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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. &amp; Tishechkin, Alexey K." docDate="2013" docId="207F9726F878A795600413B372FABD7A" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus occultus Caterino &amp; Tishechkin, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="120" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="118" updateTime="1668155382655" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caterino, Michael S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tishechkin, Alexey K.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>271</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>401</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.271.4062</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-271-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152040990" LSID="urn:lsid:zoobank.org:act:98A28E0E-C6B2-4406-9DC2-34D549763C8F" httpUri="http://treatment.plazi.org/id/207F9726F878A795600413B372FABD7A" lastPageId="119" lastPageNumber="120" pageId="117" pageNumber="118">
<subSubSection pageId="117" pageNumber="118" type="nomenclature">
<paragraph pageId="117" pageNumber="118">
<taxonomicName LSID="urn:lsid:zoobank.org:act:98A28E0E-C6B2-4406-9DC2-34D549763C8F" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus occultus" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="occultus">Operclipygus occultus</taxonomicName>
<taxonomicNameLabel pageId="117" pageNumber="118">sp. n.</taxonomicNameLabel>
Figs 34B,
<normalizedToken originalValue="DFMap">D-FMap</normalizedToken>
11
</paragraph>
</subSubSection>
<subSubSection pageId="118" pageNumber="119" type="type locality">
<paragraph pageId="118" pageNumber="119">
<pageBreakToken pageId="118" pageNumber="119" start="start">Type</pageBreakToken>
locality.
</paragraph>
<paragraph pageId="118" pageNumber="119">
ECUADOR: Orellana:
<taxonomicName lsidName="" pageId="118" pageNumber="119" rank="tribe" tribe="Tiputini">Tiputini</taxonomicName>
Biodiversity Station [
<geoCoordinate direction="south" orientation="latitude" precision="92" value="-0.63666666">0°38.2'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-76.14833">76°8.9'W</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection pageId="118" pageNumber="119" type="type material">
<paragraph pageId="118" pageNumber="119">Type material.</paragraph>
<paragraph pageId="118" pageNumber="119">
Holotype male: &quot;ECUADOR: Orellana,
<taxonomicName lsidName="" pageId="118" pageNumber="119" rank="tribe" tribe="Tiputini">Tiputini</taxonomicName>
Biodiversity Station
<geoCoordinate direction="south" orientation="latitude" precision="92" value="-0.63666666">0°38.2'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-76.14833">76°8.9'W</geoCoordinate>
. Flight intercept FIT7-1. 27-31 July 2008. A.K.Tishechkin&quot; / &quot;Caterino/Tishechkin
<taxonomicName lsidName="" pageId="118" pageNumber="119" rank="tribe" tribe="Exosternini">Exosternini</taxonomicName>
Voucher EXO-00283&quot; (FMNH).
</paragraph>
</subSubSection>
<subSubSection lastPageId="119" lastPageNumber="120" pageId="118" pageNumber="119" type="description">
<paragraph pageId="118" pageNumber="119">Diagnostic description.</paragraph>
<paragraph lastPageId="119" lastPageNumber="120" pageId="118" pageNumber="119">
This species is very similar to
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus dubius" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="dubius">Operclipygus dubius</taxonomicName>
, differing mainly in the following characters: length: 1.59 mm, width: 1.12 mm; frons weakly depressed, frontal stria complete; pronotal disk with prescutellar impression short and very narrow, few coarse punctures posterolaterad ends of anterior stria; lateral submarginal pronotal stria fine, present in anterior corners only; anterior submarginal pronotal stria fine, recurved posterolaterally to about one-third from anterior pronotal margin; elytra with dorsal stria 1 obsolete in apical third, striae 2-3 complete, 4th stria present in only apical half; prosternal keel with carinal striae subparallel in apical three-fourths, only weakly divergent basally; marginal mesoventral stria complete; metaventrite with shallow longitudinal depression; propygidium with dense, small, rather deep punctures; pygidium with fine dense ground punctation, coarser punctures dense along basal margin, sparsely scattered in apical two-thirds; marginal pygidial sulcus deep,
<pageBreakToken pageId="119" pageNumber="120" start="start">coarsely</pageBreakToken>
crenulate. Male genitalia (Figs 33B,
<normalizedToken originalValue="DF">D-F</normalizedToken>
) as for group description, the apices of S8 strongly narrowed, with only a couple inconspicuous inner apical setae, curved to ventral apex, bearing 4 or 5 long apical setae, with distinct submarginal sclerotization along apical margins; T9 with apices elongate subtriangular, inner edges parallel; T10 not divided; S9 not desclerotized along midline, truncate at base, with shallow, equilateral apical emargination; tegmen widening to near apex, narrowed to truncate apex; median lobe narrowed proximad gonopore.
</paragraph>
</subSubSection>
<subSubSection pageId="119" pageNumber="120" type="remarks">
<paragraph pageId="119" pageNumber="120">Remarks.</paragraph>
<paragraph pageId="119" pageNumber="120">
This species can only confidently be recognized by the male genitalia (Figs 33B,
<normalizedToken originalValue="DF">D-F</normalizedToken>
). The elongate apical arms of T9 and the narrow, subacute apices of S9 are both highly distinctive in the group. A few other characters, such as the median longitudinal depression of the metaventrite and the complete frontal stria may be helpful, but with only a single available specimen it is impossible to ascertain their significance.
</paragraph>
<caption pageId="119" pageNumber="120">
<paragraph pageId="119" pageNumber="120">
Figure 34.
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hospes" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="hospes">Operclipygus hospes</taxonomicName>
group. A Metaventrite and 1st abdominal ventrite of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fusistrius" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="fusistrius">Operclipygus fusistrius</taxonomicName>
B Frons of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hospes" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="hospes">Operclipygus hospes</taxonomicName>
(lectotype) C Dorsal habitus of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hospes" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="hospes">Operclipygus hospes</taxonomicName>
(lectotype) D Pygidia of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hospes" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="hospes">Operclipygus hospes</taxonomicName>
(lectotype) E Dorsal habitus of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus ignifer" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="ignifer">Operclipygus ignifer</taxonomicName>
F Dorsal habitus
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus subterraneus" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="subterraneus">Operclipygus subterraneus</taxonomicName>
G Frons of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus subterraneus" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="subterraneus">Operclipygus subterraneus</taxonomicName>
H Pygidia of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus subterraneus" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="subterraneus">Operclipygus subterraneus</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="119" pageNumber="120" type="etymology">
<paragraph pageId="119" pageNumber="120">Etymology.</paragraph>
<paragraph pageId="119" pageNumber="120">
This
<normalizedToken originalValue="species">species'</normalizedToken>
name refers to the fact that hidden characters must be observed for identification.
</paragraph>
</subSubSection>
</treatment>
</document>