405 lines
60 KiB
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405 lines
60 KiB
XML
<document id="8616C037A7539614DA6341A5C829E431" ID-DOI="10.11646/zootaxa.4040.3.10" ID-GBIF-Dataset="faff8f58-3123-4b54-8c6a-00c9ca12d3ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="237704" ID-ZooBank="40FA2451-3A46-461B-98B4-F98940FE1FE1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461336224380" checkinUser="plazi" docAuthor="Liu, Xingyue & Hajong, Sudhanya Ray" docDate="2015" docId="1D7F574AFFF2110D80D7FB90B9F7FE25" docLanguage="en" docName="zt04040p400.pdf" docOrigin="Zootaxa 4040 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Inocellia indica Liu & Hajong, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="398" masterDocId="E1462F32FFF3110B8040FFA6BB4DFFAD" masterDocTitle="Inocellia indica sp. nov. (Raphidioptera: Inocelliidae): a new snakefly species from northeastern India" masterLastPageNumber="400" masterPageNumber="393" pageNumber="394" updateTime="1698619275991" updateUser="plazi">
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<mods:titleInfo id="DB750A687DAC6C21D3671234A952208F">
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<mods:title id="1336D4992C6C425E91786B2BDA4B27EF">Inocellia indica sp. nov. (Raphidioptera: Inocelliidae): a new snakefly species from northeastern India</mods:title>
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<mods:name id="C6FD01D10619B0EC271753DBA689FE16" type="personal">
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<mods:namePart id="67F082EAD4FFB485243A703DAD89D1F0">Liu, Xingyue</mods:namePart>
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<mods:namePart id="345561341EC193489CB9F6C44B58BF53">Hajong, Sudhanya Ray</mods:namePart>
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<mods:title id="A113FE95BC33CD0A89B8D56BF5B3E473">Zootaxa</mods:title>
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<mods:part id="504588A3DB9DAD32BBD9D862013D97E2">
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<mods:date id="47D78DEB349CD24CEE180FAA5CC46E3C">2015</mods:date>
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<mods:number id="B2E8355DBA1A92BDAB474D7CB84BAAF5">4040</mods:number>
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<mods:number id="006A97A37377E60F725FB7AE54BFB181">3</mods:number>
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<mods:identifier id="777D60663A346B12D1708F7978315C47" type="DOI">10.11646/zootaxa.4040.3.10</mods:identifier>
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<treatment id="1D7F574AFFF2110D80D7FB90B9F7FE25" ID-DOI="http://doi.org/10.5281/zenodo.6112590" ID-GBIF-Taxon="119644557" ID-Zenodo-Dep="6112590" LSID="urn:lsid:plazi:treatment:1D7F574AFFF2110D80D7FB90B9F7FE25" httpUri="http://treatment.plazi.org/id/1D7F574AFFF2110D80D7FB90B9F7FE25" lastPageId="6" lastPageNumber="398" pageId="1" pageNumber="394">
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<subSubSection id="DDCCB5D7FFF2110A80D7FB90BA6DFBDF" pageId="1" pageNumber="394" type="nomenclature">
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<paragraph id="9569E65CFFF2110A80D7FB90BAF4FBFD" blockId="1.[151,441,1078,1138]" box="[151,441,1078,1104]" pageId="1" pageNumber="394">
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<heading id="CE215130FFF2110A80D7FB90BAF4FBFD" bold="true" box="[151,441,1078,1104]" fontSize="11" level="1" pageId="1" pageNumber="394" reason="1">
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<emphasis id="A7A23A4EFFF2110A80D7FB90BAF4FBFD" bold="true" box="[151,441,1078,1104]" pageId="1" pageNumber="394">
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<taxonomicName id="52D69DDFFFF2110A80D7FB90BA18FBFD" ID-CoL="3PQGQ" box="[151,341,1078,1104]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
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<emphasis id="A7A23A4EFFF2110A80D7FB90BA18FBFD" bold="true" box="[151,341,1078,1104]" italics="true" pageId="1" pageNumber="394">Inocellia indica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BC918735FFF2110A811CFB90BAF4FBFD" box="[348,441,1078,1104]" pageId="1" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="9569E65CFFF2110A80D7FBFFBA6DFBDF" blockId="1.[151,441,1078,1138]" box="[151,288,1113,1138]" pageId="1" pageNumber="394">
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(
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<figureCitation id="0DEDFAD9FFF2110A80DFFBFFBA5AFBDF" box="[159,279,1113,1138]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURES 3 – 12" captionStart-2="FIGURE 13" captionStartId-0="2.[151,264,1958,1980]" captionStartId-1="3.[151,264,1726,1748]" captionStartId-2="4.[151,250,1990,2012]" captionTargetBox-0="[316,1237,221,1933]" captionTargetBox-1="[152,1428,198,1704]" captionTargetBox-2="[210,1377,1093,1968]" captionTargetId-0="figure@2.[205,1381,195,1934]" captionTargetId-1="figure@3.[152,1435,193,1705]" captionTargetId-2="figure@4.[210,1377,1093,1969]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1 – 2. Habitus images of Inocellia indica sp. nov. 1 — holotype male; 2 — paratype female. Scale bar = 1.0 mm." captionText-1="FIGURES 3 – 12. Inocellia indica sp. nov. 3 — male genital segments, lateral view; 4 — male genital segments, dorsal view; 5 — male genital segments, ventral view; 6 — male genital segments, caudal view; 7 — male ninth gonocoxite with internal structures, lateral view; 8 — gonarcus, dorsal view; 9 — fused parameres, dorsal view; 10 — female genital segments, lateral view; 11 — female sternite 7 and subgenitale, ventral view; 12 — bursa copulatrix, lateral view. Abbreviations: ab — atrium bursae; e — ectoproct; ep — endophallus; g — gonarcus (= fused gonocoxites 11); gr — glandula receptaculi; gx 9 — gonocoxite 9; hi — hypandrium internum; o — ovipositor; p — fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10); ps — pseudostyli (= gonapophyses 9); r — receptaculum seminis; S — sternum; sb — sacculus bursae; ser — serratulum; sg — subgenitale; st — gonostylus 9; T — tergum. Arrow indicates ventral projection of male gonocoxite 9. Scale bar = 0.5 mm." captionText-2="FIGURE 13. Habitat of Inocellia indica sp. nov." httpUri-0="https://zenodo.org/record/237705/files/figure.png" httpUri-1="https://zenodo.org/record/237706/files/figure.png" httpUri-2="https://zenodo.org/record/237707/files/figure.png" pageId="1" pageNumber="394">Figs. 1–13</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="DDCCB5D7FFF2110F80D7FB07B979FEED" lastPageId="4" lastPageNumber="397" pageId="1" pageNumber="394" type="description">
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<paragraph id="9569E65CFFF2110A80D7FB07BFF4FB14" blockId="1.[151,1437,1185,2001]" box="[151,1209,1185,1210]" pageId="1" pageNumber="394">
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<emphasis id="A7A23A4EFFF2110A80D7FB07BA64FB17" bold="true" box="[151,297,1185,1210]" pageId="1" pageNumber="394">Description.</emphasis>
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Male. Body length
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<quantity id="522E4BB9FFF2110A8247FB04B923FB14" box="[519,622,1186,1210]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="1" pageNumber="394" unit="mm" value="14.3">14.3 mm</quantity>
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; forewing length
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<quantity id="522E4BB9FFF2110A8373FB07B8C6FB14" box="[819,907,1185,1210]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="1" pageNumber="394" unit="mm" value="9.6">9.6 mm</quantity>
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, hind wing length
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<quantity id="522E4BB9FFF2110A841BFB04BFF9FB14" box="[1115,1204,1186,1210]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="1" pageNumber="394" unit="mm" value="8.1">8.1 mm</quantity>
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.
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</paragraph>
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<paragraph id="9569E65CFFF2110A8087FB63BA9FFAAF" blockId="1.[151,1437,1185,2001]" pageId="1" pageNumber="394">
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Head (
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<figureCitation id="0DEDFAD9FFF2110A815AFB63BA2EFB73" box="[282,355,1221,1246]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1958,1980]" captionTargetBox="[316,1237,221,1933]" captionTargetId="figure@2.[205,1381,195,1934]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus images of Inocellia indica sp. nov. 1 — holotype male; 2 — paratype female. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/237705/files/figure.png" pageId="1" pageNumber="394">Fig. 1</figureCitation>
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) subquadrate, black throughout; clypeus pale brown. Antennae brown, but scape black. Mouthparts blackish brown.
