treatments-xml/data/35/26/B8/3526B888D3275CC2B5D1DB7E34C24AB3.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1036.66435" ID-GBIF-Dataset="bac822e0-f6b0-44f7-9ad8-17464faa8772" ID-PMC="PMC8128840" ID-Pensoft-Pub="1313-2970-1036-99" ID-Pensoft-UUID="87E9A246E0FD5DBA81318090D48DC9FD" ID-PubMed="34040482" ID-ZooBank="F4A9958818BD4B918DB43368AF1D7984" ModsDocID="1313-2970-1036-99" checkinTime="1620734415262" checkinUser="pensoft" docAuthor="Godunko, Roman J., Martynov, Alexander V. &amp; Staniczek, Arnold H." docDate="2021" docId="3526B888D3275CC2B5D1DB7E34C24AB3" docLanguage="en" docName="ZooKeys 1036: 99-120" docOrigin="ZooKeys 1036" docPubDate="2021-05-10" docSource="http://dx.doi.org/10.3897/zookeys.1036.66435" docTitle="Burmella Godunko &amp; Martynov &amp; Staniczek 2021, gen. nov." docType="treatment" docUuid="0A29850B-F977-47E1-948E-37B5C617BD95" docUuidSource="ZooBank" docVersion="4" id="87E9A246E0FD5DBA81318090D48DC9FD" lastPageNumber="99" masterDocId="87E9A246E0FD5DBA81318090D48DC9FD" masterDocTitle="First fossil record of the mayfly family Vietnamellidae (Insecta, Ephemeroptera) from Burmese Amber confirms its Oriental origin and gives new insights into its evolution" masterLastPageNumber="120" masterPageNumber="99" pageNumber="99" updateTime="1668150237642" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>First fossil record of the mayfly family Vietnamellidae (Insecta, Ephemeroptera) from Burmese Amber confirms its Oriental origin and gives new insights into its evolution</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Godunko, Roman J.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2196-3327</mods:nameIdentifier>
<mods:affiliation>Biology Centre of the Czech Academy of Sciences, Institute of Entomology, Branisovska 31, 37005 Ceske Budejovice, Czech Republic &amp; Department of Invertebrate Zoology and Hydrobiology, University of Lodz, Banacha 12 / 16, 90237 Lodz, Poland &amp; State Museum of Natural History, National Academy of Sciences of Ukraine, Teatralna 18, 79008, Lviv, Ukraine</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Martynov, Alexander V.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6506-5134</mods:nameIdentifier>
<mods:affiliation>National Museum of Natural History, National Academy of Sciences of Ukraine, Bohdan Khmelnytsky 15, 01030, Kyiv, Ukraine</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Staniczek, Arnold H.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5577-137X</mods:nameIdentifier>
<mods:affiliation>Department of Entomology, State Museum of Natural History Stuttgart, Rosenstein 1, 70191, Stuttgart, Germany</mods:affiliation>
<mods:nameIdentifier type="email">arnold.staniczek@smns-bw.de</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-05-10</mods:number>
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<mods:detail type="volume">
<mods:number>1036</mods:number>
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<mods:extent unit="page">
<mods:start>99</mods:start>
<mods:end>120</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1036.66435</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1036.66435</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1036-99</mods:identifier>
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<treatment ID-GBIF-Taxon="181974059" LSID="urn:lsid:zoobank.org:act:0A29850B-F977-47E1-948E-37B5C617BD95" httpUri="http://treatment.plazi.org/id/3526B888D3275CC2B5D1DB7E34C24AB3" lastPageNumber="99" pageId="0" pageNumber="99">
<subSubSection pageId="0" pageNumber="99" type="nomenclature">
<paragraph pageId="0" pageNumber="99">
Genus 
