatat Cave" name="Mawlamyine District" stateProvince="Mon State">Mawlamyine District</location>
|
||
,
|
||
<collectingRegion id="ADFFA602976975A4ABA3E13B65D6145A" country="Myanmar" name="Mon">Mon State</collectingRegion>
|
||
,
|
||
<collectingCountry id="C06E7E8F7D521EF5DC28127D08943991" name="Myanmar">Myanmar</collectingCountry>
|
||
(
|
||
<geoCoordinate id="9D7DFF06FF61BD1FCB8882388DBBFDEA" degrees="16" direction="north" minutes="31" orientation="latitude" precision="1" seconds="42.8" value="16.528555">16°31'42.8"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6B4DFF1974A452AF79EED15C65CAD15D" degrees="97" direction="east" minutes="43" orientation="longitude" precision="1" seconds="2.1" value="97.71725">97°43'2.1"E</geoCoordinate>
|
||
): CUMZ 13028 (7 shells).
|
||
<location id="1A9D6215B52F0D3EA8EC2CBECDA57314" LSID="urn:lsid:plazi:treatment:A9B3BC261C09572FA717DFAC7ACCC058:1A9D6215B52F0D3EA8EC2CBECDA57314" country="Myanmar" county="Mawlamyine Township" latitude="16.50639" longLatPrecision="1" longitude="97.81009" municipality="Dhammatat Cave" name="Kayon Cave" stateProvince="Mon State">Kayon Cave</location>
|
||
[previously known as Farm Caves] ~
|
||
<quantity id="4F6385F719849FA327FF7A98B4FD57AB" metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
|
||
from
|
||
<location id="845C446BA7917703F86DFE759B52320C" LSID="urn:lsid:plazi:treatment:A9B3BC261C09572FA717DFAC7ACCC058:845C446BA7917703F86DFE759B52320C" country="Myanmar" county="Mawlamyine Township" latitude="16.50639" longLatPrecision="1" longitude="97.81009" municipality="Dhammatat Cave" name="Mawlamyine Township" stateProvince="Mon State">Mawlamyine Township</location>
|
||
,
|
||
<location id="2C09970112B093DD9E751CE91EE6DC6E" LSID="urn:lsid:plazi:treatment:A9B3BC261C09572FA717DFAC7ACCC058:2C09970112B093DD9E751CE91EE6DC6E" country="Myanmar" county="Mawlamyine Township" latitude="16.50639" longLatPrecision="1" longitude="97.81009" municipality="Dhammatat Cave" name="Mawlamyine District" stateProvince="Mon State">Mawlamyine District</location>
|
||
,
|
||
<collectingRegion id="AB0817FD1B7A6AC88186DDE0B69EBA42" country="Myanmar" name="Mon">Mon State</collectingRegion>
|
||
,
|
||
<collectingCountry id="01311F39852C3EEA79D0CCFAED3902EB" name="Myanmar">Myanmar</collectingCountry>
|
||
(
|
||
<geoCoordinate id="86B375471AE749C006DC775CEA1F801C" degrees="16" direction="north" minutes="32" orientation="latitude" precision="1" seconds="0.5" value="16.533472">16°32'0.5"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3A3FDFD79212C4D9CA0320B9B297C122" degrees="97" direction="east" minutes="42" orientation="longitude" precision="1" seconds="53.5" value="97.71486">97°42'53.5"E</geoCoordinate>
|
||
: CUMZ 13029 (13 shells; Fig.
|
||
<figureCitation id="9746362B8170ECB7979DEFEC24D3DA37" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Carinartemis sankeyi A specimen NHMUK 1903.7.1.4001 from Moulmein B specimen NHMUK 1888.12.4.781 - 783 from Kwengan Hill C specimen CUMZ 13031 from Dhammatat Cave, Mawlamyine, Mon State with apertural dentition D specimen CUMZ 13029 from Kayon Cave, Mawlamyine, Mon State with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure20" httpUri="https://binary.pensoft.net/fig/710485" pageId="0" pageNumber="39">20D</figureCitation>
|
||
), CUMZ 13030 (
|
||
<specimenCount id="F1CC3AA40D3C9F1985C0203C0DB52943" type="generic">5 specimens</specimenCount>
|
||
in ethanol).
|
||
<collectingMunicipality id="432175679AEA26EB450DCDCB4EC5355F">Dhammatat Cave</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="1DF4A94BD31704C444CEA5CD948EDBB4">Mawlamyine Township</collectingCounty>
|
||
,
|
||
<collectingRegion id="D4C8C130A86CCE1A2297B144BF658112" country="Myanmar" name="Mon">Mon State</collectingRegion>
|
||
,
|
||
<collectingCountry id="B07C9D3C74CCEB79F8275E2D0376D3BD" name="Myanmar">Myanmar</collectingCountry>
|
||
(
|
||
<geoCoordinate id="96B156F72157F02F6B62C4648E660AA2" degrees="16" direction="north" minutes="30" orientation="latitude" precision="1" seconds="23.0" value="16.50639">16°30'23.0"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1E44A9C324FA876A4E2ACBCFAB961BEB" degrees="97" direction="east" minutes="48" orientation="longitude" precision="1" seconds="36.3" value="97.81009">97°48'36.3"E</geoCoordinate>
|
||
): CUMZ 13031 (12 shells; Fig.