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</paragraph>
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<paragraph id="9569E65CFFF2110A8087FAABBE18FAE7" blockId="1.[151,1437,1185,2001]" pageId="1" pageNumber="394">
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Thorax (
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<figureCitation id="0DEDFAD9FFF2110A816AFAABBA23FA8B" box="[298,366,1293,1318]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1958,1980]" captionTargetBox="[316,1237,221,1933]" captionTargetId="figure@2.[205,1381,195,1934]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus images of Inocellia indica sp. nov. 1 — holotype male; 2 — paratype female. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/237705/files/figure.png" pageId="1" pageNumber="394">Fig. 1</figureCitation>
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) entirely black. Legs yellow, slightly darker on tibiae, with yellowish setae. Wings hyaline, pterostigma dark brown, veins brown. Rs with one forked vein and one simple vein running to wing margin.
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</paragraph>
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<paragraph id="9569E65CFFF2110A8087FAF3BF81F827" blockId="1.[151,1437,1185,2001]" pageId="1" pageNumber="394">
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Abdomen (
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<figureCitation id="0DEDFAD9FFF2110A8108FAF3BAC1FAC3" box="[328,396,1365,1390]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1958,1980]" captionTargetBox="[316,1237,221,1933]" captionTargetId="figure@2.[205,1381,195,1934]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus images of Inocellia indica sp. nov. 1 — holotype male; 2 — paratype female. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/237705/files/figure.png" pageId="1" pageNumber="394">Fig. 1</figureCitation>
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) blackish brown; each pregenital segment ventrally with a yellowish transverse stripe on posterior margin, and with pleural areas continuously pale yellow; genital segments orange, but slightly paler on posterior half of gonocoxites 9. Tergum 9 (
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<figureCitation id="0DEDFAD9FFF2110A82C2FA3BB98BFA1B" box="[642,710,1437,1462]" captionStart="FIGURES 3 – 12" captionStartId="3.[151,264,1726,1748]" captionTargetBox="[152,1428,198,1704]" captionTargetId="figure@3.[152,1435,193,1705]" captionTargetPageId="3" captionText="FIGURES 3 – 12. Inocellia indica sp. nov. 3 — male genital segments, lateral view; 4 — male genital segments, dorsal view; 5 — male genital segments, ventral view; 6 — male genital segments, caudal view; 7 — male ninth gonocoxite with internal structures, lateral view; 8 — gonarcus, dorsal view; 9 — fused parameres, dorsal view; 10 — female genital segments, lateral view; 11 — female sternite 7 and subgenitale, ventral view; 12 — bursa copulatrix, lateral view. Abbreviations: ab — atrium bursae; e — ectoproct; ep — endophallus; g — gonarcus (= fused gonocoxites 11); gr — glandula receptaculi; gx 9 — gonocoxite 9; hi — hypandrium internum; o — ovipositor; p — fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10); ps — pseudostyli (= gonapophyses 9); r — receptaculum seminis; S — sternum; sb — sacculus bursae; ser — serratulum; sg — subgenitale; st — gonostylus 9; T — tergum. Arrow indicates ventral projection of male gonocoxite 9. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/237706/files/figure.png" pageId="1" pageNumber="394">Fig. 3</figureCitation>
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) approximately 1.5 times as long as sternum 9, anterior margin slightly incised medially, posterior margin shallowly incised. Sternum 9 (
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<figureCitation id="0DEDFAD9FFF2110A83ACFA67BF7FFA77" box="[1004,1074,1473,1498]" captionStart="FIGURES 3 – 12" captionStartId="3.[151,264,1726,1748]" captionTargetBox="[152,1428,198,1704]" captionTargetId="figure@3.[152,1435,193,1705]" captionTargetPageId="3" captionText="FIGURES 3 – 12. Inocellia indica sp. nov. 3 — male genital segments, lateral view; 4 — male genital segments, dorsal view; 5 — male genital segments, ventral view; 6 — male genital segments, caudal view; 7 — male ninth gonocoxite with internal structures, lateral view; 8 — gonarcus, dorsal view; 9 — fused parameres, dorsal view; 10 — female genital segments, lateral view; 11 — female sternite 7 and subgenitale, ventral view; 12 — bursa copulatrix, lateral view. Abbreviations: ab — atrium bursae; e — ectoproct; ep — endophallus; g — gonarcus (= fused gonocoxites 11); gr — glandula receptaculi; gx 9 — gonocoxite 9; hi — hypandrium internum; o — ovipositor; p — fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10); ps — pseudostyli (= gonapophyses 9); r — receptaculum seminis; S — sternum; sb — sacculus bursae; ser — serratulum; sg — subgenitale; st — gonostylus 9; T — tergum. Arrow indicates ventral projection of male gonocoxite 9. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/237706/files/figure.png" pageId="1" pageNumber="394">Fig. 5</figureCitation>
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) broadly arcuate, with anterior margin prominent. Gonocoxite 9 (
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<figureCitation id="0DEDFAD9FFF2110A826BFA43B9D8FA53" box="[555,661,1509,1534]" captionStart="FIGURES 3 – 12" captionStartId="3.[151,264,1726,1748]" captionTargetBox="[152,1428,198,1704]" captionTargetId="figure@3.[152,1435,193,1705]" captionTargetPageId="3" captionText="FIGURES 3 – 12. Inocellia indica sp. nov. 3 — male genital segments, lateral view; 4 — male genital segments, dorsal view; 5 — male genital segments, ventral view; 6 — male genital segments, caudal view; 7 — male ninth gonocoxite with internal structures, lateral view; 8 — gonarcus, dorsal view; 9 — fused parameres, dorsal view; 10 — female genital segments, lateral view; 11 — female sternite 7 and subgenitale, ventral view; 12 — bursa copulatrix, lateral view. Abbreviations: ab — atrium bursae; e — ectoproct; ep — endophallus; g — gonarcus (= fused gonocoxites 11); gr — glandula receptaculi; gx 9 — gonocoxite 9; hi — hypandrium internum; o — ovipositor; p — fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10); ps — pseudostyli (= gonapophyses 9); r — receptaculum seminis; S — sternum; sb — sacculus bursae; ser — serratulum; sg — subgenitale; st — gonostylus 9; T — tergum. Arrow indicates ventral projection of male gonocoxite 9. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/237706/files/figure.png" pageId="1" pageNumber="394">Figs 3–6</figureCitation>
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) shell-like, strongly elongate, ~1.5 times as long as width of its proximal portion, proximally with a subtriangular projection on ventral margin; stylus (gonostylus 9) present subdistad on inner surface of gonocoxite 9, flatly and broadly expanded, feebly domed, with rows of bristles; an additional bristle tuft present proximally near dorsal margin. Pseudostyli (gonapophyses 9) (
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<figureCitation id="0DEDFAD9FFF2110A8499F9F7BE0FF9C7" box="[1241,1346,1617,1642]" captionStart="FIGURES 3 – 12" captionStartId="3.[151,264,1726,1748]" captionTargetBox="[152,1428,198,1704]" captionTargetId="figure@3.[152,1435,193,1705]" captionTargetPageId="3" captionText="FIGURES 3 – 12. Inocellia indica sp. nov. 3 — male genital segments, lateral view; 4 — male genital segments, dorsal view; 5 — male genital segments, ventral view; 6 — male genital segments, caudal view; 7 — male ninth gonocoxite with internal structures, lateral view; 8 — gonarcus, dorsal view; 9 — fused parameres, dorsal view; 10 — female genital segments, lateral view; 11 — female sternite 7 and subgenitale, ventral view; 12 — bursa copulatrix, lateral view. Abbreviations: ab — atrium bursae; e — ectoproct; ep — endophallus; g — gonarcus (= fused gonocoxites 11); gr — glandula receptaculi; gx 9 — gonocoxite 9; hi — hypandrium internum; o — ovipositor; p — fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10); ps — pseudostyli (= gonapophyses 9); r — receptaculum seminis; S — sternum; sb — sacculus bursae; ser — serratulum; sg — subgenitale; st — gonostylus 9; T — tergum. Arrow indicates ventral projection of male gonocoxite 9. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/237706/files/figure.png" pageId="1" pageNumber="394">Figs 5–7</figureCitation>
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) feebly sclerotized, foliate, slightly curved anteriorly in ventral view. Fused parameres (complex of gonocoxites, gonapophyses, gonostyli 10) (