<taxonomicName LSID="http://zoobank.org/0A29850B-F977-47E1-948E-37B5C617BD95" authority="Godunko &amp; Martynov &amp; Staniczek, 2021" authorityName="Godunko &amp; Martynov &amp; Staniczek" authorityYear="2021" class="Insecta" family="Vietnamellidae" genus="Burmella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Burmella" order="Ephemeroptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="genus" status="gen. nov.">Burmella</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="99">gen. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Burmella paucivenosa sp. nov., male imago, holotype, general lateral view A left side of body B right side of body. Scale bars: 2 mm." figureDoi="10.3897/zookeys.1036.66435.figure1" httpUri="https://binary.pensoft.net/fig/541121" pageId="0" pageNumber="99">Figures 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Burmella paucivenosa sp. nov., male imago, holotype A head, right lateral view B compound eye, left lateral view C head and thorax, left lateral view D mesothorax, left lateral view. Blue line - border between portions of compound eye, da - damaged area, lp - lower portion, LPs and red line - lateroparapsidal suture, MNs and green line - mesonotal suture, MPs and yellow line - medioparapsidal suture, up - upper portion. Scale bars: 0.5 mm (A, C); 0.2 mm (B, D)." figureDoi="10.3897/zookeys.1036.66435.figure2" httpUri="https://binary.pensoft.net/fig/541122" pageId="0" pageNumber="99">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Burmella paucivenosa sp. nov., male imago, holotype A, B tarsus of right middle leg C right middle leg D left hind leg. I-V - tarsal segments, pts - patellotibial suture. Scale bars: 0.1 mm (A, B); 0.5 mm (C); 0.2 mm (D)." figureDoi="10.3897/zookeys.1036.66435.figure3" httpUri="https://binary.pensoft.net/fig/541123" pageId="0" pageNumber="99">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Burmella paucivenosa sp. nov., male imago, holotype A right and left wings position in amber B right forewing venation and size ratio of fore and hind wings. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1036.66435.figure4" httpUri="https://binary.pensoft.net/fig/541124" pageId="0" pageNumber="99">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Burmella paucivenosa sp. nov., male imago, holotype A right hind wing in amber B right hind wing venation. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1036.66435.figure5" httpUri="https://binary.pensoft.net/fig/541125" pageId="0" pageNumber="99">, 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Burmella paucivenosa sp. nov., male imago, holotype A abdomen, left lateral view B abdominal segments III-VII, lateral view C, D genitalia, ventrolateral view. III-VII - numbers of segments, white arrows mark remnants of gill sockets, pink area - penis lobes, light blue area - gonobasis (styliger plate), brown area styliger segment I, yellow area - styliger segment II, light green area - styliger segment III, purple area - styliger segment IV. Scale bars: 1 mm (A); 0.5 mm (B); 0.2 mm (C, D)." figureDoi="10.3897/zookeys.1036.66435.figure6" httpUri="https://binary.pensoft.net/fig/541126" pageId="0" pageNumber="99">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Burmella clypeata sp. nov., female imago, holotype A general dorsal view B general ventral view C, D head, dorsal view E head and thorax, dorsal view. BA - basal sclerite (basalare), LPs and red line - lateroparapsidal suture, MNs and green line - mesonotal suture, MPs and yellow line - medioparapsidal suture, rc - right cercus. Scale bars: 2 mm (A, B); 0.2 mm (C, D); 0.5 mm (E)." figureDoi="10.3897/zookeys.1036.66435.figure7" httpUri="https://binary.pensoft.net/fig/541127" pageId="0" pageNumber="99">, 7</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Burmella clypeata sp. nov., female imago, holotype A tarsus of left hind leg B tarsus of right hind leg C left forewing in amber D left forewing venation and size ratio of left fore- and right hind wings E distal part of left forewing F preserved basal part of right forewing. I-V - tarsal segments, ci - basally connected intercalary vein, fi - basally free intercalary vein. Scale bars: 0.1 mm (A, B); 1 mm (C, D); 0.5 mm (E, F)." figureDoi="10.3897/zookeys.1036.66435.figure8" httpUri="https://binary.pensoft.net/fig/541128" pageId="0" pageNumber="99">, 8</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Burmella clypeata sp. nov., female imago, holotype A right hind wing, dorsal view B right hind wing, ventral view C right hind wing venation. Scale bars: 0.2 mm (A, B); 0.5 mm (C)." figureDoi="10.3897/zookeys.1036.66435.figure9" httpUri="https://binary.pensoft.net/fig/541129" pageId="0" pageNumber="99">, 9</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Burmella clypeata sp. nov., female imago, holotype A body, ventral view B, C apical part of abdomen, ventral view D basal segments of abdomen, dorsal view. I-III and V-IX - numbers of abdominal segments, gsr - remnant of gill socket, pp - paraproct plate. Scale bars: 1 mm (A); 0.5 mm (B-D)." figureDoi="10.3897/zookeys.1036.66435.figure10" httpUri="https://binary.pensoft.net/fig/541130" pageId="0" pageNumber="99">, 10</figureCitation>