|
||
<figureCitation id="40C3F90B2B54295520BF7095F5DD2F01" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Carinartemis sankeyi A specimen NHMUK 1903.7.1.4001 from Moulmein B specimen NHMUK 1888.12.4.781 - 783 from Kwengan Hill C specimen CUMZ 13031 from Dhammatat Cave, Mawlamyine, Mon State with apertural dentition D specimen CUMZ 13029 from Kayon Cave, Mawlamyine, Mon State with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure20" httpUri="https://binary.pensoft.net/fig/710485" pageId="0" pageNumber="39">20C</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="CA3C3AD26547281AB827D3A837994B9F" doi="10.3897/zookeys.1110.85399.figure19" httpUri="https://binary.pensoft.net/fig/710484" pageId="0" pageNumber="39" start="Figure 19" startId="F19">
|
||
<paragraph id="E875E76A9CBB5EB226B127C84BB6E0AB" pageId="0" pageNumber="39">
|
||
<emphasis id="C1DCC642AFB3997229158B34684E1782" bold="true" pageId="0" pageNumber="39">Figure 19.</emphasis>
|
||
<taxonomicName id="319445DAF019A834812B3741F886D74B" baseAuthorityName="Benson" baseAuthorityYear="1859" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carinartemis sankeyi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="sankeyi">
|
||
<emphasis id="2F68B12A54A4B1C05F0FF9FFB5764ECA" italics="true" pageId="0" pageNumber="39">Carinartemis sankeyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F36CB1C5BAA1CB8BBC9A2A4652A4AFF5" bold="true" pageId="0" pageNumber="39">A</emphasis>
|
||
<typeStatus id="CC8C85D1269E7512AC8AAB0B4A38060D">syntype</typeStatus>
|
||
UMZC I.102740 from Moulmein
|
||
<emphasis id="779924E9A6293887725A093AC93E00A8" bold="true" pageId="0" pageNumber="39">B</emphasis>
|
||
specimen NHMUK 1888.12.4.792-794 from Moulmein
|
||
<emphasis id="0FEC852CC08251B994BD164D62034D5C" bold="true" pageId="0" pageNumber="39">C</emphasis>
|
||
specimen NHMUK 1906.2.2.341 from Moulmein with apertural dentition
|
||
<emphasis id="071DB2CF3F56A31331F4F21565E0C708" bold="true" pageId="0" pageNumber="39">D</emphasis>
|
||
specimen NHMUK 1871.9.23.61 from Moulmein
|
||
<emphasis id="DF49E732B732F7278C98A443FFB22AB9" bold="true" pageId="0" pageNumber="39">E</emphasis>
|
||
specimen NHMUK ex. Cuming collection from Mergui.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="D8FDBCA2D1486C10EE1FBEE338952283" doi="10.3897/zookeys.1110.85399.figure20" httpUri="https://binary.pensoft.net/fig/710485" pageId="0" pageNumber="39" start="Figure 20" startId="F20">
|
||
<paragraph id="521DDD2CF5615B1CC29F783FE042E8FE" pageId="0" pageNumber="39">
|
||
<emphasis id="634144552014D56983E883BBD0992E19" bold="true" pageId="0" pageNumber="39">Figure 20.</emphasis>
|
||
<taxonomicName id="BCAD71C52EFC32D04D475F7005DFC3E2" baseAuthorityName="Benson" baseAuthorityYear="1859" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carinartemis sankeyi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="sankeyi">
|
||
<emphasis id="DEAA6DF3BDC6AAAC5660D99B0C4B133F" italics="true" pageId="0" pageNumber="39">Carinartemis sankeyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2CE5140F4AA0E3C4320B07C337EEC024" bold="true" pageId="0" pageNumber="39">A</emphasis>
|
||
specimen NHMUK 1903.7.1.4001 from Moulmein
|
||
<emphasis id="5B9022F78686DF168F727CC016F97A0D" bold="true" pageId="0" pageNumber="39">B</emphasis>
|
||
specimen NHMUK 1888.12.4.781-783 from Kwengan Hill
|
||
<emphasis id="90343FE5B4541A5A961A07250A7B0035" bold="true" pageId="0" pageNumber="39">C</emphasis>
|
||
specimen CUMZ 13031 from Dhammatat Cave, Mawlamyine,
|
||
<collectingRegion id="3C80CB25CAABCB6739DEC5FBDC468AF8" country="Myanmar" name="Mon">Mon State</collectingRegion>
|
||
with apertural dentition
|
||
<emphasis id="109C80488A601D438931F8EAF796F23B" bold="true" pageId="0" pageNumber="39">D</emphasis>
|
||
specimen CUMZ 13029 from Kayon Cave, Mawlamyine,
|
||
<collectingRegion id="FD79F29D6AD4816486C5143D7BF50F19" country="Myanmar" name="Mon">Mon State</collectingRegion>
|
||
with apertural dentition.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="74858F6BE331F6951E1539F225611E0C" pageId="0" pageNumber="39" type="diagnosis">
|
||
<paragraph id="498655C60A767E1B4FF7F82DB4C06427" pageId="0" pageNumber="39">Diagnosis.</paragraph>
|
||
<paragraph id="738E585683CC6F929354C4FFBE778026" pageId="0" pageNumber="39">
|
||
<taxonomicName id="D7C7E929A76419AACE02D01F6F24D026" baseAuthorityName="Benson" baseAuthorityYear="1859" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carinartemis sankeyi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="sankeyi">
|
||
<emphasis id="EB0529E0693CBCEFD869FD7D0066508B" italics="true" pageId="0" pageNumber="39">Carinartemis sankeyi</emphasis>