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<figureCitation id="0DEDFAD9FFF2110A81A6F93FB929F91F" box="[486,612,1689,1714]" captionStart="FIGURES 3 – 12" captionStartId="3.[151,264,1726,1748]" captionTargetBox="[152,1428,198,1704]" captionTargetId="figure@3.[152,1435,193,1705]" captionTargetPageId="3" captionText="FIGURES 3 – 12. Inocellia indica sp. nov. 3 — male genital segments, lateral view; 4 — male genital segments, dorsal view; 5 — male genital segments, ventral view; 6 — male genital segments, caudal view; 7 — male ninth gonocoxite with internal structures, lateral view; 8 — gonarcus, dorsal view; 9 — fused parameres, dorsal view; 10 — female genital segments, lateral view; 11 — female sternite 7 and subgenitale, ventral view; 12 — bursa copulatrix, lateral view. Abbreviations: ab — atrium bursae; e — ectoproct; ep — endophallus; g — gonarcus (= fused gonocoxites 11); gr — glandula receptaculi; gx 9 — gonocoxite 9; hi — hypandrium internum; o — ovipositor; p — fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10); ps — pseudostyli (= gonapophyses 9); r — receptaculum seminis; S — sternum; sb — sacculus bursae; ser — serratulum; sg — subgenitale; st — gonostylus 9; T — tergum. Arrow indicates ventral projection of male gonocoxite 9. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/237706/files/figure.png" pageId="1" pageNumber="394">Figs 6–7, 9</figureCitation>
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) flattened on proximal portion, ventrally bearing a slender and long distal projection, which is distally vertically curved dorsad and rounded at tip. Gonarcus (fused gonocoxites 11) (
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<figureCitation id="0DEDFAD9FFF2110A8507F91BBBE8F957" captionStart="FIGURES 3 – 12" captionStartId="3.[151,264,1726,1748]" captionTargetBox="[152,1428,198,1704]" captionTargetId="figure@3.[152,1435,193,1705]" captionTargetPageId="3" captionText="FIGURES 3 – 12. Inocellia indica sp. nov. 3 — male genital segments, lateral view; 4 — male genital segments, dorsal view; 5 — male genital segments, ventral view; 6 — male genital segments, caudal view; 7 — male ninth gonocoxite with internal structures, lateral view; 8 — gonarcus, dorsal view; 9 — fused parameres, dorsal view; 10 — female genital segments, lateral view; 11 — female sternite 7 and subgenitale, ventral view; 12 — bursa copulatrix, lateral view. Abbreviations: ab — atrium bursae; e — ectoproct; ep — endophallus; g — gonarcus (= fused gonocoxites 11); gr — glandula receptaculi; gx 9 — gonocoxite 9; hi — hypandrium internum; o — ovipositor; p — fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10); ps — pseudostyli (= gonapophyses 9); r — receptaculum seminis; S — sternum; sb — sacculus bursae; ser — serratulum; sg — subgenitale; st — gonostylus 9; T — tergum. Arrow indicates ventral projection of male gonocoxite 9. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/237706/files/figure.png" pageId="1" pageNumber="394">Figs 5– 8</figureCitation>
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) broad, nearly transversely rectangular, with ventrolateral corners prominent, and with nearly truncate ventral margin in caudal view, medially bluntly produced posteriad. Endophallus (
|
||
<figureCitation id="0DEDFAD9FFF2110A83A6F8A3BF06F8B3" box="[998,1099,1797,1822]" captionStart="FIGURES 3 – 12" captionStartId="3.[151,264,1726,1748]" captionTargetBox="[152,1428,198,1704]" captionTargetId="figure@3.[152,1435,193,1705]" captionTargetPageId="3" captionText="FIGURES 3 – 12. Inocellia indica sp. nov. 3 — male genital segments, lateral view; 4 — male genital segments, dorsal view; 5 — male genital segments, ventral view; 6 — male genital segments, caudal view; 7 — male ninth gonocoxite with internal structures, lateral view; 8 — gonarcus, dorsal view; 9 — fused parameres, dorsal view; 10 — female genital segments, lateral view; 11 — female sternite 7 and subgenitale, ventral view; 12 — bursa copulatrix, lateral view. Abbreviations: ab — atrium bursae; e — ectoproct; ep — endophallus; g — gonarcus (= fused gonocoxites 11); gr — glandula receptaculi; gx 9 — gonocoxite 9; hi — hypandrium internum; o — ovipositor; p — fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10); ps — pseudostyli (= gonapophyses 9); r — receptaculum seminis; S — sternum; sb — sacculus bursae; ser — serratulum; sg — subgenitale; st — gonostylus 9; T — tergum. Arrow indicates ventral projection of male gonocoxite 9. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/237706/files/figure.png" pageId="1" pageNumber="394">Figs 5–7</figureCitation>
|
||
) short, lateral portions with a pair of bristle tufts directed posterodorsad. Scabrous membranous serratulum present, with a pair of dorsally directed bristle tufts (
|
||
<figureCitation id="0DEDFAD9FFF2110A81D6F8EBB94EF8CB" box="[406,515,1869,1894]" captionStart="FIGURES 3 – 12" captionStartId="3.[151,264,1726,1748]" captionTargetBox="[152,1428,198,1704]" captionTargetId="figure@3.[152,1435,193,1705]" captionTargetPageId="3" captionText="FIGURES 3 – 12. Inocellia indica sp. nov. 3 — male genital segments, lateral view; 4 — male genital segments, dorsal view; 5 — male genital segments, ventral view; 6 — male genital segments, caudal view; 7 — male ninth gonocoxite with internal structures, lateral view; 8 — gonarcus, dorsal view; 9 — fused parameres, dorsal view; 10 — female genital segments, lateral view; 11 — female sternite 7 and subgenitale, ventral view; 12 — bursa copulatrix, lateral view. Abbreviations: ab — atrium bursae; e — ectoproct; ep — endophallus; g — gonarcus (= fused gonocoxites 11); gr — glandula receptaculi; gx 9 — gonocoxite 9; hi — hypandrium internum; o — ovipositor; p — fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10); ps — pseudostyli (= gonapophyses 9); r — receptaculum seminis; S — sternum; sb — sacculus bursae; ser — serratulum; sg — subgenitale; st — gonostylus 9; T — tergum. Arrow indicates ventral projection of male gonocoxite 9. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/237706/files/figure.png" pageId="1" pageNumber="394">Figs 5–7</figureCitation>
|
||
). Ectoproct (
|
||
<figureCitation id="0DEDFAD9FFF2110A82DDF8EBB84BF8CB" box="[669,774,1869,1894]" captionStart="FIGURES 3 – 12" captionStartId="3.[151,264,1726,1748]" captionTargetBox="[152,1428,198,1704]" captionTargetId="figure@3.[152,1435,193,1705]" captionTargetPageId="3" captionText="FIGURES 3 – 12. Inocellia indica sp. nov. 3 — male genital segments, lateral view; 4 — male genital segments, dorsal view; 5 — male genital segments, ventral view; 6 — male genital segments, caudal view; 7 — male ninth gonocoxite with internal structures, lateral view; 8 — gonarcus, dorsal view; 9 — fused parameres, dorsal view; 10 — female genital segments, lateral view; 11 — female sternite 7 and subgenitale, ventral view; 12 — bursa copulatrix, lateral view. Abbreviations: ab — atrium bursae; e — ectoproct; ep — endophallus; g — gonarcus (= fused gonocoxites 11); gr — glandula receptaculi; gx 9 — gonocoxite 9; hi — hypandrium internum; o — ovipositor; p — fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10); ps — pseudostyli (= gonapophyses 9); r — receptaculum seminis; S — sternum; sb — sacculus bursae; ser — serratulum; sg — subgenitale; st — gonostylus 9; T — tergum. Arrow indicates ventral projection of male gonocoxite 9. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/237706/files/figure.png" pageId="1" pageNumber="394">Figs 3–4</figureCitation>
|
||
) ovoid in lateral view, and subquadrate in dorsal view. Hypandrium internum (
|
||
<figureCitation id="0DEDFAD9FFF2110A81DFF8D7B94EF827" box="[415,515,1905,1930]" captionStart="FIGURES 3 – 12" captionStartId="3.[151,264,1726,1748]" captionTargetBox="[152,1428,198,1704]" captionTargetId="figure@3.[152,1435,193,1705]" captionTargetPageId="3" captionText="FIGURES 3 – 12. Inocellia indica sp. nov. 3 — male genital segments, lateral view; 4 — male genital segments, dorsal view; 5 — male genital segments, ventral view; 6 — male genital segments, caudal view; 7 — male ninth gonocoxite with internal structures, lateral view; 8 — gonarcus, dorsal view; 9 — fused parameres, dorsal view; 10 — female genital segments, lateral view; 11 — female sternite 7 and subgenitale, ventral view; 12 — bursa copulatrix, lateral view. Abbreviations: ab — atrium bursae; e — ectoproct; ep — endophallus; g — gonarcus (= fused gonocoxites 11); gr — glandula receptaculi; gx 9 — gonocoxite 9; hi — hypandrium internum; o — ovipositor; p — fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10); ps — pseudostyli (= gonapophyses 9); r — receptaculum seminis; S — sternum; sb — sacculus bursae; ser — serratulum; sg — subgenitale; st — gonostylus 9; T — tergum. Arrow indicates ventral projection of male gonocoxite 9. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/237706/files/figure.png" pageId="1" pageNumber="394">Figs 5, 7</figureCitation>
|
||
) small with lateral lobes foliate and slightly sinuate posteriorly.