, 
<figureCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements of fossil representatives of the genus Burmella gen. nov." pageId="0" pageNumber="99">Table 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="type species">
<paragraph pageId="0" pageNumber="99">Type species.</paragraph>
<paragraph pageId="0" pageNumber="99">
<taxonomicName authorityName="Godunko &amp; Martynov &amp; Staniczek" authorityYear="2021" class="Insecta" family="Vietnamellidae" genus="Burmella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Burmella paucivenosa" order="Ephemeroptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="paucivenosa">
<emphasis italics="true" pageId="0" pageNumber="99">Burmella paucivenosa</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="derivation of name">
<paragraph pageId="0" pageNumber="99">Derivation of name.</paragraph>
<paragraph pageId="0" pageNumber="99">
The generic name of female gender is a composition of &quot; 
<taxonomicName genus="Burmar" lsidName="Burmar" pageId="0" pageNumber="99" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="99">Burmar</emphasis>
</taxonomicName>
&quot; as an ancient term for Myanmar, combined with &quot; 
<emphasis italics="true" pageId="0" pageNumber="99">ella</emphasis>
&quot;, a common ending of generic names in mayflies, and especially so within 
<taxonomicName lsidName="" pageId="0" pageNumber="99" rank="superFamily" superFamily="Ephemerelloidea">Ephemerelloidea</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="diagnosis">
<paragraph pageId="0" pageNumber="99">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="99">
Adults of 
<taxonomicName authorityName="Godunko &amp; Martynov &amp; Staniczek" authorityYear="2021" class="Insecta" family="Vietnamellidae" genus="Burmella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Burmella" order="Ephemeroptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="99">Burmella</emphasis>
</taxonomicName>
gen. nov. differ from other mayfly genera by the following combination of features: 
<emphasis italics="true" pageId="0" pageNumber="99">forewings</emphasis>
(a) with small number of cross veins; (b) pterostigma with simple veins, not anastomosed; (c) CuP smoothly curved towards wing base; (d) two secondary bifurcate veins in cubital field; (e) at least several free marginal intercalaries along ventral margin; 
<emphasis italics="true" pageId="0" pageNumber="99">hind wings</emphasis>
(f) strongly rounded, small, as long as 0.08-0.14 of forewing length; (g) small number of cross veins; (h) triad RS present or absent; no MA and MP triads; (i) no secondary branches of cubital veins; (j) costal process developed, rounded apically, situated centrally; 
<emphasis italics="true" pageId="0" pageNumber="99">abdomen</emphasis>
(k) with vestigial gill sockets recognizable at least on segments II-VI; 
<emphasis italics="true" pageId="0" pageNumber="99">genitalia</emphasis>
(l) with large median projection of styliger plate, widely rounded apically; (m) three distal segments of forceps strongly elongated and slender; segment II longest, 5 
<normalizedToken originalValue="×">x</normalizedToken>
as long as segment III; segments III and IV approximately of equal length; segment IV expanding apically; (n) penis lobes widely separated by V-shaped cleft; (o) no trace of paracercus. Additionally, 
<emphasis italics="true" pageId="0" pageNumber="99">in female</emphasis>
(p) anterior part of eyes covered by anterolaterally expanded clypeal shield.
</paragraph>
<paragraph pageId="0" pageNumber="99">Subimago and larva unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="species composition">
<paragraph pageId="0" pageNumber="99">Species composition.</paragraph>
<paragraph pageId="0" pageNumber="99">
<taxonomicName authorityName="Godunko &amp; Martynov &amp; Staniczek" authorityYear="2021" class="Insecta" family="Vietnamellidae" genus="Burmella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Burmella paucivenosa" order="Ephemeroptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="paucivenosa">
<emphasis italics="true" pageId="0" pageNumber="99">Burmella paucivenosa</emphasis>
</taxonomicName>
sp. nov. (SMNS; BU-179); 
<taxonomicName authorityName="Godunko &amp; Martynov &amp; Staniczek" authorityYear="2021" class="Insecta" family="Vietnamellidae" genus="Burmella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Burmella clypeata" order="Ephemeroptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="clypeata">
<emphasis italics="true" pageId="0" pageNumber="99">Burmella clypeata</emphasis>
</taxonomicName>
sp. nov. (SMNS; BU-321).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="locality and horizon">
<paragraph pageId="0" pageNumber="99">Locality and horizon.</paragraph>
<paragraph pageId="0" pageNumber="99">Hukawng Valley, Kachin State, Myanmar (Burma); Cenomanian, mid-Cretaceous.</paragraph>
<caption ID-Table-UUID="73C5555D08B56DEEA903CE63EDD2896A" httpUri="http://table.plazi.org/id/73C5555D08B56DEEA903CE63EDD2896A" pageId="0" pageNumber="99" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">Table 1.</emphasis>
Measurements of fossil representatives of the genus 
<taxonomicName authorityName="Godunko &amp; Martynov &amp; Staniczek" authorityYear="2021" class="Insecta" family="Vietnamellidae" genus="Burmella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Burmella" order="Ephemeroptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="99">Burmella</emphasis>
</taxonomicName>
gen. nov.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="99">
<table pageId="0" pageNumber="99">
<tr pageId="0" pageNumber="99">
<th colspan="1" pageId="0" pageNumber="99" rowspan="1">Adult characters</th>
<th colspan="1" pageId="0" pageNumber="99" rowspan="1">
<taxonomicName authorityName="Godunko &amp; Martynov &amp; Staniczek" authorityYear="2021" class="Insecta" family="Vietnamellidae" genus="Burmella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Burmella paucivenosa" order="Ephemeroptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="paucivenosa">
<emphasis italics="true" pageId="0" pageNumber="99">Burmella paucivenosa</emphasis>
</taxonomicName>
sp. nov. [SMNS, BU-179, male imago] (mm)
</th>
<th colspan="1" pageId="0" pageNumber="99" rowspan="1">
<taxonomicName authorityName="Godunko &amp; Martynov &amp; Staniczek" authorityYear="2021" class="Insecta" family="Vietnamellidae" genus="Burmella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Burmella clypeata" order="Ephemeroptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="clypeata">
<emphasis italics="true" pageId="0" pageNumber="99">Burmella clypeata</emphasis>
</taxonomicName>
sp. nov. [SMNS, BU-321, female imago] (mm)
</th>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of body</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">5.75</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">7.00</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of right foreleg</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">2.51*</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">1.14*</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of femur</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.83</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.42</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of tibia</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">1.68</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.72*</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of tarsus</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment I</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment II</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment III</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment IV</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment V</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of left foreleg</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">2.52*</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">1.64*</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of femur</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.85</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.46</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of tibia</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">1.67</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">1.18*</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of tarsus</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment I</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment II</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment III</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment IV</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment V</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of right middle leg</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">2.78</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of femur</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">1.45</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of tibia</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">1.03</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of tarsus</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.30</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment I</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.08</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment II</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.10</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment III</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.10</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment IV</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.11</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment V</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.14</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of left middle leg</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">2.70*</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">2.36</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of femur</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">1.20</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.60</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of tibia</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">1.02</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">1.34</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of tarsus</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.48*</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.42</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment I</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.08</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment II</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.07</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment III</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.07</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment IV</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.08</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment V</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.12</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of right hind leg</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">2.47</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">1.71</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of femur</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">1.02</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.71</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of tibia</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.93</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.56</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of tarsus</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.52</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.44</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment I</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.07</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.09</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment II</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.09</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.08</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment III</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.10</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.07</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment IV</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.12</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.08</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment V</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.14</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.12</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of left hind leg</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">2.07</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">2.60</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of femur</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">1.00</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.98</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of tibia</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.66</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">1.06</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of tarsus</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.41</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.56</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment I</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.05</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.11</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment II</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.07</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.11</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment III</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.07</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.10</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment IV</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.10</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.10</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Segment V</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.11</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.14</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of right forewing</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">4.64</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">1.80</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of left forewing</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">4.68</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">5.12</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of right hind wing</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.66</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.45</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of left hind wing</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.64</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Hind/Fore wings length ratio</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.14</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">0.08</td>
</tr>
<tr pageId="0" pageNumber="99">
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">Length of cerci [right/left]</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">2.35*/1.24*</td>
<td colspan="1" pageId="0" pageNumber="99" rowspan="1">8.12/-</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="99">
<tableNote pageId="0" pageNumber="99">* preserved part.</tableNote>
</paragraph>
</subSubSection>
</treatment>
</document>