|
||
</taxonomicName>
|
||
is superficially similar to
|
||
<taxonomicName id="621D9DFCF3F12EDE71FB0AF8DCDE5DB7" lsidName="C. vesperus" pageId="0" pageNumber="39" rank="species" species="vesperus">
|
||
<emphasis id="C28467CB2172156163AF435BF0828079" italics="true" pageId="0" pageNumber="39">C. vesperus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="62B1F38595B34946F7741CE039D4ECA7" lsidName="C. striatus" pageId="0" pageNumber="39" rank="species" species="striatus">
|
||
<emphasis id="A44C44F960C4546A1FA87D0D0A0ED200" italics="true" pageId="0" pageNumber="39">C. striatus</emphasis>
|
||
</taxonomicName>
|
||
in having a subquadrangular aperture, and less expanded and continuous peristome, slender atrium, vas deferens does not pass through a penial sheath, proximal penial hooks located on laterally flattened penial papillae, and distal penial hooks located on laterally compressed penial papillae separated by reticulated folds. In contrast, the shell of
|
||
<taxonomicName id="2FE502AD0DF5701A42CB3FE54C590908" lsidName="C. vesperus" pageId="0" pageNumber="39" rank="species" species="vesperus">
|
||
<emphasis id="F30E31126FBDB28FB3222BD1D24AFD2D" italics="true" pageId="0" pageNumber="39">C. vesperus</emphasis>
|
||
</taxonomicName>
|
||
has fine transverse ridges (nearly smooth) with few transverse ridges near peristome, varices absent, periphery more extended beyond the diameter of last whorl, lacking parietal lamella, and penial papillae absent. Meanwhile,
|
||
<taxonomicName id="D3D9F6FF67C51EA1E1EF50BD6E41F9A2" lsidName="C. striatus" pageId="0" pageNumber="39" rank="species" species="striatus">
|
||
<emphasis id="4D41569236BCC7AD3F7BFD5515C0D767" italics="true" pageId="0" pageNumber="39">C. striatus</emphasis>
|
||
</taxonomicName>
|
||
has vas deferens attached to the distal end of penial sheath with very thin connective tissue, and hooks located on papillae without connected longitudinal folds.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BC6005E1340C7CA901FC8E73ED7875BA" pageId="0" pageNumber="39" type="description">
|
||
<paragraph id="21E903001E4A0119B7FA6EC04652217A" pageId="0" pageNumber="39">Description.</paragraph>
|
||
<paragraph id="4CE9B81ECB04153C5612694911FBB37D" pageId="0" pageNumber="39">
|
||
Shell oblique-heliciform, white, translucent; whorls 6-7; spire conical with distinct suture. Shell surface glossy with fine transverse ridge across the entire shell; varices present. Embryonic shell ~
|
||
<normalizedToken id="85890DF981B2A392307CF49DD8AD1059" originalValue="2½">21/2</normalizedToken>
|
||
whorls with smooth surface; following whorls regularly coiled. Shell periphery wide and sharply keeled around nearly the entire penultimate whorl; last whorl axially deflected. Aperture subquadrangular; peristome continuous, thickened, expanded, and reflected. Apertural dentition with one strong parietal lamella. Umbilicus open and deep (Figs
|
||
<figureCitation id="EF99739D6D06F0D7DCD44C8329F6B188" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Carinartemis sankeyi A syntype UMZC I. 102740 from Moulmein B specimen NHMUK 1888.12.4.792 - 794 from Moulmein C specimen NHMUK 1906.2.2.341 from Moulmein with apertural dentition D specimen NHMUK 1871.9.23.61 from Moulmein E specimen NHMUK ex. Cuming collection from Mergui." figureDoi="10.3897/zookeys.1110.85399.figure19" httpUri="https://binary.pensoft.net/fig/710484" pageId="0" pageNumber="39">19</figureCitation>
|
||
,
|
||
<figureCitation id="AD2AB56FE2F31A164E4312E7452B3C1E" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Carinartemis sankeyi A specimen NHMUK 1903.7.1.4001 from Moulmein B specimen NHMUK 1888.12.4.781 - 783 from Kwengan Hill C specimen CUMZ 13031 from Dhammatat Cave, Mawlamyine, Mon State with apertural dentition D specimen CUMZ 13029 from Kayon Cave, Mawlamyine, Mon State with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure20" httpUri="https://binary.pensoft.net/fig/710485" pageId="0" pageNumber="39">20</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B52F898D94116EA121DC20DD22B5EE5F" pageId="0" pageNumber="39">
|
||
<emphasis id="3892F32A43DCD7864ECD568F0038E4F4" bold="true" pageId="0" pageNumber="39">
|
||
<emphasis id="C1DFDC5670E216329037C8E1675CD2CD" italics="true" pageId="0" pageNumber="39">Genital organs</emphasis>
|
||
.
|
||
</emphasis>
|
||
Atrium (at) short, thin and slender. Penis (p) very thin and long. Penial sheath (ps) thin and extending nearly entire length of penis; penial sheath retractor muscle (psr) thin, originating near genital orifice and attached to atrium with short and thin connective tissue, and inserting distally on penial sheath (Fig.
|
||
<figureCitation id="30EB16CE13C3F3E1619F620E627259C7" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Genital anatomy of A, B Carinartemis exacutus, specimen CUMZ 13021 A reproductive system and B insertion of vas deferens into penial sheath C, D Carinartemis sankeyi, specimen CUMZ 13029 C reproductive system and D insertion of vas deferens into penial sheath." figureDoi="10.3897/zookeys.1110.85399.figure21" httpUri="https://binary.pensoft.net/fig/710486" pageId="0" pageNumber="39">21C</figureCitation>
|
||
). Vas deferens (vd) runs downwards to curved portion, ~ 1/3 of the penial sheath length, without insertion before entering penis distally (Fig.
|
||
<figureCitation id="5809FC309B61202413F67F9D746AD474" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Genital anatomy of A, B Carinartemis exacutus, specimen CUMZ 13021 A reproductive system and B insertion of vas deferens into penial sheath C, D Carinartemis sankeyi, specimen CUMZ 13029 C reproductive system and D insertion of vas deferens into penial sheath." figureDoi="10.3897/zookeys.1110.85399.figure21" httpUri="https://binary.pensoft.net/fig/710486" pageId="0" pageNumber="39">21D</figureCitation>
|
||
). Penial retractor muscle (pr) thin, very long, inserting at penis and vas deferens junction.
|
||
</paragraph>
|
||
<caption id="612E7DE7FC11B602FBA7DCB7CABE295B" doi="10.3897/zookeys.1110.85399.figure21" httpUri="https://binary.pensoft.net/fig/710486" pageId="0" pageNumber="39" start="Figure 21" startId="F21">
|
||
<paragraph id="FF6324D2D399367745CEE0A844905DA3" pageId="0" pageNumber="39">
|
||
<emphasis id="07B7C0C9B4F031F98F76E89AECFB3269" bold="true" pageId="0" pageNumber="39">Figure 21.</emphasis>
|
||
Genital anatomy of
|
||
<emphasis id="5496EE7E8F8DA444B0C0E412EBBBD6B7" bold="true" pageId="0" pageNumber="39">A, B</emphasis>
|
||
<taxonomicName id="41894BBB303BCF4AA9F4938FFC033DE3" baseAuthorityName="Gould" baseAuthorityYear="1856" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carinartemis exacutus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="exacutus">
|
||
<emphasis id="DE82FBD701739752B8F8882AAE1F3C0B" italics="true" pageId="0" pageNumber="39">Carinartemis exacutus</emphasis>
|
||
</taxonomicName>
|
||
, specimen CUMZ 13021
|
||
<emphasis id="021E9AC16C3EEEBA5DB8F2FBA43C8580" bold="true" pageId="0" pageNumber="39">A</emphasis>
|
||
reproductive system and
|
||
<emphasis id="3C5993AE45EE5F0310D4315B92E83521" bold="true" pageId="0" pageNumber="39">B</emphasis>
|
||
insertion of vas deferens into penial sheath
|
||
<emphasis id="D54AD38566341DDC6FDB26CA8B1A9CB3" bold="true" pageId="0" pageNumber="39">C, D</emphasis>
|
||
<taxonomicName id="363A266FDDFB41DEDBE38A89A1334FF1" baseAuthorityName="Benson" baseAuthorityYear="1859" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carinartemis sankeyi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="sankeyi">
|
||
<emphasis id="442931BBE7F26DE63178221B5A641450" italics="true" pageId="0" pageNumber="39">Carinartemis sankeyi</emphasis>
|
||
</taxonomicName>
|
||
, specimen CUMZ 13029
|
||
<emphasis id="DCD2E82D81EF66702A745D6CBCB8A2CF" bold="true" pageId="0" pageNumber="39">C</emphasis>
|
||
reproductive system and
|
||
<emphasis id="2067A9B9906704A3A19DBF05EDEF98B0" bold="true" pageId="0" pageNumber="39">D</emphasis>
|
||
insertion of vas deferens into penial sheath.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="BB2F777E6C7799B7C1CC1A99FCEDA6D1" doi="10.3897/zookeys.1110.85399.figure22" httpUri="https://binary.pensoft.net/fig/710487" pageId="0" pageNumber="39" start="Figure 22" startId="F22">
|
||
<paragraph id="3505B1B1F29DB2EA6D8515639A42CE2E" pageId="0" pageNumber="39">
|
||
<emphasis id="1A7983059DF7AA7C747D62E5BA84DFE1" bold="true" pageId="0" pageNumber="39">Figure 22.</emphasis>
|
||
Internal sculpture of genitalia of
|
||
<taxonomicName id="2CC57754549CF78CA0829DC7A4D0680D" baseAuthorityName="Gould" baseAuthorityYear="1856" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carinartemis exacutus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="exacutus">
|
||
<emphasis id="828ECE082294D4D050366F9F1EB34843" italics="true" pageId="0" pageNumber="39">Carinartemis exacutus</emphasis>
|
||
</taxonomicName>
|
||
, specimen CUMZ 13021
|
||
<emphasis id="06AAE8ECBF7112344210F7BDD0C5E326" bold="true" pageId="0" pageNumber="39">A</emphasis>
|
||
atrium surface
|
||
<emphasis id="0DC66EF52AD09D560DB1C160EC1007D7" bold="true" pageId="0" pageNumber="39">B</emphasis>
|
||
overview of internal penial wall
|
||
<emphasis id="C3E5D8BD8BE9D43839731D2EF5939F3B" bold="true" pageId="0" pageNumber="39">C</emphasis>
|
||
arrangement of penial hooks on proximal part of penis
|
||
<emphasis id="72CCD138BF9496D73C213DCA10F60A70" bold="true" pageId="0" pageNumber="39">D</emphasis>
|
||
lateral view of penial hooks
|
||
<emphasis id="1390DEBBF566D3D4A97C613729286799" bold="true" pageId="0" pageNumber="39">E</emphasis>
|
||
arrangement of penial hooks on middle part of penis
|
||
<emphasis id="898EC466FD99E53E92500E415B0A1A1A" bold="true" pageId="0" pageNumber="39">F</emphasis>
|
||
lateral view penial hooks
|
||
<emphasis id="6507A4001546B12484FD9CC1896CCBF4" bold="true" pageId="0" pageNumber="39">G</emphasis>
|
||
arrangement of longitudinal oblique vaginal folds.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="73871672961EC26E56C6467B7EA97D8D" pageId="0" pageNumber="39">
|
||
Internal wall of atrium generally smooth with sparse atrial pores (Fig.