|
||
</paragraph>
|
||
<paragraph id="9569E65CFFF2110A8087F833BB87F87C" blockId="1.[151,1437,1185,2001]" pageId="1" pageNumber="394">
|
||
Female. Body length
|
||
<quantity id="522E4BB9FFF2110A81FAF830B952F800" box="[442,543,1942,1966]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="1" pageNumber="394" unit="mm" value="12.9">12.9 mm</quantity>
|
||
(without ovipositor)/
|
||
<quantity id="522E4BB9FFF2110A8352F833B835F800" box="[786,888,1941,1966]" metricMagnitude="-2" metricUnit="m" metricValue="1.88" pageId="1" pageNumber="394" unit="mm" value="18.8">18.8 mm</quantity>
|
||
; forewing length
|
||
<quantity id="522E4BB9FFF2110A847FF830BFEBF800" box="[1087,1190,1942,1966]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="1" pageNumber="394" unit="mm" value="12.1">12.1 mm</quantity>
|
||
, hind wing length
|
||
<quantity id="522E4BB9FFF2110A8539F830BB8BF87C" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="1" pageNumber="394" unit="mm" value="9.9">9.9 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<caption id="C1A9B6D4FFF1110980D7F800BE2CF811" box="[151,1377,1958,1980]" httpUri="https://zenodo.org/record/237705/files/figure.png" pageId="2" pageNumber="395" targetBox="[316,1237,221,1933]" targetPageId="2">
|
||
<paragraph id="9569E65CFFF1110980D7F800BE2CF811" blockId="2.[151,1377,1958,1980]" box="[151,1377,1958,1980]" pageId="2" pageNumber="395">
|
||
<emphasis id="A7A23A4EFFF1110980D7F800BA77F811" bold="true" box="[151,314,1958,1980]" pageId="2" pageNumber="395">FIGURES 1–2.</emphasis>
|
||
Habitus images of
|
||
<taxonomicName id="52D69DDFFFF1110981BFF800B9D3F811" box="[511,670,1958,1980]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="2" pageNumber="395" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
|
||
<emphasis id="A7A23A4EFFF1110981BFF800B9D3F811" box="[511,670,1958,1980]" italics="true" pageId="2" pageNumber="395">Inocellia indica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A7A23A4EFFF1110982E4F800B845F811" bold="true" box="[676,776,1958,1980]" pageId="2" pageNumber="395">
|
||
<taxonomicNameLabel id="BC918735FFF1110982E4F800B9B8F811" box="[676,757,1958,1980]" pageId="2" pageNumber="395" rank="species">sp. nov.</taxonomicNameLabel>
|
||
1
|
||
</emphasis>
|
||
—holotype male;
|
||
<emphasis id="A7A23A4EFFF1110983FFF800B886F811" bold="true" box="[959,971,1958,1980]" pageId="2" pageNumber="395">2</emphasis>
|
||
—paratype female. Scale bar = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C1A9B6D4FFF0110880D7F918BE2CF806" httpUri="https://zenodo.org/record/237706/files/figure.png" pageId="3" pageNumber="396" targetBox="[152,1428,198,1704]" targetPageId="3">
|
||
<paragraph id="9569E65CFFF0110880D7F918BE2CF806" blockId="3.[151,1436,1726,1963]" pageId="3" pageNumber="396">
|
||
<emphasis id="A7A23A4EFFF0110880D7F918BA0BF979" bold="true" box="[151,326,1726,1748]" pageId="3" pageNumber="396">FIGURES 3–12.</emphasis>
|
||
<taxonomicName id="52D69DDFFFF01108810CF918BAA1F979" box="[332,492,1726,1748]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
|
||
<emphasis id="A7A23A4EFFF01108810CF918BAA1F979" box="[332,492,1726,1748]" italics="true" pageId="3" pageNumber="396">Inocellia indica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A7A23A4EFFF0110881B2F918B918F979" bold="true" box="[498,597,1726,1748]" pageId="3" pageNumber="396">
|
||
<taxonomicNameLabel id="BC918735FFF0110881B2F918B90EF979" box="[498,579,1726,1748]" pageId="3" pageNumber="396" rank="species">sp. nov.</taxonomicNameLabel>
|
||
3
|
||
</emphasis>
|
||
—male genital segments, lateral view;
|
||
<emphasis id="A7A23A4EFFF0110883A0F918B8A1F979" bold="true" box="[992,1004,1726,1748]" pageId="3" pageNumber="396">4</emphasis>
|
||
—male genital segments, dorsal view;
|
||
<emphasis id="A7A23A4EFFF011088536F918BECEF979" bold="true" box="[1398,1411,1726,1748]" pageId="3" pageNumber="396">5</emphasis>
|
||
— male genital segments, ventral view;
|
||
<emphasis id="A7A23A4EFFF011088273F97BB90DF95E" bold="true" box="[563,576,1757,1779]" pageId="3" pageNumber="396">6</emphasis>
|
||
—male genital segments, caudal view;
|
||
<emphasis id="A7A23A4EFFF0110883B0F97BB8B0F95E" bold="true" box="[1008,1021,1757,1779]" pageId="3" pageNumber="396">7</emphasis>
|
||
—male ninth gonocoxite with internal structures, lateral view;
|
||
<emphasis id="A7A23A4EFFF0110881D6F95DBAEEF8BC" bold="true" box="[406,419,1787,1809]" pageId="3" pageNumber="396">8</emphasis>
|
||
—gonarcus, dorsal view;
|
||
<emphasis id="A7A23A4EFFF0110882F3F95DB9F2F8BC" bold="true" box="[691,703,1787,1809]" pageId="3" pageNumber="396">9</emphasis>
|
||
—fused parameres, dorsal view;
|
||
<emphasis id="A7A23A4EFFF01108845DF95DBF7BF8BC" bold="true" box="[1053,1078,1787,1809]" pageId="3" pageNumber="396">10</emphasis>
|
||
—female genital segments, lateral view;
|
||
<emphasis id="A7A23A4EFFF01108809AF8BCBBBCF89D" bold="true" box="[218,241,1818,1840]" pageId="3" pageNumber="396">11</emphasis>
|
||
—female sternite 7 and subgenitale, ventral view;
|
||
<emphasis id="A7A23A4EFFF01108834EF8BCB86AF89D" bold="true" box="[782,807,1818,1840]" pageId="3" pageNumber="396">12</emphasis>
|
||
—bursa copulatrix, lateral view. Abbreviations:
|
||
<emphasis id="A7A23A4EFFF011088565F8BCBE0DF89D" bold="true" box="[1317,1344,1818,1840]" pageId="3" pageNumber="396">ab</emphasis>
|
||
—atrium bursae;
|
||
<emphasis id="A7A23A4EFFF0110880A7F89FBBBFF8E2" bold="true" box="[231,242,1849,1871]" pageId="3" pageNumber="396">e</emphasis>
|
||
—ectoproct;
|
||
<emphasis id="A7A23A4EFFF011088137F89FBADDF8E2" bold="true" box="[375,400,1849,1871]" pageId="3" pageNumber="396">ep</emphasis>
|
||
—endophallus;
|
||
<emphasis id="A7A23A4EFFF011088272F89FB973F8E2" bold="true" box="[562,574,1849,1871]" pageId="3" pageNumber="396">g</emphasis>
|
||
—gonarcus (= fused gonocoxites 11);
|
||
<emphasis id="A7A23A4EFFF011088385F89FB890F8E2" bold="true" box="[965,989,1849,1871]" pageId="3" pageNumber="396">gr</emphasis>
|
||
—glandula receptaculi;
|
||
<emphasis id="A7A23A4EFFF011088492F89FBFBAF8E2" bold="true" box="[1234,1271,1849,1871]" pageId="3" pageNumber="396">gx9</emphasis>
|
||
—gonocoxite 9;
|
||
<emphasis id="A7A23A4EFFF0110880D7F8F1BBE1F8C0" bold="true" box="[151,172,1879,1901]" pageId="3" pageNumber="396">hi</emphasis>
|
||
—hypandrium internum;
|
||
<emphasis id="A7A23A4EFFF0110881F3F8F1BAF2F8C0" bold="true" box="[435,447,1879,1901]" pageId="3" pageNumber="396">o</emphasis>
|
||
—ovipositor;
|
||
<emphasis id="A7A23A4EFFF01108820EF8F1B911F8C0" bold="true" box="[590,604,1879,1901]" pageId="3" pageNumber="396">p</emphasis>
|
||
—fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10);
|
||
<emphasis id="A7A23A4EFFF011088086F8D0BB93F821" bold="true" box="[198,222,1910,1932]" pageId="3" pageNumber="396">ps</emphasis>
|
||
—pseudostyli (= gonapophyses 9);
|
||
<emphasis id="A7A23A4EFFF011088207F8D0B91FF821" bold="true" box="[583,594,1910,1932]" pageId="3" pageNumber="396">r</emphasis>
|
||
—receptaculum seminis;
|
||
<emphasis id="A7A23A4EFFF011088315F8D0B82EF821" bold="true" box="[853,867,1910,1932]" pageId="3" pageNumber="396">S</emphasis>
|
||
—sternum;
|
||
<emphasis id="A7A23A4EFFF01108839CF8D0B8B9F821" bold="true" box="[988,1012,1910,1932]" pageId="3" pageNumber="396">sb</emphasis>
|
||
—sacculus bursae;
|
||
<emphasis id="A7A23A4EFFF0110884F9F8D0BF94F821" bold="true" box="[1209,1241,1910,1932]" pageId="3" pageNumber="396">ser</emphasis>
|
||
—serratulum;
|
||
<emphasis id="A7A23A4EFFF01108852CF8D0BECFF821" bold="true" box="[1388,1410,1910,1932]" pageId="3" pageNumber="396">sg</emphasis>
|
||
— subgenitale;
|
||
<emphasis id="A7A23A4EFFF011088157F833BA64F806" bold="true" box="[279,297,1941,1963]" pageId="3" pageNumber="396">st</emphasis>
|
||
—gonostylus 9;
|
||
<emphasis id="A7A23A4EFFF011088191F833BAAFF806" bold="true" box="[465,482,1941,1963]" pageId="3" pageNumber="396">T</emphasis>
|
||
—tergum. Arrow indicates ventral projection of male gonocoxite 9. Scale bar = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9569E65CFFF7110F8087FF3EB979FEED" blockId="4.[151,1436,151,1040]" pageId="4" pageNumber="397">
|
||
Sternum 7 (
|
||
<figureCitation id="0DEDFAD9FFF7110F8109FF31BA85FF1D" box="[329,456,151,176]" captionStart="FIGURES 3 – 12" captionStartId="3.[151,264,1726,1748]" captionTargetBox="[152,1428,198,1704]" captionTargetId="figure@3.[152,1435,193,1705]" captionTargetPageId="3" captionText="FIGURES 3 – 12. Inocellia indica sp. nov. 3 — male genital segments, lateral view; 4 — male genital segments, dorsal view; 5 — male genital segments, ventral view; 6 — male genital segments, caudal view; 7 — male ninth gonocoxite with internal structures, lateral view; 8 — gonarcus, dorsal view; 9 — fused parameres, dorsal view; 10 — female genital segments, lateral view; 11 — female sternite 7 and subgenitale, ventral view; 12 — bursa copulatrix, lateral view. Abbreviations: ab — atrium bursae; e — ectoproct; ep — endophallus; g — gonarcus (= fused gonocoxites 11); gr — glandula receptaculi; gx 9 — gonocoxite 9; hi — hypandrium internum; o — ovipositor; p — fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10); ps — pseudostyli (= gonapophyses 9); r — receptaculum seminis; S — sternum; sb — sacculus bursae; ser — serratulum; sg — subgenitale; st — gonostylus 9; T — tergum. Arrow indicates ventral projection of male gonocoxite 9. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/237706/files/figure.png" pageId="4" pageNumber="397">Figs 10–11</figureCitation>
|
||
) nearly trapezoidal in lateral view, posterolateral margins slightly emarginate, posterior margin slightly concave. Subgenital plate (fused gonocoxites 8) (
|
||
<figureCitation id="0DEDFAD9FFF7110F8334FF1AB8BEFF79" box="[884,1011,188,213]" captionStart="FIGURES 3 – 12" captionStartId="3.[151,264,1726,1748]" captionTargetBox="[152,1428,198,1704]" captionTargetId="figure@3.[152,1435,193,1705]" captionTargetPageId="3" captionText="FIGURES 3 – 12. Inocellia indica sp. nov. 3 — male genital segments, lateral view; 4 — male genital segments, dorsal view; 5 — male genital segments, ventral view; 6 — male genital segments, caudal view; 7 — male ninth gonocoxite with internal structures, lateral view; 8 — gonarcus, dorsal view; 9 — fused parameres, dorsal view; 10 — female genital segments, lateral view; 11 — female sternite 7 and subgenitale, ventral view; 12 — bursa copulatrix, lateral view. Abbreviations: ab — atrium bursae; e — ectoproct; ep — endophallus; g — gonarcus (= fused gonocoxites 11); gr — glandula receptaculi; gx 9 — gonocoxite 9; hi — hypandrium internum; o — ovipositor; p — fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10); ps — pseudostyli (= gonapophyses 9); r — receptaculum seminis; S — sternum; sb — sacculus bursae; ser — serratulum; sg — subgenitale; st — gonostylus 9; T — tergum. Arrow indicates ventral projection of male gonocoxite 9. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/237706/files/figure.png" pageId="4" pageNumber="397">Figs 10–11</figureCitation>
|
||
) mostly membranous, anteriorly with a pair of longitudinal sclerotized areas, posteriorly with a narrow, transverse sclerite. Atrium bursae (
|
||
<figureCitation id="0DEDFAD9FFF7110F857EFF79BED9FF55" box="[1342,1428,223,248]" captionStart="FIGURES 3 – 12" captionStartId="3.[151,264,1726,1748]" captionTargetBox="[152,1428,198,1704]" captionTargetId="figure@3.[152,1435,193,1705]" captionTargetPageId="3" captionText="FIGURES 3 – 12. Inocellia indica sp. nov. 3 — male genital segments, lateral view; 4 — male genital segments, dorsal view; 5 — male genital segments, ventral view; 6 — male genital segments, caudal view; 7 — male ninth gonocoxite with internal structures, lateral view; 8 — gonarcus, dorsal view; 9 — fused parameres, dorsal view; 10 — female genital segments, lateral view; 11 — female sternite 7 and subgenitale, ventral view; 12 — bursa copulatrix, lateral view. Abbreviations: ab — atrium bursae; e — ectoproct; ep — endophallus; g — gonarcus (= fused gonocoxites 11); gr — glandula receptaculi; gx 9 — gonocoxite 9; hi — hypandrium internum; o — ovipositor; p — fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10); ps — pseudostyli (= gonapophyses 9); r — receptaculum seminis; S — sternum; sb — sacculus bursae; ser — serratulum; sg — subgenitale; st — gonostylus 9; T — tergum. Arrow indicates ventral projection of male gonocoxite 9. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/237706/files/figure.png" pageId="4" pageNumber="397">Fig. 12</figureCitation>
|
||
) subtriangular, membranous, laterally rugose; sacculus bursae sac-like; receptaculum seminis short, distally with a pair of tubular glandulae receptaculi.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DDCCB5D7FFF7110F8087FEEABAFDFD28" pageId="4" pageNumber="397" type="materials_examined">
|
||
<paragraph id="9569E65CFFF7110F8087FEEABAFDFD28" blockId="4.[151,1436,151,1040]" pageId="4" pageNumber="397">
|
||
<emphasis id="A7A23A4EFFF7110F8087FEEABAD8FEC8" bold="true" box="[199,405,332,357]" pageId="4" pageNumber="397">
|
||
<typeStatus id="4A6D58FEFFF7110F8087FEEABA4FFEC8" box="[199,258,332,357]" pageId="4" pageNumber="397">Type</typeStatus>
|
||
specimens.
|
||
</emphasis>
|
||
<typeStatus id="4A6D58FEFFF7110F81E8FEEAB95DFEC8" box="[424,528,332,357]" pageId="4" pageNumber="397" type="holotype">Holotype</typeStatus>
|
||
♂,
|
||
<collectingCountry id="EDC1A6CCFFF7110F8212FEEAB9EBFEC9" box="[594,678,332,356]" name="India" pageId="4" pageNumber="397">INDIA</collectingCountry>
|
||
: Meghalaya, East Khasi Hills, Shillong, Umshing,
|
||
<quantity id="522E4BB9FFF7110F8574FEEABEDAFEC9" box="[1332,1431,332,356]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="4" pageNumber="397" unit="m" value="1420.0">1420 m</quantity>
|
||
, 25°36′43.55″N, 91°53′58.86″E,
|
||
<date id="E168C09CFFF7110F824DFEC9B9CEFE25" box="[525,643,367,392]" pageId="4" pageNumber="397" value="2015-05-28">28.V.2015</date>
|
||
, S.R. Hajong and E.S. Lyngchiang (
|
||
<collectionCode id="F3C77E99FFF7110F8400FED6BFC2FE25" box="[1088,1167,368,392]" pageId="4" pageNumber="397">NCBS</collectionCode>
|
||
).
|
||
<typeStatus id="4A6D58FEFFF7110F84E8FED6BE54FE25" box="[1192,1305,368,392]" pageId="4" pageNumber="397" type="paratype">Paratypes</typeStatus>
|
||
: 1♀, same collecting site and collectors as
|
||
<typeStatus id="4A6D58FEFFF7110F81B8FE32B910FE00" box="[504,605,404,429]" pageId="4" pageNumber="397" type="holotype">holotype</typeStatus>
|
||
, but collected on
|
||
<date id="E168C09CFFF7110F8360FE32B8DEFE00" box="[800,915,404,429]" pageId="4" pageNumber="397" value="2015-05-27">27.V.2015</date>
|
||
(
|
||
<collectionCode id="F3C77E99FFF7110F83E1FE32B8BDFE01" box="[929,1008,404,428]" pageId="4" pageNumber="397">NCBS</collectionCode>
|
||
);
|
||
<specimenCount id="83D02DD5FFF7110F8442FE32BF6AFE01" box="[1026,1063,404,428]" pageId="4" pageNumber="397" type="generic">1♂</specimenCount>
|
||
, same collecting data as
|
||
<typeStatus id="4A6D58FEFFF7110F857AFE32BED1FE00" box="[1338,1436,404,429]" pageId="4" pageNumber="397" type="holotype">holotype</typeStatus>
|
||
(collection S.R. Hajong);
|
||
<specimenCount id="83D02DD5FFF7110F81F6FE1EBA90FE7D" box="[438,477,440,464]" pageId="4" pageNumber="397" type="generic">2♂</specimenCount>
|
||
2♀, same collecting site and collectors as
|
||
<typeStatus id="4A6D58FEFFF7110F83ECFE11BF5CFE7D" box="[940,1041,439,464]" pageId="4" pageNumber="397" type="holotype">holotype</typeStatus>
|
||
, but collected on
|
||
<date id="E168C09CFFF7110F8494FE11BE0BFE7D" box="[1236,1350,439,464]" pageId="4" pageNumber="397" value="2015-05-29">29.V.2015</date>
|
||
(
|
||
<specimenCount id="83D02DD5FFF7110F8518FE1EBE33FE7D" box="[1368,1406,440,464]" pageId="4" pageNumber="397" type="generic">1♂</specimenCount>
|
||
1♀ in NHM;
|
||
<specimenCount id="83D02DD5FFF7110F8150FE7ABA7EFE59" box="[272,307,476,500]" pageId="4" pageNumber="397" type="generic">1♂</specimenCount>
|
||
1♀ in
|
||
<collectionCode id="F3C77E99FFF7110F81C6FE7ABA86FE59" box="[390,459,476,500]" pageId="4" pageNumber="397">NFIC</collectionCode>
|
||
); 1♀, same collecting site and collectors as
|
||
<typeStatus id="4A6D58FEFFF7110F83A7FE7ABF01FE59" box="[999,1100,476,500]" pageId="4" pageNumber="397" type="holotype">holotype</typeStatus>
|
||
, but collected on
|
||
<date id="E168C09CFFF7110F856AFE7ABED1FE59" box="[1322,1436,476,500]" pageId="4" pageNumber="397" value="2015-05-30">30.V.2015</date>
|
||
(collection S.R. Hajong); 2♀, same collecting site and collectors as
|
||
<typeStatus id="4A6D58FEFFF7110F83A0FDA6BF08FDB5" box="[992,1093,512,536]" pageId="4" pageNumber="397" type="holotype">holotype</typeStatus>
|
||
, but collected on
|
||
<date id="E168C09CFFF7110F856BFDA6BED1FDB5" box="[1323,1436,512,536]" pageId="4" pageNumber="397" value="2015-06-01">1.VI.2015</date>
|
||
(collection S.R. Hajong); 1♀, same collecting site as
|
||
<typeStatus id="4A6D58FEFFF7110F8346FD82B826FD91" box="[774,875,548,572]" pageId="4" pageNumber="397" type="holotype">holotype</typeStatus>
|
||
, 25°36′43.90″N, 91°54′14.3″E,
|
||
<date id="E168C09CFFF7110F849BFD82BE10FD91" box="[1243,1373,548,572]" pageId="4" pageNumber="397" value="2012-06-18">18.VI.2012</date>
|
||
, S.R. Hajong (collection S.R. Hajong); 2♀, same collecting site as
|
||
<typeStatus id="4A6D58FEFFF7110F831AFDEEB8F2FDCD" box="[858,959,584,608]" pageId="4" pageNumber="397" type="holotype">holotype</typeStatus>
|
||
, but collected on
|
||
<date id="E168C09CFFF7110F84CDFDEEBE4EFDCD" box="[1165,1283,584,608]" pageId="4" pageNumber="397" value="2014-05-28">28.V.2014</date>
|
||
, S.R. Hajong (collection S.R. Hajong).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DDCCB5D7FFF7110F8087FD29B940FD60" pageId="4" pageNumber="397" type="etymology">
|
||
<paragraph id="9569E65CFFF7110F8087FD29B940FD60" blockId="4.[151,1436,151,1040]" pageId="4" pageNumber="397">
|
||
<emphasis id="A7A23A4EFFF7110F8087FD29BA03FD05" bold="true" box="[199,334,655,680]" pageId="4" pageNumber="397">Etymology.</emphasis>
|
||
The specific epithet “
|
||
<taxonomicName id="52D69DDFFFF7110F820BFD36B9DCFD05" box="[587,657,656,680]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="4" pageNumber="397" phylum="Arthropoda" rank="species" species="indica">
|
||
<emphasis id="A7A23A4EFFF7110F820BFD36B9DCFD05" box="[587,657,656,680]" italics="true" pageId="4" pageNumber="397">indica</emphasis>
|
||
</taxonomicName>
|
||
” refers to the geographical distribution of this new species which is currently known only from
|
||
<collectingCountry id="EDC1A6CCFFF7110F818CFD12B944FD60" box="[460,521,692,717]" name="India" pageId="4" pageNumber="397">India</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DDCCB5D7FFF7110F8087FD71BE7CFBBD" pageId="4" pageNumber="397" type="distribution">
|
||
<paragraph id="9569E65CFFF7110F8087FD71BA9FFCB9" blockId="4.[151,1436,151,1040]" pageId="4" pageNumber="397">
|
||
<emphasis id="A7A23A4EFFF7110F8087FD71BA2DFD5D" bold="true" box="[199,352,727,752]" pageId="4" pageNumber="397">Distribution.</emphasis>
|
||
This new species is currently known only from Shillong of East Khasi Hills in Meghalaya, northeastern
|
||
<collectingCountry id="EDC1A6CCFFF7110F8168FD5ABA2CFCB8" box="[296,353,764,789]" name="India" pageId="4" pageNumber="397">India</collectingCountry>
|
||
(
|
||
<figureCitation id="0DEDFAD9FFF7110F8131FD5ABA89FCB9" box="[369,452,764,789]" captionStart="FIGURE 14" captionStartId="5.[151,250,1681,1703]" captionTargetBox="[226,1360,193,1659]" captionTargetId="figure@5.[226,1360,193,1660]" captionTargetPageId="5" captionText="FIGURE 14. Geographic distribution of inocelliid species from South Asia. ● — Inocellia fulvostigmata; ■ — Inocellia bhutana; ★ — Inocellia indica sp. nov. [Records of I. fulvostigmata and I. bhutana follow H. Aspöck et al. (1991)]" httpUri="https://zenodo.org/record/237708/files/figure.png" pageId="4" pageNumber="397">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="9569E65CFFF7110F8087FCB9BE7CFBBD" blockId="4.[151,1436,151,1040]" pageId="4" pageNumber="397">
|
||
<emphasis id="A7A23A4EFFF7110F8087FCB9BADDFC95" bold="true" box="[199,400,799,824]" pageId="4" pageNumber="397">Ecological notes.</emphasis>
|
||
Collection of the new species was done by using a light trap, which consisted of a 125W HPMV Mercury vapor lamp hung against a white sheet of cloth. The trap was run from 6 pm in the evening till 10 pm. Not more than 3–4 individuals could be collected during this period. The collecting site (
|
||
<figureCitation id="0DEDFAD9FFF7110F848BFCC1BE50FC2D" box="[1227,1309,871,896]" captionStart="FIGURE 13" captionStartId="4.[151,250,1990,2012]" captionTargetBox="[210,1377,1093,1968]" captionTargetId="figure@4.[210,1377,1093,1969]" captionTargetPageId="4" captionText="FIGURE 13. Habitat of Inocellia indica sp. nov." httpUri="https://zenodo.org/record/237707/files/figure.png" pageId="4" pageNumber="397">Fig. 13</figureCitation>
|
||
) is located adjacent to a forest patch consisting of tall Khasi pine trees (
|
||
<taxonomicName id="52D69DDFFFF7110F837FFC2BBFDAFC08" authority="Royle ex Gordon" authorityName="Royle ex Gordon" box="[831,1175,908,933]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="397" phylum="Tracheophyta" rank="species" species="kesiya">
|
||
<emphasis id="A7A23A4EFFF7110F837FFC2BB881FC09" box="[831,972,908,932]" italics="true" pageId="4" pageNumber="397">Pinus kesiya</emphasis>
|
||
Royle ex Gordon
|
||
</taxonomicName>
|
||
) which were of around more than 30 meters tall. The new snakeflies appear to inhabit the tree canopy because attempts to collect them using sweeping net from lower branches of small pine trees were unsuccessful. Nothing is known about the biology of the new species and no immatures could be collected from the tree bark where they purportedly dwell.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="C1A9B6D4FFF7110F80D7F860B9C5F871" box="[151,648,1990,2012]" httpUri="https://zenodo.org/record/237707/files/figure.png" pageId="4" pageNumber="397" targetBox="[210,1377,1093,1968]" targetPageId="4">
|
||
<paragraph id="9569E65CFFF7110F80D7F860B9C5F871" blockId="4.[151,648,1990,2012]" box="[151,648,1990,2012]" pageId="4" pageNumber="397">
|
||
<emphasis id="A7A23A4EFFF7110F80D7F860BA54F871" bold="true" box="[151,281,1990,2012]" pageId="4" pageNumber="397">FIGURE 13</emphasis>
|
||
. Habitat of
|
||
<taxonomicName id="52D69DDFFFF7110F81D2F860B97CF871" box="[402,561,1990,2012]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="4" pageNumber="397" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
|
||
<emphasis id="A7A23A4EFFF7110F81D2F860B97CF871" box="[402,561,1990,2012]" italics="true" pageId="4" pageNumber="397">Inocellia indica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BC918735FFF7110F8278F860B9C5F871" box="[568,648,1990,2012]" pageId="4" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C1A9B6D4FFF6110E80D7F937BE69F966" httpUri="https://zenodo.org/record/237708/files/figure.png" pageId="5" pageNumber="398" targetBox="[226,1360,193,1659]" targetPageId="5">
|
||
<paragraph id="9569E65CFFF6110E80D7F937BE69F966" blockId="5.[151,1436,1681,1739]" pageId="5" pageNumber="398">
|
||
<emphasis id="A7A23A4EFFF6110E80D7F937BA6DF90A" bold="true" box="[151,288,1681,1703]" pageId="5" pageNumber="398">FIGURE 14</emphasis>
|
||
. Geographic distribution of inocelliid species from South Asia. ●—
|
||
<taxonomicName id="52D69DDFFFF6110E8453F937BE48F90A" box="[1043,1285,1681,1703]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="5" pageNumber="398" phylum="Arthropoda" rank="species" species="fulvostigmata">
|
||
<emphasis id="A7A23A4EFFF6110E8453F937BE48F90A" box="[1043,1285,1681,1703]" italics="true" pageId="5" pageNumber="398">Inocellia fulvostigmata</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="A7A23A4EFFF6110E8559F934BE0CF90A" bold="true" box="[1305,1345,1682,1703]" pageId="5" pageNumber="398">■—</emphasis>
|
||
<taxonomicName id="52D69DDFFFF6110E8502F937BBA7F966" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="5" pageNumber="398" phylum="Arthropoda" rank="species" species="bhutana">
|
||
<emphasis id="A7A23A4EFFF6110E8502F937BBA7F966" italics="true" pageId="5" pageNumber="398">Inocellia bhutana</emphasis>
|
||
</taxonomicName>
|
||
; ★—
|
||
<taxonomicName id="52D69DDFFFF6110E8167F913BA8BF966" box="[295,454,1717,1739]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="5" pageNumber="398" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
|
||
<emphasis id="A7A23A4EFFF6110E8167F913BA8BF966" box="[295,454,1717,1739]" italics="true" pageId="5" pageNumber="398">Inocellia indica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BC918735FFF6110E818DF913B950F966" box="[461,541,1717,1739]" pageId="5" pageNumber="398" rank="species">
|
||
<emphasis id="A7A23A4EFFF6110E818DF913B950F966" bold="true" box="[461,541,1717,1739]" pageId="5" pageNumber="398">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
[Records of
|
||
<taxonomicName id="52D69DDFFFF6110E82EAF910B807F966" box="[682,842,1717,1739]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="5" pageNumber="398" phylum="Arthropoda" rank="species" species="fulvostigmata">I. fulvostigmata</taxonomicName>
|
||
and
|
||
<taxonomicName id="52D69DDFFFF6110E833BF910B8ADF966" box="[891,992,1717,1739]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="5" pageNumber="398" phylum="Arthropoda" rank="species" species="bhutana">I. bhutana</taxonomicName>
|
||
follow H. Aspöck et al. (1991)]
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="DDCCB5D7FFF6110D8087F95AB9F7FE25" lastPageId="6" lastPageNumber="399" pageId="5" pageNumber="398" type="discussion">
|
||
<paragraph id="9569E65CFFF6110D8087F95AB9F7FE25" blockId="5.[151,1437,1788,2029]" lastBlockId="6.[151,1436,151,392]" lastPageId="6" lastPageNumber="399" pageId="5" pageNumber="398">
|
||
<emphasis id="A7A23A4EFFF6110E8087F95ABA64F8B9" box="[199,297,1788,1812]" italics="true" pageId="5" pageNumber="398">Remarks</emphasis>
|
||
. The new species belongs to the genus
|
||
<taxonomicName id="52D69DDFFFF6110E82A4F95AB805F8B9" box="[740,840,1788,1812]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="5" pageNumber="398" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A7A23A4EFFF6110E82A4F95AB805F8B9" box="[740,840,1788,1812]" italics="true" pageId="5" pageNumber="398">Inocellia</emphasis>
|
||
</taxonomicName>
|
||
as evidenced the absence of the MA base in the hind wing and the presence of the foliate pseudostyli.
|
||
<bibRefCitation id="F1479BADFFF6110E8287F886B820F895" author="Hajong" box="[711,877,1824,1849]" pageId="5" pageNumber="398" refString="Hajong, S. R. (2013) Record of a rare snake fly (Raphidioptera: Inocelliidae) from Meghalaya. NeBIO, 4, 50 - 51." type="journal article" year="2013">Hajong (2013)</bibRefCitation>
|
||
mentioned the possibility that the new species is conspecific with
|
||
<taxonomicName id="52D69DDFFFF6110E8118F8E2B89AF8F0" authority="U. Aspock and H. Aspock, 1968" authorityName="U. Aspock and H. Aspock" authorityYear="1968" box="[344,983,1860,1885]" class="Insecta" family="Inocelliidae" genus="Parainocellia" kingdom="Animalia" order="Raphidioptera" pageId="5" pageNumber="398" phylum="Arthropoda" rank="species" species="burmana">
|
||
<emphasis id="A7A23A4EFFF6110E8118F8E2B912F8F1" box="[344,607,1860,1884]" italics="true" pageId="5" pageNumber="398">Parainocellia burmana</emphasis>
|
||
U. Aspöck and H. Aspöck, 1968
|
||
</taxonomicName>
|
||
from northern
|
||
<collectingCountry id="EDC1A6CCFFF6110E84C3F8E3BFBFF8F0" box="[1155,1266,1861,1885]" name="Myanmar" pageId="5" pageNumber="398">Myanmar</collectingCountry>
|
||
. However, it is now clear that they are distantly related because of the male genitalic structures, especially the pseudostyli which are claw-like in
|
||
<taxonomicName id="52D69DDFFFF6110E8115F82BB9B5F808" authority="H. Aspock et al. 1991" authorityName="H. Aspock et al." authorityYear="1991" box="[341,760,1932,1957]" class="Insecta" family="Inocelliidae" genus="Parainocellia" kingdom="Animalia" order="Raphidioptera" pageId="5" pageNumber="398" phylum="Arthropoda" rank="species" species="burmana">
|
||
<emphasis id="A7A23A4EFFF6110E8115F82BBA94F809" box="[341,473,1932,1956]" italics="true" pageId="5" pageNumber="398">P. burmana</emphasis>
|
||
(H.
|
||
<bibRefCitation id="F1479BADFFF6110E8252F82AB9BDF808" author="Aspock" box="[530,752,1932,1957]" pageId="5" pageNumber="398" refString="Aspock, H., Aspock, U. & Rausch, H. (1991) Die Raphidiopteren der Erde. Eine monographische Darstellung der Systematik, Taxonomie, Biologie, Okologie und Chorologie der rezenten Raphidiopteren der Erde, mit einer zusammenfassenden Ubersicht der fossilen Raphidiopteren (Insecta: Neuropteroidea). Vols. 2. Goecke & Evers, Krefeld, 730 pp. & 550 pp." type="book" year="1991">
|
||
Aspöck
|
||
<emphasis id="A7A23A4EFFF6110E8232F82BB9E0F809" box="[626,685,1932,1956]" italics="true" pageId="5" pageNumber="398">et al.</emphasis>
|
||
1991
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
but foliate in the new species as in the other members of
|
||
<taxonomicName id="52D69DDFFFF6110E80D7F816BBB6F865" box="[151,251,1968,1992]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="5" pageNumber="398" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A7A23A4EFFF6110E80D7F816BBB6F865" box="[151,251,1968,1992]" italics="true" pageId="5" pageNumber="398">Inocellia</emphasis>
|
||
</taxonomicName>
|
||
. The new species should be placed within the
|
||
<taxonomicName id="52D69DDFFFF6110E834BF817B8F1F865" box="[779,956,1968,1992]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="5" pageNumber="398" phylum="Arthropoda" rank="species" species="fulvostigmata">
|
||
<emphasis id="A7A23A4EFFF6110E834BF817B8F1F865" box="[779,956,1968,1992]" italics="true" pageId="5" pageNumber="398">I. fulvostigmata</emphasis>
|
||
</taxonomicName>
|
||
species group due to the form of the male gonocoxite 9, which is wider than long, and the presence of a scabrous membranous serratulum [see identification key to species of
|
||
<taxonomicName id="52D69DDFFFF5110D8117FF3EBAF6FF1D" box="[343,443,152,176]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="6" pageNumber="399" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A7A23A4EFFF5110D8117FF3EBAF6FF1D" box="[343,443,152,176]" italics="true" pageId="6" pageNumber="399">Inocellia</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="F1479BADFFF5110D819DFF31B9EBFF1D" box="[477,678,151,176]" pageId="6" pageNumber="399" refString="Liu, X. Y., Aspock, H., Zhang, W. W. & Aspock, U. (2012 b) New species of the snakefly genus Inocellia Schneider, 1843 (Raphidioptera: Inocelliidae) from Yunnan, China. Zootaxa, 3298, 43 - 52." type="journal article">
|
||
Liu
|
||
<emphasis id="A7A23A4EFFF5110D824AFF3FB90FFF1D" box="[522,578,152,176]" italics="true" pageId="6" pageNumber="399">et al.</emphasis>
|
||
(2012b)
|
||
</bibRefCitation>
|
||
]. The new species appears to be closely related to
|
||
<taxonomicName id="52D69DDFFFF5110D8493FF3FBE0BFF1D" box="[1235,1350,152,176]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="6" pageNumber="399" phylum="Arthropoda" rank="species" species="bhutana">
|
||
<emphasis id="A7A23A4EFFF5110D8493FF3FBE0BFF1D" box="[1235,1350,152,176]" italics="true" pageId="6" pageNumber="399">I. bhutana</emphasis>
|
||
</taxonomicName>
|
||
as both have a similar male gonocoxite 9 ventrally with a subtriangular projection, and by having a generally similar gonarcus, which is prominent at its ventrolateral corners and posteriorly produced medially. However,
|
||
<taxonomicName id="52D69DDFFFF5110D8556FF47BE3EFF55" box="[1302,1395,224,248]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="6" pageNumber="399" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
|
||
<emphasis id="A7A23A4EFFF5110D8556FF47BE3EFF55" box="[1302,1395,224,248]" italics="true" pageId="6" pageNumber="399">I. indica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BC918735FFF5110D853AFF79BB85FEB0" pageId="6" pageNumber="399" rank="species">
|
||
<emphasis id="A7A23A4EFFF5110D853AFF79BB85FEB0" bold="true" pageId="6" pageNumber="399">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be distinguished from
|
||
<taxonomicName id="52D69DDFFFF5110D81B5FEA3B925FEB1" box="[501,616,260,284]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="6" pageNumber="399" phylum="Arthropoda" rank="species" species="bhutana">
|
||
<emphasis id="A7A23A4EFFF5110D81B5FEA3B925FEB1" box="[501,616,260,284]" italics="true" pageId="6" pageNumber="399">I. bhutana</emphasis>
|
||
</taxonomicName>
|
||
by its male gonocoxite 9 possessing a broad, flat stylus, the presence of a bristle tuft on the anterodorsal portion of the inner surface of gonocoxite 9, and the presence of a pair of bristle tufts respectively on the endophallus and serratulum. In
|
||
<taxonomicName id="52D69DDFFFF5110D82A1FEEBB81BFEC9" box="[737,854,332,356]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="6" pageNumber="399" phylum="Arthropoda" rank="species" species="bhutana">
|
||
<emphasis id="A7A23A4EFFF5110D82A1FEEBB81BFEC9" box="[737,854,332,356]" italics="true" pageId="6" pageNumber="399">I. bhutana</emphasis>
|
||
</taxonomicName>
|
||
, the stylus of gonocoxite 9 is slenderly digitiform, and the endophallus and serratulum lack bristles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |