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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1190.109539" ID-Pensoft-Pub="1313-2970-1190-39" ID-Pensoft-UUID="D1C6B72C21B050CCA36EF15C44F982C2" ID-ZooBank="9CE8F6C628B74CDA8140BA78B59FF68F" ModsDocID="1313-2970-1190-39" checkinTime="1705976937398" checkinUser="pensoft" docAuthor="Li, Yihang, Li, Haoyuan &amp; Shi, Hongliang" docDate="2024" docId="65D41756CD6153C483504D9630DFF938" docLanguage="en" docName="ZooKeys 1190: 39-73" docOrigin="ZooKeys 1190" docPubDate="2024-01-22" docSource="http://dx.doi.org/10.3897/zookeys.1190.109539" docTitle="Amara (Curtonotus) hyperborea Dejean 1831" docType="treatment" docVersion="1" id="D1C6B72C21B050CCA36EF15C44F982C2" lastPageNumber="39" masterDocId="D1C6B72C21B050CCA36EF15C44F982C2" masterDocTitle="Revision of the macropterous subgenus Curtonotus from east China, with the description of a new species (Carabidae, Zabrini, Amara)" masterLastPageNumber="73" masterPageNumber="39" pageNumber="39" updateTime="1705976937398" updateUser="pensoft">
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<mods:title id="4FCEDBCFB1A82160E73744A70F93612B">Revision of the macropterous subgenus Curtonotus from east China, with the description of a new species (Carabidae, Zabrini, Amara)</mods:title>
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<mods:namePart id="2AFDF574C5916AF27E854DDA866BA6FA">Li, Yihang</mods:namePart>
<mods:nameIdentifier id="354333AF13B0930A2B5E1F8A953BAA1C" type="ORCID">https://orcid.org/0000-0001-5677-0440</mods:nameIdentifier>
<mods:affiliation id="9236FBF22CC0BB7DFBDA8D492AE0BA91">College of Agriculture, Purdue University, West Lafayette, Indiana 47906, USA</mods:affiliation>
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<mods:namePart id="3994A58649676070DA2CD873853437F1">Li, Haoyuan</mods:namePart>
<mods:nameIdentifier id="2C1378C782726E8567C7E63B7BF96B1E" type="ORCID">https://orcid.org/0009-0005-0314-9542</mods:nameIdentifier>
<mods:affiliation id="E8C9F9C57E361B7283D228D1E804C526">College of Life Science, Capital Normal University, Beijing 100048, China</mods:affiliation>
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<mods:namePart id="85BE9A10D57954AF4023A1C15B34D87F">Shi, Hongliang</mods:namePart>
<mods:nameIdentifier id="59114BF612D4F8CACD4BBD98E5E9AD70" type="ORCID">https://orcid.org/0000-0002-9989-5830</mods:nameIdentifier>
<mods:affiliation id="C8C2CEC9846974D98C9A8433E90F4FC0">College of Forestry, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
<mods:nameIdentifier id="91C4EB7EB50DC622BFC84A2033897906" type="email">shihl@bjfu.edu.cn</mods:nameIdentifier>
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<mods:date id="D9913F35E8B97A22E1466EA92135CC56">2024</mods:date>
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<subSubSection id="1475CAD9A695EB871E2067BE29C6C449" pageId="0" pageNumber="39" type="nomenclature">
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<taxonomicName id="9438F179610165DD93B374E347A36806" ID-CoL="CFBV" LSID="65D41756-CD61-53C4-8350-4D9630DFF938" authority="Dejean, 1831" authorityName="Dejean" authorityYear="1831" class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara (Curtonotus) hyperborea" order="Coleoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="hyperborea" subGenus="Curtonotus">Amara (Curtonotus) hyperborea Dejean, 1831</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="8AC84787BFA7ED7768EED5762BD9E923" pageId="0" pageNumber="39" type="reference_group">
<paragraph id="016C66544C3745CADCD554D46BE73E4C" pageId="0" pageNumber="39">
<taxonomicName id="3B7B41CA692F13D4E60F84D8B217B64D" authorityName="Dejean" authorityYear="1831" class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara hyperborea" order="Coleoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="hyperborea">Amara hyperborea</taxonomicName>
Dejean, 1831: 800 (type locality:
<normalizedToken id="69D11D29BB75ACE6F613F56D0913BB78" originalValue="“Labrador”">&quot;Labrador&quot;</normalizedToken>
[= Newfoundland and Labrador, Canada]; holotype in MNHN);
<bibRefCitation id="6331AA6D89584560FBAB76FA1EE0A8BB" author="Putzeys, JAAH" journalOrPublisher="Stettin" pageId="0" pageNumber="39" pagination="332 - 344" refId="B45" refString="Putzeys, JAAH, 1865. Remarques sur les amaroides. Entomologische Zeitung. Stettin 26: 332 - 344" title="Remarques sur les amaroides. Entomologische Zeitung." volume="26" year="1865">Putzeys 1865</bibRefCitation>
: 338;
<bibRefCitation id="0CB9AB5341A5F0F480693B7337F16B64" author="Putzeys, JAAH" journalOrPublisher="Memoires de la Societe Royale des Sciences de Liege" pageId="0" pageNumber="39" pagination="171 - 283" refId="B46" refString="Putzeys, JAAH, 1866. Etude sur les Amara de la collection de Mr. le Baron de Chaudoir. Memoires de la Societe Royale des Sciences de Liege 1 (2): 171 - 283" title="Etude sur les Amara de la collection de Mr. le Baron de Chaudoir." volume="1" year="1866">Putzeys 1866</bibRefCitation>
: 257;
<bibRefCitation id="8DE602AF08E9115CEB0667B1B40DCF73" author="Jakobson, G" journalOrPublisher="Devrien, St. Petersburg" pageId="0" pageNumber="39" refId="B24" refString="Jakobson, G, 1906. Shuki Rossii i sapadnoj Ewropy 1. Devrien, St. Petersburg" title="Shuki Rossii i sapadnoj Ewropy 1." year="1906">Jakobson 1906</bibRefCitation>
: 262;
<bibRefCitation id="1BF9A77CB4EA2529C162C738262B8917" author="Lindroth, CH" journalOrPublisher="Opuscula Entomologica" pageId="0" pageNumber="39" pagination="17 - 21" refId="B32" refString="Lindroth, CH, 1953. Fennoskandicus Carabider. Ett. litet Supplement. Opuscula Entomologica 18: 17 - 21" title="Fennoskandicus Carabider. Ett. litet Supplement." volume="18" year="1953">Lindroth 1953</bibRefCitation>
: 18;
<bibRefCitation id="07AE82D0651DDEE8AD9152EFDF798128" author="Lindroth, CH" journalOrPublisher="Bulletin of the Museum of Comparative Zoology at Harvard College" pageId="0" pageNumber="39" pagination="117 - 161" refId="B33" refString="Lindroth, CH, 1954. Random notes on North American Carabidae (Coleopt.). Bulletin of the Museum of Comparative Zoology at Harvard College 111: 117 - 161" title="Random notes on North American Carabidae (Coleopt.)." volume="111" year="1954">Lindroth 1954</bibRefCitation>
: 134, 135;
<bibRefCitation id="617C6006AA00B5208102C7D5529F118F" author="Lindroth, CH" journalOrPublisher="Bulletin of the Museum of Comparative Zoology at Harvard College" pageId="0" pageNumber="39" refId="B34" refString="Lindroth, CH, 1955. The Carabid beetles of Newfoundland including the French islands St. Pierre and Miquelon. Opuscula Entomologica (Suppl. 12): 1-160." title="The Carabid beetles of Newfoundland including the French islands St. Pierre and Miquelon. Opuscula Entomologica (Suppl. 12): 1 - 160." year="1955">Lindroth 1955</bibRefCitation>
: 98;
<bibRefCitation id="FF91702DCD11652E762601B25996E75A" DOI="https://doi.org/10.1002/mmnz.19900660202" author="Hieke, F" journalOrPublisher="Mitteilungen aus dem Zoologischen Museum in Berlin" pageId="0" pageNumber="39" pagination="195 - 292" refId="B17" refString="Hieke, F, 1990. Neue und wenig bekannte Amara -Arten aus Amerika und Asien (Coleoptera, Carabidae). Mitteilungen aus dem Zoologischen Museum in Berlin 66 (2): 195 - 292, DOI: https://doi.org/10.1002/mmnz.19900660202" title="Neue und wenig bekannte Amara - Arten aus Amerika und Asien (Coleoptera, Carabidae)." url="https://doi.org/10.1002/mmnz.19900660202" volume="66" year="1990">Hieke 1990</bibRefCitation>
: 252;
<bibRefCitation id="117DE9885A5A0E765A96165C79FADC97" DOI="https://doi.org/10.1002/mmnd.19930400102" author="Hieke, F" journalOrPublisher=") (Berlin)]" pageId="0" pageNumber="39" pagination="1 - 160" refId="B18" refString="Hieke, F, 1993. Die Untergattung Harpalodema Reitter 1888, von Amara Bon. sowie ueber Amara -Arten anderer Subgenera (Coleoptera, Carabidae). Deutsche Entomologische Zeitschrift [(N. F.) (Berlin)] 40 (1): 1 - 160, DOI: https://doi.org/10.1002/mmnd.19930400102" title="Die Untergattung Harpalodema Reitter 1888, von Amara Bon. sowie ueber Amara - Arten anderer Subgenera (Coleoptera, Carabidae). Deutsche Entomologische Zeitschrift [(N. F." url="https://doi.org/10.1002/mmnd.19930400102" volume="40" year="1993">Hieke 1993</bibRefCitation>
: 147.
</paragraph>
<paragraph id="2BD5CD4F947D0E99AE38832110139337" pageId="0" pageNumber="39">
<taxonomicName id="FC5A46F6CCB6728C2DEE7B735D06B987" authorityName="LeConte" authorityYear="1850" class="Insecta" family="Carabidae" genus="Curtonotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Curtonotus elongatus" order="Coleoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="elongatus">Curtonotus elongatus</taxonomicName>
LeConte, 1850: 207 (type locality: &quot;Lake Superior&quot; [Probably in USA]; holotype in CUMZ); synonymized by
<bibRefCitation id="8D759102B850CC55E20425440AB0C6E0" author="Lindroth, CH" journalOrPublisher="Bulletin of the Museum of Comparative Zoology at Harvard College" pageId="0" pageNumber="39" pagination="117 - 161" refId="B33" refString="Lindroth, CH, 1954. Random notes on North American Carabidae (Coleopt.). Bulletin of the Museum of Comparative Zoology at Harvard College 111: 117 - 161" title="Random notes on North American Carabidae (Coleopt.)." volume="111" year="1954">Lindroth 1954</bibRefCitation>
: 134.
</paragraph>
<paragraph id="F349B7966F1B939798823593FBEF2286" pageId="0" pageNumber="39">
<taxonomicName id="6FE3F819479FCADDFB5701A20C2C7068" authorityName="Motschulsky" authorityYear="1859" class="Insecta" family="Carabidae" genus="Leirus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leirus ovipennis" order="Coleoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="ovipennis">Leirus ovipennis</taxonomicName>
Motschulsky, 1859: 156 (type locality:
<normalizedToken id="5ECE5597DD458A6A8875A491A7AA38BF" originalValue="“Californie”">&quot;Californie&quot;</normalizedToken>
[actually in western Alaska]; syntypes in ZMUM); synonymized by
<bibRefCitation id="4368AF511FF8DAFE6B31187E63BB530E" DOI="https://doi.org/10.1002/mmnd.19930400102" author="Hieke, F" journalOrPublisher=") (Berlin)]" pageId="0" pageNumber="39" pagination="1 - 160" refId="B18" refString="Hieke, F, 1993. Die Untergattung Harpalodema Reitter 1888, von Amara Bon. sowie ueber Amara -Arten anderer Subgenera (Coleoptera, Carabidae). Deutsche Entomologische Zeitschrift [(N. F.) (Berlin)] 40 (1): 1 - 160, DOI: https://doi.org/10.1002/mmnd.19930400102" title="Die Untergattung Harpalodema Reitter 1888, von Amara Bon. sowie ueber Amara - Arten anderer Subgenera (Coleoptera, Carabidae). Deutsche Entomologische Zeitschrift [(N. F." url="https://doi.org/10.1002/mmnd.19930400102" volume="40" year="1993">Hieke 1993</bibRefCitation>
: 147.
</paragraph>
<paragraph id="9881C04153F7C62C91F1ACB977FD32E0" pageId="0" pageNumber="39">
<taxonomicName id="794B425A26DB250C8AD4E1AE30B8711D" authorityName="Motschulsky" authorityYear="1860" class="Insecta" family="Carabidae" genus="Leirus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leirus longicollis" order="Coleoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="longicollis">Leirus longicollis</taxonomicName>
Motschulsky, 1860: 95 (type locality: &quot;Daourie orientale&quot; [in Kamchatka, Russia]; syntypes in ZMUM);
<bibRefCitation id="53E9064E41B987B54137580365E56A6B" author="Tschitscherine, T" journalOrPublisher="Horae Societatis Entomologicae Rossicae" pageId="0" pageNumber="39" pagination="366 - 435" refId="B55" refString="Tschitscherine, T, 1894. Materiaux pour servir a l'etude des Feroniens. II. Horae Societatis Entomologicae Rossicae 28 (1893-1894): 366 - 435" title="Materiaux pour servir a l'etude des Feroniens. II." volume="28" year="1894">
<normalizedToken id="8A297F14895B4437097217B51FA1204E" originalValue="Tschitschérine">Tschitscherine</normalizedToken>
1894
</bibRefCitation>
: 389; synonymized by
<bibRefCitation id="834994DE2F6DE5EA11F4693D15FF294A" author="Putzeys, JAAH" journalOrPublisher="Stettin" pageId="0" pageNumber="39" pagination="332 - 344" refId="B45" refString="Putzeys, JAAH, 1865. Remarques sur les amaroides. Entomologische Zeitung. Stettin 26: 332 - 344" title="Remarques sur les amaroides. Entomologische Zeitung." volume="26" year="1865">Putzeys 1865</bibRefCitation>
: 338.
</paragraph>
<paragraph id="0CC5735F25178211317FE40FAA69CF78" pageId="0" pageNumber="39">
<taxonomicName id="9CF5FC4745E2AD8C729EF3659699D8EF" authorityName="A. Morawitz" authorityYear="1862" class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara (Leirus) peregrina" order="Coleoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="peregrina" subGenus="Leirus">Amara (Leirus) peregrina</taxonomicName>
Morawitz, 1862: 258 (type locality:
<normalizedToken id="6D0A28964794A647C8ABE70B51956118" originalValue="“Kulussutai”">&quot;Kulussutai&quot;</normalizedToken>
[in southeastern Siberia]; syntypes in ZRAS); synonymized by
<bibRefCitation id="D42F12A818DD8F6794D7E42699A074D7" author="Lindroth, CH" journalOrPublisher="Opuscula Entomologica" pageId="0" pageNumber="39" pagination="17 - 21" refId="B32" refString="Lindroth, CH, 1953. Fennoskandicus Carabider. Ett. litet Supplement. Opuscula Entomologica 18: 17 - 21" title="Fennoskandicus Carabider. Ett. litet Supplement." volume="18" year="1953">Lindroth 1953</bibRefCitation>
: 18.
</paragraph>
<paragraph id="9091848C82E5C2B7955F94F0308A289B" pageId="0" pageNumber="39">
<taxonomicName id="F3CB0FBC2BA7AE0EAAEC1DF17A0D7856" authorityName="Putzeys" authorityYear="1866" class="Insecta" family="Carabidae" genus="Curtonotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Curtonotus canadensis" order="Coleoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="canadensis">Curtonotus canadensis</taxonomicName>
Putzeys, 1866: 256 (type locality: &quot;Canada
<normalizedToken id="07BB5D69C7B596F824C3B0A5EAE17816" originalValue="boréal.”">boreal.&quot;</normalizedToken>
[= northern Canada]; holotype in MNHN); synonymized by
<bibRefCitation id="EDDEEB5CF55AF1A742EF524C3A852351" DOI="https://doi.org/10.1093/besa/13.1.77" author="Lindroth, CH" journalOrPublisher="Bulletin of the Museum of Comparative Zoology at Harvard College" pageId="0" pageNumber="39" refId="B35" refString="Lindroth, CH, 1968. The Ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska Part 5. Opuscula Entomologica (Suppl. 33): 649-944. https://doi.org/10.1093/besa/13.1.77" title="The Ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska Part 5. Opuscula Entomologica (Suppl. 33): 649 - 944." url="https://doi.org/10.1093/besa/13.1.77" year="1968">Lindroth 1968</bibRefCitation>
: 678.
</paragraph>
<paragraph id="8F14786D8ABFB330C90ED9FD68DFC2BF" pageId="0" pageNumber="39">
<taxonomicName id="E156B6015150BF4A3AE5BB6D6C15DC77" authorityName="Putzeys" authorityYear="1866" class="Insecta" family="Carabidae" genus="Curtonotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Curtonotus dejeani" order="Coleoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="dejeani">Curtonotus dejeani</taxonomicName>
Putzeys, 1866: 258 (type locality:
<normalizedToken id="C89CDA8A589A62F21EDCEDF0C1B6153E" originalValue="“Kamchatka”">&quot;Kamchatka&quot;</normalizedToken>
; holotype in MNHN); synonymized to
<taxonomicName id="2689C463AB2B17343AD2E932F4C51542" family="Carabidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="A. longicollis" order="Coleoptera" pageId="0" pageNumber="39" rank="species" species="longicollis">A. longicollis</taxonomicName>
by
<bibRefCitation id="4E8494FD33424D6441CB61A3AD3847EA" author="Tschitscherine, T" journalOrPublisher="Horae Societatis Entomologicae Rossicae" pageId="0" pageNumber="39" pagination="366 - 435" refId="B55" refString="Tschitscherine, T, 1894. Materiaux pour servir a l'etude des Feroniens. II. Horae Societatis Entomologicae Rossicae 28 (1893-1894): 366 - 435" title="Materiaux pour servir a l'etude des Feroniens. II." volume="28" year="1894">
<normalizedToken id="D985F214461E0D211711B119FE9F4F61" originalValue="Tschitschérine">Tschitscherine</normalizedToken>
1894
</bibRefCitation>
: 389.
</paragraph>
<paragraph id="AFE8E7BE0B538CF2E9F1AD42BE552CCA" pageId="0" pageNumber="39">
<taxonomicName id="596FFCEA1AD2745067F2DBDF02148EB3" authorityName="Putzeys" authorityYear="1866" class="Insecta" family="Carabidae" genus="Curtonotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Curtonotus pedestris" order="Coleoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="pedestris">Curtonotus pedestris</taxonomicName>
Putzeys, 1866: 254 (type locality: &quot;Udskoe Okhotsk&quot; [= Udskoye, Region Chabarowsk, Russia; holotype in MNHN); synonymized to
<taxonomicName id="A321FBF922383D323643FD76905589D7" family="Carabidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="A. (C.) hyperborea" order="Coleoptera" pageId="0" pageNumber="39" rank="species" species="hyperborea" subGenus="C.">A. (C.) hyperborea</taxonomicName>
Dejean by
<bibRefCitation id="4A9FB809DE850D60C823C66E7DE737C9" DOI="https://doi.org/10.1002/mmnz.19900660202" author="Hieke, F" journalOrPublisher="Mitteilungen aus dem Zoologischen Museum in Berlin" pageId="0" pageNumber="39" pagination="195 - 292" refId="B17" refString="Hieke, F, 1990. Neue und wenig bekannte Amara -Arten aus Amerika und Asien (Coleoptera, Carabidae). Mitteilungen aus dem Zoologischen Museum in Berlin 66 (2): 195 - 292, DOI: https://doi.org/10.1002/mmnz.19900660202" title="Neue und wenig bekannte Amara - Arten aus Amerika und Asien (Coleoptera, Carabidae)." url="https://doi.org/10.1002/mmnz.19900660202" volume="66" year="1990">Hieke 1990</bibRefCitation>
: 252.
</paragraph>
<paragraph id="1ADBCF97EC8FBCBE80F7FBE92A06E681" pageId="0" pageNumber="39">
<taxonomicName id="3FAC7A36D9153734E04111C21D2622AF" authorityName="Putzeys" authorityYear="1866" class="Insecta" family="Carabidae" genus="Curtonotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Curtonotus tristis" order="Coleoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="tristis">Curtonotus tristis</taxonomicName>
Putzeys, 1866: 255 (type locality:
<normalizedToken id="82C570FA6446E164C4E8B6E29754E679" originalValue="“Oowho-Bay”">&quot;Oowho-Bay&quot;</normalizedToken>
[= Hudson Bay, Canada]; holotype in MNHN); synonymized to
<taxonomicName id="506D7141AAD5B2A6A8C447BF6CFF5F54" family="Carabidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="A. (C.) hyperborea" order="Coleoptera" pageId="0" pageNumber="39" rank="species" species="hyperborea" subGenus="C.">A. (C.) hyperborea</taxonomicName>
by
<bibRefCitation id="51DAE015FCD3E5E80882AE0E1BFB4080" DOI="https://doi.org/10.1093/besa/13.1.77" author="Lindroth, CH" journalOrPublisher="Bulletin of the Museum of Comparative Zoology at Harvard College" pageId="0" pageNumber="39" refId="B35" refString="Lindroth, CH, 1968. The Ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska Part 5. Opuscula Entomologica (Suppl. 33): 649-944. https://doi.org/10.1093/besa/13.1.77" title="The Ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska Part 5. Opuscula Entomologica (Suppl. 33): 649 - 944." url="https://doi.org/10.1093/besa/13.1.77" year="1968">Lindroth 1968</bibRefCitation>
: 253.
</paragraph>
<paragraph id="1A3BF4A6BB23EB069E210C1472FBCFF4" pageId="0" pageNumber="39">
<taxonomicName id="6AFD7CA807B4A4442E3FA639EE180000" authorityName="J. Sahlberg" authorityYear="1880" class="Insecta" family="Carabidae" genus="Harpalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpalus simulans" order="Coleoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="simulans">Harpalus simulans</taxonomicName>
Sahlberg, 1880: 44 (type locality: &quot;Tschornaja ostrow&quot; [= Yenisey range, Russia]; syntypes in NHRS); synonymized to
<taxonomicName id="CAF8965A69D7A417512D381C13333038" family="Carabidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="A. (C.) hyperborea" order="Coleoptera" pageId="0" pageNumber="39" rank="species" species="hyperborea" subGenus="C.">A. (C.) hyperborea</taxonomicName>
by
<bibRefCitation id="73AF2E43245C45C12B5344702B267ECB" author="Jakobson, G" journalOrPublisher="Devrien, St. Petersburg" pageId="0" pageNumber="39" refId="B24" refString="Jakobson, G, 1906. Shuki Rossii i sapadnoj Ewropy 1. Devrien, St. Petersburg" title="Shuki Rossii i sapadnoj Ewropy 1." year="1906">Jakobson 1906</bibRefCitation>
: 262.
</paragraph>
<paragraph id="3F1C1691CF9C74699ADBE501EDAB195C" pageId="0" pageNumber="39">
<taxonomicName id="3B59A273D46E7FE71AA56D21D6376212" authorityName="Brown" authorityYear="1930" class="Insecta" family="Carabidae" genus="Curtonotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Curtonotus imperfectus" order="Coleoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="imperfectus">Curtonotus imperfectus</taxonomicName>
Brown, 1930: 232 (type locality: &quot;Bradore Bay&quot; [in Quebec, Canada]; holotype in CMNC); synonymized to
<taxonomicName id="DCFBFE6E4D43A083C7523DCCAA2DFC8E" family="Carabidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="A. (C.) hyperborea" order="Coleoptera" pageId="0" pageNumber="39" rank="species" species="hyperborea" subGenus="C.">A. (C.) hyperborea</taxonomicName>
by
<bibRefCitation id="D7F5B46F9FFFEC9392BA9034154DF283" author="Lindroth, CH" journalOrPublisher="Bulletin of the Museum of Comparative Zoology at Harvard College" pageId="0" pageNumber="39" pagination="117 - 161" refId="B33" refString="Lindroth, CH, 1954. Random notes on North American Carabidae (Coleopt.). Bulletin of the Museum of Comparative Zoology at Harvard College 111: 117 - 161" title="Random notes on North American Carabidae (Coleopt.)." volume="111" year="1954">Lindroth 1954</bibRefCitation>
: 135.
</paragraph>
<paragraph id="F974F63DC5B270D3010C6BB9C1D98B74" pageId="0" pageNumber="39">
<taxonomicName id="18E79C9400A7696808DDA36AA1F3677F" authorityName="Baliani" authorityYear="1937" class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara (Curtonotus) coreana" order="Coleoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="coreana" subGenus="Curtonotus">Amara (Curtonotus) coreana</taxonomicName>
Baliani, 1937: 182 (type locality:
<normalizedToken id="8F608C2FF5BF6E131AA7DF07D060E665" originalValue="“Ompo”">&quot;Ompo&quot;</normalizedToken>
[= Onbo, North Hamgyeong Province, North Korea]; holotype in MSNG); synonymized to
<taxonomicName id="106BB67D396EAE1242AB8C8B54BF997D" family="Carabidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="A. (C.) hyperborea" order="Coleoptera" pageId="0" pageNumber="39" rank="species" species="hyperborea" subGenus="C.">A. (C.) hyperborea</taxonomicName>
by
<bibRefCitation id="A4FA1F75910EE648FC5530715BE37784" DOI="https://doi.org/10.1002/mmnz.19900660202" author="Hieke, F" journalOrPublisher="Mitteilungen aus dem Zoologischen Museum in Berlin" pageId="0" pageNumber="39" pagination="195 - 292" refId="B17" refString="Hieke, F, 1990. Neue und wenig bekannte Amara -Arten aus Amerika und Asien (Coleoptera, Carabidae). Mitteilungen aus dem Zoologischen Museum in Berlin 66 (2): 195 - 292, DOI: https://doi.org/10.1002/mmnz.19900660202" title="Neue und wenig bekannte Amara - Arten aus Amerika und Asien (Coleoptera, Carabidae)." url="https://doi.org/10.1002/mmnz.19900660202" volume="66" year="1990">Hieke 1990</bibRefCitation>
: 255.
</paragraph>
</subSubSection>
<subSubSection id="7767A1CE1BBCC7356BB8D5337F2DB14F" pageId="0" pageNumber="39" type="chinese common name">
<paragraph id="3B970C2D667DAAA6DFBDE8DC24DCDCA2" pageId="0" pageNumber="39">Chinese common name.</paragraph>
<paragraph id="AD3F709112A7997FE1A9BB19793E0EA6" pageId="0" pageNumber="39">
<normalizedToken id="D86D4EB0AFDA8964BAFA5AB4AD329BB4" originalValue="极北暗步甲">极北暗步甲</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection id="6C2A0386F776B57554B2436D9B6BCD66" pageId="0" pageNumber="39" type="diagnosis">
<paragraph id="AE77288FC3A9B06AFD46AC5ADAB91291" pageId="0" pageNumber="39">Diagnosis.</paragraph>
<paragraph id="8A3C0DBAA514EC2727656D76DEEA905F" pageId="0" pageNumber="39">
Small- to medium-sized species, BL = 9.0-13.0 mm; dorsum brown, legs reddish brown; head small, ~ 1/2 of the pronotum maximum width; head with one supraorbital seta. Pronotum cordate, punctate at base, sporadically punctate at mid-anterior region; lateral margins nearly straight before posterior angles; posterior angles nearly rectangular, slightly laterally protruded. Elytra long, widest after middle; lateral sides of abdominal sternites smooth. Male mesotibiae projection composed of two denticles; proximal denticle acutely pointed beyond middle; distal denticle smaller, present between the proximal one and tibiae apex. Male genitalia with short apical lamella, slightly declined toward right, narrowed to apex; gonocoxite 2 of ovipositor elongated, length ~ 1.7-2.0
<normalizedToken id="4B6B8A838607EF9201FEFC88CF25E152" originalValue="×">x</normalizedToken>
as greatest width, narrowed to apex.
</paragraph>
</subSubSection>
<subSubSection id="0888ED448CCE25D37C455F03AD13A8B0" pageId="0" pageNumber="39" type="comparisons">
<paragraph id="B2174D5CEF30D3E0398F469B075BBD97" pageId="0" pageNumber="39">Comparison.</paragraph>
<paragraph id="4C29BF6CEEE6C84A26E5DA147A2161F9" pageId="0" pageNumber="39">this species can be distinguished by having relatively elongated body, impunctate sides of abdominal sternites, nearly smooth pronotum anterior portion, and nearly straight pronotum lateral edge before posterior angle.</paragraph>
</subSubSection>
<subSubSection id="0A4CB17ABAA03C56D88A1AB7851C77F0" pageId="0" pageNumber="39" type="distribution">
<paragraph id="900EDD778548264808583850B3777A0E" pageId="0" pageNumber="39">Distribution.</paragraph>
<paragraph id="BBE4C3A1E8BA9C9E1053E0D7222603BF" pageId="0" pageNumber="39">China (Heilongjiang, Jilin, Xinjiang), Russia (Far East, east Siberia, west Siberia), Mongolia, North Korea, Europe, North America.</paragraph>
</subSubSection>
<subSubSection id="5AC5664CB35C15DCDC99FE9EE7C32C56" pageId="0" pageNumber="39" type="remarks">
<paragraph id="92302EA8192286F4FC6C6C490DD26F97" pageId="0" pageNumber="39">Remarks.</paragraph>
<paragraph id="04BA7B6800DEBDCBD49E13064F33ADBA" pageId="0" pageNumber="39">
This species, as well as the following two species (
<taxonomicName id="7340DB58AED1D87557A23CBC9C1A4441" family="Carabidae" kingdom="Animalia" lsidName="A. (C.) tumida" order="Coleoptera" pageId="0" pageNumber="39" rank="species" species="tumida" subGenus="C.">A. (C.) tumida</taxonomicName>
,
<taxonomicName id="FAE0BF3563C845C9C1FBEB9A93D8E8C4" family="Carabidae" kingdom="Animalia" lsidName="A. (C.) shinanensis" order="Coleoptera" pageId="0" pageNumber="39" rank="species" species="shinanensis" subGenus="C.">A. (C.) shinanensis</taxonomicName>
), was recorded from the provinces of northeastern China (
<bibRefCitation id="AC0D0BF68FD55D86495FCD72F7C309BD" author="Habu, A" journalOrPublisher="Kontyu" pageId="0" pageNumber="39" pagination="39 - 44" refId="B16" refString="Habu, A, 1953. Notes on the species of the genus Curtonotus from Japan (Coleoptera, Carabidae). Kontyu 20 (1-2): 39 - 44" title="Notes on the species of the genus Curtonotus from Japan (Coleoptera, Carabidae)." volume="20" year="1953">Habu 1953</bibRefCitation>
;
<bibRefCitation id="5AB203BB5E99117D2DEAD6F42F1129B7" DOI="https://doi.org/10.1002/mmnz.19900660202" author="Hieke, F" journalOrPublisher="Mitteilungen aus dem Zoologischen Museum in Berlin" pageId="0" pageNumber="39" pagination="195 - 292" refId="B17" refString="Hieke, F, 1990. Neue und wenig bekannte Amara -Arten aus Amerika und Asien (Coleoptera, Carabidae). Mitteilungen aus dem Zoologischen Museum in Berlin 66 (2): 195 - 292, DOI: https://doi.org/10.1002/mmnz.19900660202" title="Neue und wenig bekannte Amara - Arten aus Amerika und Asien (Coleoptera, Carabidae)." url="https://doi.org/10.1002/mmnz.19900660202" volume="66" year="1990">Hieke 1990</bibRefCitation>
). However, we did not find any specimens collected from eastern China that accord with these three species although several specimens of
<taxonomicName id="9959E802524ED2B9C262F2AF780AE7B6" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara (Curtonotus)" order="Coleoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="subGenus" subGenus="Curtonotus">
<emphasis id="28EA7A16205F4CDFA033A2896B13C98F" italics="true" pageId="0" pageNumber="39">Amara Curtonotus</emphasis>
</taxonomicName>
were examined from northeastern China. Considering many species of this subgenus are superficially similar externally, we suspect that these three species records from China could be based on misidentifications of similar species. We hope future research can corroborate their precise distribution in China.
</paragraph>
</subSubSection>
</treatment>
</document>
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