|
||
<figureCitation id="2AAE5A2ABEE0BA11C8A33B909447D86B" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Internal sculpture of genitalia of Carinartemis sankeyi, specimen CUMZ 13029 A atrium surface B arrangement of penial hooks on proximal part of penis C lateral view of exposed proximal penial hooks D penial hooks on middle part of penis E arrangement of penial hooks on distal part of penis F uncovered distal penial hooks with pointed tip G distal penial hooks embedded in penial papillae H vaginal surface with vaginal pores." figureDoi="10.3897/zookeys.1110.85399.figure23" httpUri="https://binary.pensoft.net/fig/710488" pageId="0" pageNumber="39">23A</figureCitation>
|
||
). Proximal penial wall covered with scattered and pale brownish penial hooks, ~ 12 hooks/200
|
||
<normalizedToken id="500257FE0927C77CF4B872FB01760275" originalValue="μm">μm</normalizedToken>
|
||
2. Proximal penial hooks located on laterally flattened penial papillae; hooks small and short (<0.03 mm in length), slightly expanded at base, tips obtuse and curved towards genital orifice (Fig.
|
||
<figureCitation id="4817880A622ACBAC0C57DAC0B54640AD" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Internal sculpture of genitalia of Carinartemis sankeyi, specimen CUMZ 13029 A atrium surface B arrangement of penial hooks on proximal part of penis C lateral view of exposed proximal penial hooks D penial hooks on middle part of penis E arrangement of penial hooks on distal part of penis F uncovered distal penial hooks with pointed tip G distal penial hooks embedded in penial papillae H vaginal surface with vaginal pores." figureDoi="10.3897/zookeys.1110.85399.figure23" httpUri="https://binary.pensoft.net/fig/710488" pageId="0" pageNumber="39">23B, C</figureCitation>
|
||
). Middle and distal penial walls densely covered with pale brownish hooks, ~ 20 hooks/200
|
||
<normalizedToken id="1DBEDC9B986123264917F370B9EF6102" originalValue="μm">μm</normalizedToken>
|
||
2. Middle and distal hooks located on laterally compressed penial papillae separated by reticulated folds; hooks small, short (<0.01 mm in length), slightly expanded at base, and tips pointed (Fig.
|
||
<figureCitation id="D75678BEA97473FD29103AF6B42C1054" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Internal sculpture of genitalia of Carinartemis sankeyi, specimen CUMZ 13029 A atrium surface B arrangement of penial hooks on proximal part of penis C lateral view of exposed proximal penial hooks D penial hooks on middle part of penis E arrangement of penial hooks on distal part of penis F uncovered distal penial hooks with pointed tip G distal penial hooks embedded in penial papillae H vaginal surface with vaginal pores." figureDoi="10.3897/zookeys.1110.85399.figure23" httpUri="https://binary.pensoft.net/fig/710488" pageId="0" pageNumber="39">23D-G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="4ED0D8E33BE31DD8C261983046A9F4C0" doi="10.3897/zookeys.1110.85399.figure23" httpUri="https://binary.pensoft.net/fig/710488" pageId="0" pageNumber="39" start="Figure 23" startId="F23">
|
||
<paragraph id="2D60A58642413AE640D1A8D159D5FE91" pageId="0" pageNumber="39">
|
||
<emphasis id="A4CDF4A9945B32E0A591CEECBBBF79BA" bold="true" pageId="0" pageNumber="39">Figure 23.</emphasis>
|
||
Internal sculpture of genitalia of
|
||
<taxonomicName id="3C3154325926C366B2F7FDD8D156F783" baseAuthorityName="Benson" baseAuthorityYear="1859" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carinartemis sankeyi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="sankeyi">
|
||
<emphasis id="DAA26C67F32B2FB3EA0B3B4CA5FC6BA9" italics="true" pageId="0" pageNumber="39">Carinartemis sankeyi</emphasis>
|
||
</taxonomicName>
|
||
, specimen CUMZ 13029
|
||
<emphasis id="731B21AEDCF7DCDB7D331AE0FA0F19A5" bold="true" pageId="0" pageNumber="39">A</emphasis>
|
||
atrium surface
|
||
<emphasis id="EFA93EBF1D3616EE755487F006C501A9" bold="true" pageId="0" pageNumber="39">B</emphasis>
|
||
arrangement of penial hooks on proximal part of penis
|
||
<emphasis id="F79E1459BD03024E48F393C0871963FB" bold="true" pageId="0" pageNumber="39">C</emphasis>
|
||
lateral view of exposed proximal penial hooks
|
||
<emphasis id="B4CD5E12E426736229CC031AC92C6EB8" bold="true" pageId="0" pageNumber="39">D</emphasis>
|
||
penial hooks on middle part of penis
|
||
<emphasis id="9692D99A3549615B8819592849CF2AAC" bold="true" pageId="0" pageNumber="39">E</emphasis>
|
||
arrangement of penial hooks on distal part of penis
|
||
<emphasis id="9ED2225FD26E7B42291498C415E97D15" bold="true" pageId="0" pageNumber="39">F</emphasis>
|
||
uncovered distal penial hooks with pointed tip
|
||
<emphasis id="237C02B1F1488DF63503C3057B08CEFC" bold="true" pageId="0" pageNumber="39">G</emphasis>
|
||
distal penial hooks embedded in penial papillae
|
||
<emphasis id="834A7F17910BE232A5C9C733DFCC89EF" bold="true" pageId="0" pageNumber="39">H</emphasis>
|
||
vaginal surface with vaginal pores.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A2BC19AAC38CB516F8FD25A64113091C" pageId="0" pageNumber="39">
|
||
Vagina (v) short, stout and ~ 1/5 of penis length. Gametolytic duct (gd) a long tube extending as far as albumin gland; gametolytic sac (gs) ovate. Proximal free oviduct (fo) enlarged then tapering to smaller diameter in middle section, and slightly enlarged distally. Oviduct (ov) enlarged and folded; prostate gland inconspicuous and bound to oviduct. Talon (ta) small, short and club shaped. Hermaphroditic duct (hd) bearing long seminal vesicle (sv) ca. the same length as from talon to branching point of seminal vesicle (Fig.
|
||
<figureCitation id="E490EC87A3AD10D48B0C7C47BD2AB52B" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Genital anatomy of A, B Carinartemis exacutus, specimen CUMZ 13021 A reproductive system and B insertion of vas deferens into penial sheath C, D Carinartemis sankeyi, specimen CUMZ 13029 C reproductive system and D insertion of vas deferens into penial sheath." figureDoi="10.3897/zookeys.1110.85399.figure21" httpUri="https://binary.pensoft.net/fig/710486" pageId="0" pageNumber="39">21C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B74543BB8388966C8AE590B0CA66C048" pageId="0" pageNumber="39">
|
||
Vaginal wall generally smooth with vaginal pores, and vaginal hook absent (Fig.
|
||
<figureCitation id="BF60504014CC3CA00833311A32E35F3D" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Internal sculpture of genitalia of Carinartemis sankeyi, specimen CUMZ 13029 A atrium surface B arrangement of penial hooks on proximal part of penis C lateral view of exposed proximal penial hooks D penial hooks on middle part of penis E arrangement of penial hooks on distal part of penis F uncovered distal penial hooks with pointed tip G distal penial hooks embedded in penial papillae H vaginal surface with vaginal pores." figureDoi="10.3897/zookeys.1110.85399.figure23" httpUri="https://binary.pensoft.net/fig/710488" pageId="0" pageNumber="39">23H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1C7066E74E30A393382C72A8B3506B99" pageId="0" pageNumber="39" type="distribution">
|
||
<paragraph id="4525FF4699095FACAF6368AE27021233" pageId="0" pageNumber="39">Distribution.</paragraph>
|
||
<paragraph id="7CEF5003625489B8F9436E0B7FB79091" pageId="0" pageNumber="39">This species occurs from three localities in Mon State, southern Myanmar and is likely to be endemic to this area.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F3E0F71C82C9F569A8077EB463CE0AFC" pageId="0" pageNumber="39" type="radula">
|
||
<paragraph id="37EE960311C4AAA9B264ED0131CB3453" pageId="0" pageNumber="39">
|
||
<taxonomicName id="BCB4AAA01F1EA156B632F367546ED7DA" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Plantae" lsidName="Radula" order="Porellales" pageId="0" pageNumber="39" phylum="Marchantiophyta" rank="genus">Radula</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="2A1DFFBC54F124A117E2A1796408C111" pageId="0" pageNumber="39">
|
||
Each row consists of 43-49 teeth with formula (21-24)-1-(21-24). Central tooth small with pointed cusp. Lateral and marginal teeth undifferentiated, unicuspid and lanceolate. Latero-marginal teeth gradually reduced in size, with outermost teeth much smaller and shorter than inner teeth (Fig.
|
||
<figureCitation id="AA1F306E290476273C3858C2A2C8BD54" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Radula of specimens A Discartemon tonywhitteni, paratype CUMZ 13001 B Discartemon paurodeviatus sp. nov., paratype CUMZ 13003 C Haploptychius heliakosus sp. nov., paratype CUMZ 13014 D Carinartemis exacutus, specimen CUMZ 13021 E, F Carinartemis sankeyi, specimen CUMZ 13029 E overview of radula and F high magnification of central teeth." figureDoi="10.3897/zookeys.1110.85399.figure24" httpUri="https://binary.pensoft.net/fig/710489" pageId="0" pageNumber="39">24D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="ADF881F1646AE9BA79076174346CD004" doi="10.3897/zookeys.1110.85399.figure24" httpUri="https://binary.pensoft.net/fig/710489" pageId="0" pageNumber="39" start="Figure 24" startId="F24">
|
||
<paragraph id="B6C05AEA9F6647C779268B15A865F0C2" pageId="0" pageNumber="39">
|
||
<emphasis id="C5EAD58FBC989D1B80282AF5C11C8EFC" bold="true" pageId="0" pageNumber="39">Figure 24.</emphasis>
|
||
<taxonomicName id="42D66E7865EA7880CE6FD2F459C411C7" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Plantae" lsidName="Radula" order="Porellales" pageId="0" pageNumber="39" phylum="Marchantiophyta" rank="genus">Radula</taxonomicName>
|
||
of specimens
|
||
<emphasis id="8FCFE1F96C2FBEC01F54C1444F109519" bold="true" pageId="0" pageNumber="39">A</emphasis>
|
||
<taxonomicName id="D01346611BC320741B667852309C9445" authorityName="Sutcharit & Panha" authorityYear="2020" class="Gastropoda" family="Streptaxidae" genus="Discartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discartemon tonywhitteni" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="tonywhitteni">
|
||
<emphasis id="8AA1B138C446FED7B220FE40D195EADD" italics="true" pageId="0" pageNumber="39">Discartemon tonywhitteni</emphasis>
|
||
</taxonomicName>
|
||
, paratype CUMZ 13001
|
||
<emphasis id="49217B6CC28C4668741D467642F28F7F" bold="true" pageId="0" pageNumber="39">B</emphasis>
|
||
<taxonomicName id="374706C1479516EE058039DD32D53FD5" authorityName="Man & Panha" authorityYear="2022" class="Gastropoda" family="Streptaxidae" genus="Discartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discartemon paurodeviatus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="paurodeviatus">
|
||
<emphasis id="CA45107B5A3F2BBB7D76F10521C8921E" italics="true" pageId="0" pageNumber="39">Discartemon paurodeviatus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype CUMZ 13003
|
||
<emphasis id="498C4041198496ADBE51615AE38B439F" bold="true" pageId="0" pageNumber="39">C</emphasis>
|
||
<taxonomicName id="132F30A1479310F8C71AAA42696DF894" authorityName="Man & Panha" authorityYear="2022" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius heliakosus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="heliakosus">
|
||
<emphasis id="F0C32A9D1B8FF688EE6B1512EE5186E4" italics="true" pageId="0" pageNumber="39">Haploptychius heliakosus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype CUMZ 13014
|
||
<emphasis id="59C5EE07430EE30986A2EBB49320E073" bold="true" pageId="0" pageNumber="39">D</emphasis>
|
||
<taxonomicName id="15184140A2233A47AFB3881EFC72FCD4" baseAuthorityName="Gould" baseAuthorityYear="1856" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carinartemis exacutus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="exacutus">
|
||
<emphasis id="14339B17ED051B3E776353644296F0E0" italics="true" pageId="0" pageNumber="39">Carinartemis exacutus</emphasis>
|
||
</taxonomicName>
|
||
, specimen CUMZ 13021
|
||
<emphasis id="E0F91A7671FBBF08A5EDE4B50888D0B3" bold="true" pageId="0" pageNumber="39">E, F</emphasis>
|
||
<taxonomicName id="4CEAC6E9A5D7715E37FBFCB402CD784F" baseAuthorityName="Benson" baseAuthorityYear="1859" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carinartemis sankeyi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="sankeyi">
|
||
<emphasis id="B43939D137A2260C764BBE4B236B5B47" italics="true" pageId="0" pageNumber="39">Carinartemis sankeyi</emphasis>
|
||
</taxonomicName>
|
||
, specimen CUMZ 13029
|
||
<emphasis id="2CA5FF3EC1A31CBC06A909FB1519D1EB" bold="true" pageId="0" pageNumber="39">E</emphasis>
|
||
overview of radula and
|
||
<emphasis id="CB9E15E901593747427F6B0C410C69B7" bold="true" pageId="0" pageNumber="39">F</emphasis>
|
||
high magnification of central teeth.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="67B17F93637715C787F4D731FA2374BE" pageId="0" pageNumber="39" type="remarks">
|
||
<paragraph id="4A12498C0496C98079CCA027A1C95733" pageId="0" pageNumber="39">Remarks.</paragraph>
|
||
<paragraph id="DE8CCE82821C621A9C1C967265899C67" pageId="0" pageNumber="39">
|
||
<bibRefCitation id="216BAFEE41B8A8463B07543A5DD0C57A" author="Benson, WH" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="39" refId="B5" refString="Benson, WH, 1859a. Characters of a new Burmese Streptaxis, and of two forms belonging to a peculiar section of Helix collected by Capt. Richard H. Sankey, Madras Engineers. Annals and Magazine of Natural History, Series 3, Volume 3: 471-474." title="Characters of a new Burmese Streptaxis, and of two forms belonging to a peculiar section of Helix collected by Capt. Richard H. Sankey, Madras Engineers. Annals and Magazine of Natural History, Series 3, Volume 3: 471 - 474." year="1859 a">Benson (1859a)</bibRefCitation>
|
||
introduced
|
||
<taxonomicName id="F01AD2A5C9BDDC8394EB19DEA3A50CC3" lsidName="C. sankeyi" pageId="0" pageNumber="39" rank="species" species="sankeyi">
|
||
<emphasis id="9B44B975AF9BB677D2D0D2AE3A0EBEAE" italics="true" pageId="0" pageNumber="39">C. sankeyi</emphasis>
|
||
</taxonomicName>
|
||
based on a single specimen from
|
||
<normalizedToken id="FE59FDEF1C2885B156F97F18B3373AF3" originalValue="‘Moulmein’">'Moulmein'</normalizedToken>
|
||
. Later,
|
||
<bibRefCitation id="6915A504C8EC9ECCE377A1B58AEF9DF4" author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="39" pagination="143 - 177" refId="B80" refString="Stoliczka, F, 1871. Notes on the terrestrial Mollusca from the neighbourhood of Moulmein, with descriptions of new species. Journal of the Asiatic Society of Bengal 40: 143 - 177" title="Notes on the terrestrial Mollusca from the neighbourhood of Moulmein, with descriptions of new species." volume="40" year="1871">Stoliczka (1871)</bibRefCitation>
|
||
provided some diagnostic characters with a description of the soft body color and suggested that the Farm-Cave is possibly the correct type locality. This historical locality name refers to a group of caves located on a karst ridge, namely Kayon Hill, situated on the west bank of the Attaran River. At the same time,
|
||
<bibRefCitation id="EBD331A0A193BD04F4A0586451C88431" author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="39" pagination="143 - 177" refId="B80" refString="Stoliczka, F, 1871. Notes on the terrestrial Mollusca from the neighbourhood of Moulmein, with descriptions of new species. Journal of the Asiatic Society of Bengal 40: 143 - 177" title="Notes on the terrestrial Mollusca from the neighbourhood of Moulmein, with descriptions of new species." volume="40" year="1871">Stoliczka (1871)</bibRefCitation>
|
||
proposed
|
||
<taxonomicName id="558CB234EB81581278AB462C8963F6A8" authorityName="Stoliczka" authorityYear="1871" class="Gastropoda" family="Streptaxidae" genus="Streptaxis" kingdom="Animalia" lsidName="Streptaxis hanleyanus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="hanleyanus">
|
||
<emphasis id="DF6D797323EB25B4357713FE2FA9E83F" italics="true" pageId="0" pageNumber="39">Streptaxis hanleyanus</emphasis>
|
||
</taxonomicName>
|
||
based on a single specimen with a small and depressed shell, wide umbilicus, and almost rectangular aperture. However, this holotype specimen was collected from the same geographical area as
|
||
<taxonomicName id="3BFD9B53F49A41839C03884B9E08F038" lsidName="C. sankeyi" pageId="0" pageNumber="39" rank="species" species="sankeyi">
|
||
<emphasis id="69B6DF9FC04ACC811758714C310F46CA" italics="true" pageId="0" pageNumber="39">C. sankeyi</emphasis>
|
||
</taxonomicName>
|
||
. Therefore, we consider this nominal species as a smaller shell form and so treat it as a junior synonym with
|
||
<taxonomicName id="4D452865FB10F7A3E19BCE6B643F5A02" lsidName="C. sankeyi" pageId="0" pageNumber="39" rank="species" species="sankeyi">
|
||
<emphasis id="8B165951F1D478F28D83F3C2EF8A229C" italics="true" pageId="0" pageNumber="39">C. sankeyi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D3B9122AEB8B4151B46F00344B6F6A78" pageId="0" pageNumber="39">
|
||
The original spelling of this nominal species was
|
||
<taxonomicName id="B730B21C66D4004A2011D726CDDFCCE7" baseAuthorityName="Benson" baseAuthorityYear="1859" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carinartemis sankeyi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="sankeyi">
|
||
<emphasis id="CB9B2D151F7CD29E3EC41A4E9BEAD63F" italics="true" pageId="0" pageNumber="39">Carinartemis sankeyi</emphasis>
|
||
</taxonomicName>
|
||
, which was intentionally modified to
|
||
<taxonomicName id="60810F338362392362581E6FB743A107" authorityName="Stoliczka" authorityYear="1871" class="Gastropoda" family="Streptaxidae" genus="Streptaxis" kingdom="Animalia" lsidName="Streptaxis sankeyanus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="sankeyanus">
|
||
<emphasis id="F4BEF65871B62F8C9B5F4BDDADD80F59" italics="true" pageId="0" pageNumber="39">Streptaxis sankeyanus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="6333FF7E68E0BC21C53FCDDD52FCD430" author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="39" pagination="143 - 177" refId="B80" refString="Stoliczka, F, 1871. Notes on the terrestrial Mollusca from the neighbourhood of Moulmein, with descriptions of new species. Journal of the Asiatic Society of Bengal 40: 143 - 177" title="Notes on the terrestrial Mollusca from the neighbourhood of Moulmein, with descriptions of new species." volume="40" year="1871">Stoliczka (1871)</bibRefCitation>
|
||
without a clear reason. Therefore, this unjustified emendation name was made available with its authorship and date and became a junior objective synonym (ICZN 1999: Arts 32.3, 33.2.3, 50.5).
|
||
</paragraph>
|
||
<paragraph id="229B73A76ECE8D430A5241D620ABBF6D" pageId="0" pageNumber="39">
|
||
Recently, the population collected from Dhammatat Cave showed a more rectangular aperture, slightly compact penultimate whorl, and larger parietal lamella (Fig.
|
||
<figureCitation id="9E0D7F9E880580B670053AB8456797FF" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Carinartemis sankeyi A specimen NHMUK 1903.7.1.4001 from Moulmein B specimen NHMUK 1888.12.4.781 - 783 from Kwengan Hill C specimen CUMZ 13031 from Dhammatat Cave, Mawlamyine, Mon State with apertural dentition D specimen CUMZ 13029 from Kayon Cave, Mawlamyine, Mon State with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure20" httpUri="https://binary.pensoft.net/fig/710485" pageId="0" pageNumber="39">20C</figureCitation>
|
||
), but this species could only be examined from the empty shells. Therefore, we have to conclude that these differences are an intraspecific variation until more specimens are available, or at least the genitalia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |