432 lines
41 KiB
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432 lines
41 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.366.6172" ID-PMC="PMC3890666" ID-Pensoft-Pub="1313-2970-366-1" ID-Pensoft-UUID="FFD8DC462108382BCB68FFC9FF97F235" ID-PubMed="24453569" ID-Zenodo-Dep="577560" ModsDocID="1313-2970-366-1" checkinTime="1451246487361" checkinUser="pensoft" docAuthor="Kolibac, Jiri" docDate="2013" docId="E5D1FD5D7B890215315C4190D710C58D" docLanguage="en" docName="ZooKeys 366: 1-194" docOrigin="ZooKeys 366" docPubDate="2013-12-31" docSource="http://dx.doi.org/10.3897/zookeys.366.6172" docTitle="Thymalus Latreille 1802" docType="treatment" docVersion="5" id="FFD8DC462108382BCB68FFC9FF97F235" lastPageNumber="101" masterDocId="FFD8DC462108382BCB68FFC9FF97F235" masterDocTitle="Trogossitidae: A review of the beetle family, with a catalogue and keys" masterLastPageNumber="194" masterPageNumber="1" pageNumber="99" updateTime="1668157348086" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Trogossitidae: A review of the beetle family, with a catalogue and keys</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kolibac, Jiri</mods:namePart>
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<mods:affiliation>Moravian Museum, Department of Entomology, Hviezdoslavova 29 a, 627 00 Brno, Czech Republic</mods:affiliation>
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<mods:date>2013</mods:date>
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<mods:number>2013-12-31</mods:number>
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<mods:detail type="volume">
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<mods:number>366</mods:number>
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<mods:start>1</mods:start>
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<mods:end>194</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.366.6172</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.366.6172</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-366-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFD8DC462108382BCB68FFC9FF97F235</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577560</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152050242" LSID="urn:lsid:plazi:treatment:E5D1FD5D7B890215315C4190D710C58D" httpUri="http://treatment.plazi.org/id/E5D1FD5D7B890215315C4190D710C58D" lastPageId="100" lastPageNumber="101" pageId="98" pageNumber="99">
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<subSubSection pageId="98" pageNumber="99" type="nomenclature">
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<paragraph pageId="98" pageNumber="99">
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<pageBreakToken pageId="98" pageNumber="99" start="start">Genus</pageBreakToken>
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<taxonomicName LSID="E5D1FD5D-7B89-0215-315C-4190D710C58D" authority="Latreille, 1802" authorityName="Latreille" authorityYear="1802" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thymalus" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">Thymalus Latreille, 1802</taxonomicName>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A Thymalus limbatus B Phloiophilus edwardsi C Eronyxa marginicollis D Decamerus haemorhoidalis E Diontolobus punctatipennis F Afrocyrona ciskeiensis G Afrocyrona dwesae H Grynoma sp., New Zealand I Grynoma diluta." httpUri="https://binary.pensoft.net/fig/21100" pageId="98" pageNumber="99">Figs 10</figureCitation>
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. A-F Colydiopeltis compactum (A ventral surface B procoxal area C head ventrally D detail of mouthparts E protarsus F apex of metatibia with row of spines) G-H Thymalus limbatus (G antenna H apex of protibia with straight spur)." httpUri="https://binary.pensoft.net/fig/21111" pageId="98" pageNumber="99">, 16</figureCitation>
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. A Thymalus limbatus, elytral apex with interlocking mechanism B-E Decamerus haemorhoidalis (B prothorax with partially closed procoxal cavities C metatarsal claws with denticle D protarsus with projecting empodium E irregular punctation of elytra) F Grynocharis oblonga, elytral sculpture G-H Lophocateres pusillus (G head in frontal view with deeply emarginate frontoclypeal suture H detail)." httpUri="https://binary.pensoft.net/fig/21113" pageId="98" pageNumber="99">, 18</figureCitation>
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<figureCitation captionStart="Map 10" captionStartId="F10m" captionText="Map 10. A distribution of the tribe Thymalini including Rentonium - group." httpUri="https://binary.pensoft.net/fig/21102" pageId="98" pageNumber="99">; Map 10</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="98" pageNumber="99" type="reference_group">
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<paragraph pageId="98" pageNumber="99">
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<taxonomicName authorityName="Latreille" authorityYear="1802" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">Thymalus</taxonomicName>
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Latreille, 1802: 133.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="98" pageNumber="99" type="type species">
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<paragraph pageId="98" pageNumber="99">Type species.</paragraph>
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<paragraph pageId="98" pageNumber="99">
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<taxonomicName class="Hexapoda" family="Trogossitidae" genus="Peltis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="brunnea">
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<emphasis italics="true" pageId="98" pageNumber="99">Peltis brunnea</emphasis>
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</taxonomicName>
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Thunberg, 1794 (=
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<taxonomicName authorityName="Fabricius" authorityYear="1787" class="Hexapoda" family="Chrysomelidae" genus="Cassida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="limbata">
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<emphasis italics="true" pageId="98" pageNumber="99">Cassida limbata</emphasis>
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</taxonomicName>
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Fabricius, 1787) [by original designation and monotypy]
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</paragraph>
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<paragraph pageId="98" pageNumber="99">
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<normalizedToken originalValue="Léveillé">Leveille</normalizedToken>
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, A. 1910: 32. Barron, J. R. 1971: 35. Crowson, R. A. 1964a: 296.
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<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
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, J. 2005: 85 (redescription).
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<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
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, J. 2006: 111 (phylogeny).
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<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
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, J. 2007a: 366. Nikitsky, N. B. et al. 1998: 29 (key). Lafer, G. Sh. 1992: 83. Reitter, E. 1876: 64
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</paragraph>
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<paragraph pageId="98" pageNumber="99">
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<taxonomicName class="Insecta" family="Trogositidae" genus="Thymalops" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="98" pageNumber="99">Thymalops</emphasis>
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</taxonomicName>
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Iablokoff-Khnzorian, 1962 [Type species:
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<taxonomicName authorityName="Fabricius" authorityYear="1787" class="Hexapoda" family="Chrysomelidae" genus="Cassida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="limbata">
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<emphasis italics="true" pageId="98" pageNumber="99">Cassida limbata</emphasis>
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</taxonomicName>
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Fabricius, 1787]
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</paragraph>
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<paragraph pageId="98" pageNumber="99">Barron, J. R. 1971: 35. Iablokoff-Khnzorian, S. M. 1962: 421</paragraph>
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</subSubSection>
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<subSubSection pageId="98" pageNumber="99" type="remarks">
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<paragraph pageId="98" pageNumber="99">Remarks.</paragraph>
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<paragraph pageId="98" pageNumber="99">
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Comparing the larvae as well as adults of
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<taxonomicName authorityName="Latreille" authorityYear="1802" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="98" pageNumber="99">Thymalus</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Crowson" authorityYear="1964" class="Insecta" family="Trogossitidae" genus="Protopeltis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="98" pageNumber="99">Protopeltis</emphasis>
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</taxonomicName>
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, I found some interesting similarities, which led me to consideration of their phylogenetic relationship. Later character analysis (
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<bibRefCitation author="Kolibac, J" journalOrPublisher="Entomologica Basiliensia et Collectionis Frey" pageId="155" pageNumber="156" pagination="105 - 153" refId="B146" refString="Kolibac, J, 2006. A review of the Trogossitidae. Part 2: Larval morphology, phylogeny and taxonomy (Coleoptera, Cleroidea). Entomologica Basiliensia et Collectionis Frey 28: 105 - 153" title="A review of the Trogossitidae. Part 2: Larval morphology, phylogeny and taxonomy (Coleoptera, Cleroidea)." volume="28" year="2006">
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<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
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2006
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</bibRefCitation>
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) showed a relationship of
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<taxonomicName authorityName="Latreille" authorityYear="1802" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="98" pageNumber="99">Thymalus</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Crowson" authorityYear="1964" class="Insecta" family="Trogossitidae" genus="Protopeltis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="98" pageNumber="99">Protopeltis</emphasis>
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</taxonomicName>
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with the former
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<taxonomicName lsidName="" pageId="98" pageNumber="99" rank="tribe" tribe="Rentoniini">Rentoniini</taxonomicName>
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Crowson, 1966. This in turn led to the establishment of the tribe
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<taxonomicName authorityName="Leveille" authorityYear="1888" lsidName="" pageId="98" pageNumber="99" rank="tribe" tribe="Thymalini">Thymalini</taxonomicName>
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for the group. However,
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<bibRefCitation author="Crowson, RA" journalOrPublisher="Proceedings of the Royal entomological Society of London" pageId="149" pageNumber="150" pagination="119 - 127" refId="B46" refString="Crowson, RA, 1966. Further observations on Peltidae (Coleoptera: Cleroidea), with definitions of a new subfamily and of four new genera. Proceedings of the Royal entomological Society of London 35: 119 - 127" title="Further observations on Peltidae (Coleoptera: Cleroidea), with definitions of a new subfamily and of four new genera." volume="35" year="1966">Crowson (1966</bibRefCitation>
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,
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<bibRefCitation author="Crowson, RA" journalOrPublisher="Proceedings of the Royal entomological Society of London (B)" pageId="149" pageNumber="150" pagination="1 - 20" refId="B48" refString="Crowson, RA, 1970. Further observations on Cleroidea (Coleoptera). Proceedings of the Royal entomological Society of London (B) 39: 1 - 20" title="Further observations on Cleroidea (Coleoptera)." volume="39" year="1970">1970</bibRefCitation>
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) also associated the former monotypic tribe
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<taxonomicName authorityName="Crowson" authorityYear="1966" lsidName="" pageId="98" pageNumber="99" rank="tribe" tribe="Protopeltini">Protopeltini</taxonomicName>
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Crowson, 1966 with the rentoniins. Recently, such a classification was called into question by
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<bibRefCitation author="Lawrence, JF" journalOrPublisher="Annales Zoologici" pageId="156" pageNumber="157" pagination="1 - 217" publicationUrl="10.3161/000345411X576725" refId="B176" refString="Lawrence, JF, Ślipinski, A, Seago, SE, Thayer, MK, Newton, AF, Marvaldi, AE, 2011. Phylogeny of the Coleoptera based on morphological characters of adults and larvae. Annales Zoologici 61: 1 - 217, 10.3161/000345411X576725" title="Phylogeny of the Coleoptera based on morphological characters of adults and larvae." url="10.3161/000345411X576725" volume="61" year="2011">Lawrence et al. (2011)</bibRefCitation>
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and
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<bibRefCitation author="Leschen, RAB" journalOrPublisher="Systematic Entomology" pageId="156" pageNumber="157" refId="B185" refString="Leschen, RAB, Lawrence, JF, Ślipinski, AS, Gimmel, M, 2012. Phylogenetic relationships of Cleroidea.XXIVth International Congress of Entomology, August 19-25, 2012, Daegu, Korea." title="Phylogenetic relationships of Cleroidea. XXIVth International Congress of Entomology, August 19 - 25, 2012, Daegu, Korea." year="2012">Leschen et al. (2012)</bibRefCitation>
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, who found
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<taxonomicName authorityName="Latreille" authorityYear="1802" class="Hexapoda" family="Trogossitidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="family">Trogossitidae</taxonomicName>
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polyphyletic in their character analyses; however, both analyses were based on the same character states. Their model genera
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<taxonomicName authorityName="Latreille" authorityYear="1802" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="98" pageNumber="99">Thymalus</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Crowson" authorityYear="1966" class="Insecta" family="Trogossitidae" genus="Rentonellum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="98" pageNumber="99">Rentonellum</emphasis>
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</taxonomicName>
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are classified outside
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<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Corydiidae" genus="Cleroidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Blattodea" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">Cleroidea</taxonomicName>
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in
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<bibRefCitation author="Lawrence, JF" journalOrPublisher="Annales Zoologici" pageId="156" pageNumber="157" pagination="1 - 217" publicationUrl="10.3161/000345411X576725" refId="B176" refString="Lawrence, JF, Ślipinski, A, Seago, SE, Thayer, MK, Newton, AF, Marvaldi, AE, 2011. Phylogeny of the Coleoptera based on morphological characters of adults and larvae. Annales Zoologici 61: 1 - 217, 10.3161/000345411X576725" title="Phylogeny of the Coleoptera based on morphological characters of adults and larvae." url="10.3161/000345411X576725" volume="61" year="2011">Lawrence et al. (2011)</bibRefCitation>
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trees whereas
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<bibRefCitation author="Leschen, RAB" journalOrPublisher="Systematic Entomology" pageId="156" pageNumber="157" refId="B185" refString="Leschen, RAB, Lawrence, JF, Ślipinski, AS, Gimmel, M, 2012. Phylogenetic relationships of Cleroidea.XXIVth International Congress of Entomology, August 19-25, 2012, Daegu, Korea." title="Phylogenetic relationships of Cleroidea. XXIVth International Congress of Entomology, August 19 - 25, 2012, Daegu, Korea." year="2012">Leschen et al. (2012)</bibRefCitation>
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, using a restricted character set, removed them only from
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<taxonomicName authorityName="Latreille" authorityYear="1802" class="Hexapoda" family="Trogossitidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="family">Trogossitidae</taxonomicName>
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and/or suggested subfamily rank for rentoniins again, without necessarily believing in a mutual relationship between the two genera. Some more detail appears in
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<normalizedToken originalValue="“Remarks”">"Remarks"</normalizedToken>
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with the tribe
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<taxonomicName authorityName="Leveille" authorityYear="1888" lsidName="" pageId="98" pageNumber="99" rank="tribe" tribe="Thymalini">Thymalini</taxonomicName>
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.
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</paragraph>
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<paragraph pageId="98" pageNumber="99">
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<bibRefCitation author="Leveille, A" journalOrPublisher="Bulletin de la Societe entomologique de France" pageId="156" pageNumber="157" pagination="cxi - cxii" refId="B186" refString="Leveille, A, 1877. Description d'une nouvelle espece de trogositidae. Bulletin de la Societe entomologique de France 1877: cxi - cxii" title="Description d'une nouvelle espece de trogositidae." volume="1877" year="1877">
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<normalizedToken originalValue="Léveillé">Leveille</normalizedToken>
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(1877)
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</bibRefCitation>
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described the Caucasian species
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<taxonomicName authorityName="Leveille" authorityYear="1877" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="aubei">
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<emphasis italics="true" pageId="98" pageNumber="99">Thymalus aubei</emphasis>
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</taxonomicName>
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as
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<taxonomicName class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="fulgidus">
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<emphasis italics="true" pageId="98" pageNumber="99">Thymalus fulgidus</emphasis>
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</taxonomicName>
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var.
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<emphasis italics="true" pageId="98" pageNumber="99">aubei</emphasis>
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<normalizedToken originalValue="Léveillé">Leveille</normalizedToken>
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, 1877. However,
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<taxonomicName class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="fulgidus">
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<emphasis italics="true" pageId="98" pageNumber="99">Thymalus fulgidus</emphasis>
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</taxonomicName>
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Erichson, 1844 was originally described from North America and
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<bibRefCitation author="Barron, JR" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="147" pageNumber="148" pagination="1 - 143" publicationUrl="10.4039/entm10375fv" refId="B13" refString="Barron, JR, 1971. A revision of the Trogositidae of America North of Mexico (Coleoptera: Cleroidea). Memoirs of the Entomological Society of Canada 75: 1 - 143, 10.4039/entm10375fv" title="A revision of the Trogositidae of America North of Mexico (Coleoptera: Cleroidea)." url="10.4039/entm10375fv" volume="75" year="1971">Barron (1971)</bibRefCitation>
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synonymized this species with
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<taxonomicName authorityName="Chevrolat" authorityYear="1842" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="marginicollis">
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<emphasis italics="true" pageId="98" pageNumber="99">Thymalus marginicollis</emphasis>
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</taxonomicName>
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Chevrolat, 1842. That is perhaps why the latter author also synonymized the taxon
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<emphasis italics="true" pageId="98" pageNumber="99">aubei</emphasis>
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as a synonym of
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<emphasis italics="true" pageId="98" pageNumber="99">marginicollis</emphasis>
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, probably without examination of the holotype or even Caucasian specimens. Russian entomologists, for example
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<bibRefCitation author="Nikitsky, NB" journalOrPublisher="Sbornik Trudov Zoologicheskogo Muzeya MGU" pageId="159" pageNumber="160" pagination="1 - 60" refId="B226" refString="Nikitsky, NB, Semenov, VB, Dolgin, MM, 1998. The beetles of the Prioksko-Terrasny Biosphere Reserve - xylobiontes, mycetobiontes, and Scarabaeidae (with the review of the Moscow Region fauna of these groups). Supplement 1. (With remarks on nomenclature and systematics of some Melandryidae of the World fauna). Sbornik Trudov Zoologicheskogo Muzeya MGU 36: 1 - 60" title="The beetles of the Prioksko-Terrasny Biosphere Reserve - xylobiontes, mycetobiontes, and Scarabaeidae (with the review of the Moscow Region fauna of these groups). Supplement 1. (With remarks on nomenclature and systematics of some Melandryidae of the World fauna)." volume="36" year="1998">Nikitsky et al. (1998)</bibRefCitation>
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, consider
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<taxonomicName authorityName="Leveille" authorityYear="1877" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="aubei">
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<emphasis italics="true" pageId="98" pageNumber="99">Thymalus aubei</emphasis>
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</taxonomicName>
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a valid species, with the synonym
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<taxonomicName authorityName="Reitter" authorityYear="1889" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="subtilis">
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<emphasis italics="true" pageId="98" pageNumber="99">Thymalus subtilis</emphasis>
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|
</taxonomicName>
|
|
Reitter, 1889.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection lastPageId="99" lastPageNumber="100" pageId="98" pageNumber="99" type="description">
|
|
<paragraph pageId="98" pageNumber="99">Description.</paragraph>
|
|
<paragraph lastPageId="99" lastPageNumber="100" pageId="98" pageNumber="99">
|
|
Body size: 4.3-7.5 mm. Body shape convex (not conglobate). Gular sutures wide, subparallel. Frontoclypeal suture absent. Frons: longitudinal groove or depression absent. Cranium ventrally: tufts of long setae at sides absent. Submentum: ctenidium absent. Antennal groove absent. Eyes: size large, lateral. Eyes number: two. Epicranial acumination moderate. Lacinial hooks: two. Galea: shape elongate. Galea: ciliate setae absent. Mediostipes-Lacinia not fused. Palpifer: outer edge even. Mandibular apical teeth number: two, horizontally situated. Mola present. Penicillus (at base) absent. Pubescence above mola or cutting edge absent. Ventral furrow absent. Basal notch moderate. Labrum-Cranium not fused. Epipharyngial sclerite absent. Lat
|
|
<pageBreakToken pageId="99" pageNumber="100" start="start">eral</pageBreakToken>
|
|
tormal process: projection projection not developed (all remaining). Ligula: ciliate setae absent. Ligula rigid, not retroflexed, weakly emarginate. Hypopharyngeal sclerite H-shaped. Antenna 11-segmented. Antennal club symmetrical, sensorial fields absent. Front coxal cavities externally open, internally closed. Pronotum transverse. Prepectus absent. Middle coxal cavities open. Elytra: long hairs absent. Epipleuron wide. Elytral interlocking mechanism absent, carinae reduced. Elytral punctation regular, scales absent. Wing: radial cell oblong (or reduced), wedge cell present, cross vein MP3-4 present, cross vein AA1+2-3+4 absent. Front tibiae: spines along side reduced. Hooked spur absent, apical spurs not hooked or weakly hooked. Claws: denticle absent. Parasternites number along ventrites III-VII: one. Spiculum gastrale absent. Tegmen composed of three parts.
|
|
</paragraph>
|
|
<paragraph pageId="99" pageNumber="100">Larva: Frontal arms V-shaped. Epicranial stem reduced. Endocarina absent. Gular sutures inconspicuous. Gula: anterior apodemes present. Paragular sclerites absent. Hypostomal rods present. Stemmata number: five. Mandibular apical teeth number: two, horizontally situated. Lacinia mandibulae with several small spines. Mola absent. Maxillary palpi 3-segmented. Palpifer present. Pedunculate seta absent. Mala simple. Mala: bidentate protrusion present. Cardo-Stipes partially fused. Cardo: size much smaller than stipes. Ligula present. Labial palpi 2-segmented. Prementum in single part, anterior margin even. Torma: two separate lateral sclerites. Antennal joints 1, 2 transverse. Sensory appendix larger than half of joint 3. Thoracic sclerites pattern (dorsally) 2-0-0. Thoracic sclerites pattern (ventrally) 0+0+0. Trochanter triangular. Abdominal segment IX not divided. Tergite IX flat. Urogomphi present, hooked; median process absent.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="99" pageNumber="100" type="biology">
|
|
<paragraph pageId="99" pageNumber="100">Biology.</paragraph>
|
|
<paragraph pageId="99" pageNumber="100">
|
|
The beetles are not associated with any particular tree species and are found on both deciduous and coniferous trees.
|
|
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1787" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="limbatus">
|
|
<emphasis italics="true" pageId="99" pageNumber="100">Thymalus limbatus</emphasis>
|
|
</taxonomicName>
|
|
is known from the trunks of birch, beech, linden, and spruce, mostly under bark. It is assumed that the larvae feed on fungi in rotten or decaying wood (
|
|
<bibRefCitation author="Kolibac, J" journalOrPublisher="Entomologica Basiliensia et Collectionis Frey" pageId="155" pageNumber="156" pagination="39 - 159" refId="B145" refString="Kolibac, J, 2005. A review of the Trogositidae. Part 1: Morphology of the genera (Coleoptera, Cleroidea). Entomologica Basiliensia et Collectionis Frey 27: 39 - 159" title="A review of the Trogositidae. Part 1: Morphology of the genera (Coleoptera, Cleroidea)." volume="27" year="2005">
|
|
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
|
|
et al. 2005
|
|
</bibRefCitation>
|
|
).
|
|
<taxonomicName authorityName="Chevrolat" authorityYear="1842" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="marginicollis">
|
|
<emphasis italics="true" pageId="99" pageNumber="100">Thymalus marginicollis</emphasis>
|
|
</taxonomicName>
|
|
has been collected from the fungi
|
|
<taxonomicName class="Agaricomycetes" family="Polyporaceae" genus="Polyporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="" order="Polyporales" pageId="99" pageNumber="100" phylum="Basidiomycota" rank="species" species="betulinus">
|
|
<emphasis italics="true" pageId="99" pageNumber="100">Polyporus betulinus</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName class="Agaricomycetes" family="Polyporaceae" genus="Polyporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="" order="Polyporales" pageId="99" pageNumber="100" phylum="Basidiomycota" rank="species" species="versicolor">
|
|
<emphasis italics="true" pageId="99" pageNumber="100">Polyporus versicolor</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName class="Agaricomycetes" family="Fomitopsidaceae" genus="Daedalea" higherTaxonomySource="GBIF" kingdom="Fungi" lsidName="" order="Polyporales" pageId="99" pageNumber="100" phylum="Basidiomycota" rank="species" species="confragosa">
|
|
<emphasis italics="true" pageId="99" pageNumber="100">Daedalea confragosa</emphasis>
|
|
</taxonomicName>
|
|
, on the trunks of birch and also on "
|
|
<emphasis italics="true" pageId="99" pageNumber="100">wild flowers in plant press</emphasis>
|
|
" (
|
|
<bibRefCitation author="Barron, JR" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="147" pageNumber="148" pagination="1 - 143" publicationUrl="10.4039/entm10375fv" refId="B13" refString="Barron, JR, 1971. A revision of the Trogositidae of America North of Mexico (Coleoptera: Cleroidea). Memoirs of the Entomological Society of Canada 75: 1 - 143, 10.4039/entm10375fv" title="A revision of the Trogositidae of America North of Mexico (Coleoptera: Cleroidea)." url="10.4039/entm10375fv" volume="75" year="1971">Barron 1971</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="99" pageNumber="100" type="distribution">
|
|
<paragraph pageId="99" pageNumber="100">Distribution.</paragraph>
|
|
<paragraph pageId="99" pageNumber="100">Holoarctic: Northern states of USA, Canada, Europe, North Africa, Siberia to China and Japan. Some specimens, probably a new species, have recently been collected in Chinese Sichuan and Yunnan and also in northern Thailand.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection lastPageId="100" lastPageNumber="101" pageId="99" pageNumber="100" type="species">
|
|
<paragraph pageId="99" pageNumber="100">Species:</paragraph>
|
|
<paragraph pageId="99" pageNumber="100">
|
|
<taxonomicName authorityName="Leveille" authorityYear="1877" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="aubei">
|
|
<emphasis italics="true" pageId="99" pageNumber="100">Thymalus aubei</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken originalValue="Léveillé">Leveille</normalizedToken>
|
|
, 1877;
|
|
<normalizedToken originalValue="“Batum”">"Batum"</normalizedToken>
|
|
, Caucasus (varA)
|
|
</paragraph>
|
|
<paragraph pageId="99" pageNumber="100">
|
|
<normalizedToken originalValue="Léveillé">Leveille</normalizedToken>
|
|
, A. 1910: 32. Klausnitzer, B. 1996: 156. Barron, J. R. 1971: 36 (syn.
|
|
<taxonomicName authorityName="Leveille" authorityYear="1877" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="aubei">
|
|
<emphasis italics="true" pageId="99" pageNumber="100">Thymalus aubei</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken originalValue="Léveillé">Leveille</normalizedToken>
|
|
, 1877 with
|
|
<taxonomicName authorityName="Chevrolat" authorityYear="1842" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="marginicollis">
|
|
<emphasis italics="true" pageId="99" pageNumber="100">Thymalus marginicollis</emphasis>
|
|
</taxonomicName>
|
|
Chevrolat, 1842). Barron, J. R. 1971: 36 (syn.
|
|
<taxonomicName class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="fulgidus">
|
|
<emphasis italics="true" pageId="99" pageNumber="100">Thymalus fulgidus</emphasis>
|
|
</taxonomicName>
|
|
var.
|
|
<emphasis italics="true" pageId="99" pageNumber="100">aubei</emphasis>
|
|
<normalizedToken originalValue="Léveillé">Leveille</normalizedToken>
|
|
, 1877 with
|
|
<taxonomicName authorityName="Chevrolat" authorityYear="1842" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="marginicollis">
|
|
<emphasis italics="true" pageId="99" pageNumber="100">Thymalus marginicollis</emphasis>
|
|
</taxonomicName>
|
|
Chevrolat, 1842).
|
|
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
|
|
, J. 2007a: 366 (syn.
|
|
<taxonomicName authorityName="Reitter" authorityYear="1889" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="subtilis">
|
|
<emphasis italics="true" pageId="99" pageNumber="100">Thymalus subtilis</emphasis>
|
|
</taxonomicName>
|
|
Reitter, 1889). Lafer, G. Sh. 1992: 86 (
|
|
<taxonomicName authorityName="Reitter" authorityYear="1889" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="subtilis">
|
|
<emphasis italics="true" pageId="99" pageNumber="100">Thymalus subtilis</emphasis>
|
|
</taxonomicName>
|
|
Reitter, 1889).
|
|
<normalizedToken originalValue="Léveillé">Leveille</normalizedToken>
|
|
, A. 1910: 33 (
|
|
<taxonomicName authorityName="Reitter" authorityYear="1889" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="subtilis">
|
|
<emphasis italics="true" pageId="99" pageNumber="100">Thymalus subtilis</emphasis>
|
|
</taxonomicName>
|
|
Reitter, 1889). Nikitsky, N. B. et al. 1998: 28 (syn.
|
|
<taxonomicName authorityName="Reitter" authorityYear="1889" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="subtilis">
|
|
<emphasis italics="true" pageId="99" pageNumber="100">Thymalus subtilis</emphasis>
|
|
</taxonomicName>
|
|
Reitter, 1889; lectotype designated). Nikitsky, N. B. & Semenov, V. B. 2001: 49
|
|
</paragraph>
|
|
<paragraph pageId="100" pageNumber="101">
|
|
<taxonomicName class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="chinensis">
|
|
<emphasis italics="true" pageId="100" pageNumber="101">
|
|
<pageBreakToken pageId="100" pageNumber="101" start="start">Thymalus</pageBreakToken>
|
|
chinensis
|
|
</emphasis>
|
|
</taxonomicName>
|
|
Fairmaire, 1900; China: Fujian (JK)
|
|
</paragraph>
|
|
<paragraph pageId="100" pageNumber="101">
|
|
<normalizedToken originalValue="Léveillé">Leveille</normalizedToken>
|
|
, A. 1910: 32.
|
|
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
|
|
, J. 2007a: 366
|
|
</paragraph>
|
|
<paragraph pageId="100" pageNumber="101">
|
|
<taxonomicName class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="laticeps">
|
|
<emphasis italics="true" pageId="100" pageNumber="101">Thymalus laticeps</emphasis>
|
|
</taxonomicName>
|
|
Lewis, 1894; Japan (varA)
|
|
</paragraph>
|
|
<paragraph pageId="100" pageNumber="101">
|
|
<normalizedToken originalValue="Léveillé">Leveille</normalizedToken>
|
|
, A. 1910: 32. Esaki, T. et al. 1951: 1063.
|
|
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
|
|
, J. 2007a: 366. Lafer, G. Sh. 1992: 86. Nakane, T. et al. 1963: 181
|
|
</paragraph>
|
|
<paragraph pageId="100" pageNumber="101">
|
|
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1787" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="limbatus">
|
|
<emphasis italics="true" pageId="100" pageNumber="101">Thymalus limbatus</emphasis>
|
|
</taxonomicName>
|
|
Fabricius, 1787; Europe, North Africa: Tunisia (JK)
|
|
</paragraph>
|
|
<paragraph pageId="100" pageNumber="101">
|
|
<normalizedToken originalValue="Léveillé">Leveille</normalizedToken>
|
|
, A. 1910: 32. Alexander, K. N. A. 1996: 90 (biology). Bahillo de la Puebla, P. &
|
|
<normalizedToken originalValue="López-Colón">Lopez-Colon</normalizedToken>
|
|
, J. I. 1999: 13. Bahillo de la Puebla, P. &
|
|
<normalizedToken originalValue="López-Colón">Lopez-Colon</normalizedToken>
|
|
, J. I. 2004: 129. Bercedo, P. et al. 2006: 180 (distribution). Borowiec, L. 1983: 15. Burakowski, B. et al. 1986: 123. Cunev, J. 1999: 76. Franz, H. 1981: 51-52 (distribution). Klausnitzer, B. 1976: 8. Klausnitzer, B. 1978: 176. Klausnitzer, B. 1996: 155.
|
|
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
|
|
, J. 1993a: 20.
|
|
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
|
|
, J. 1993b: 90.
|
|
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
|
|
, J. 2002: 55 (larva).
|
|
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
|
|
, J. 2005: 85 (redescription).
|
|
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
|
|
, J. 2006: 110 (larva).
|
|
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
|
|
, J. 2007a: 366 (distribution).
|
|
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
|
|
, J. 2007a: 366 (syn.
|
|
<emphasis italics="true" pageId="100" pageNumber="101">brunneus</emphasis>
|
|
Thunberg, 1794).
|
|
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
|
|
, J. 2007a: 366 (syn.
|
|
<emphasis italics="true" pageId="100" pageNumber="101">rubiginosus</emphasis>
|
|
Gmelin, 1790). Krasutskii, B. V. 1996: 274. Mitter, H. 1998: 561. Pileckis, S. &
|
|
<normalizedToken originalValue="Monsevičius">Monsevicius</normalizedToken>
|
|
, V. 1995: 273. Ratti, E. 1997: 178. Reitter, E. 1876: 64. Theunert, R. 2006: 113-114 (distribution). Vogt, H. 1967: 18
|
|
</paragraph>
|
|
<paragraph pageId="100" pageNumber="101">
|
|
<taxonomicName authorityName="Chevrolat" authorityYear="1842" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="marginicollis">
|
|
<emphasis italics="true" pageId="100" pageNumber="101">Thymalus marginicollis</emphasis>
|
|
</taxonomicName>
|
|
Chevrolat, 1842; Canada, USA (JRB)
|
|
</paragraph>
|
|
<paragraph pageId="100" pageNumber="101">
|
|
Barron, J. R. 1971: 36 (syn.
|
|
<taxonomicName authorityName="Leveille" authorityYear="1877" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="aubei">
|
|
<emphasis italics="true" pageId="100" pageNumber="101">Thymalus aubei</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken originalValue="Léveillé">Leveille</normalizedToken>
|
|
, 1877). Barron, J. R. 1971: 36 (syn.
|
|
<taxonomicName class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="fulgidus">
|
|
<emphasis italics="true" pageId="100" pageNumber="101">Thymalus fulgidus</emphasis>
|
|
</taxonomicName>
|
|
Erichson, 1844). Barron, J. R. 1971: 36 (syn.
|
|
<taxonomicName class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="fulgidus">
|
|
<emphasis italics="true" pageId="100" pageNumber="101">Thymalus fulgidus</emphasis>
|
|
</taxonomicName>
|
|
var.
|
|
<emphasis italics="true" pageId="100" pageNumber="101">aubei</emphasis>
|
|
<normalizedToken originalValue="Léveillé">Leveille</normalizedToken>
|
|
, 1877).
|
|
<normalizedToken originalValue="Böving">Boeving</normalizedToken>
|
|
, A. G. & Craighead, F. C. 1931: 273 (larva). Dajoz, R. 1997: 44 (biology). Reitter, E. 1876: 64 (
|
|
<taxonomicName class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="fulgidus">
|
|
<emphasis italics="true" pageId="100" pageNumber="101">Thymalus fulgidus</emphasis>
|
|
</taxonomicName>
|
|
Erichson, 1844: "Amer. bor.")
|
|
</paragraph>
|
|
<paragraph pageId="100" pageNumber="101">
|
|
<taxonomicName authorityName="Reitter" authorityYear="1889" class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="oblongus">
|
|
<emphasis italics="true" pageId="100" pageNumber="101">Thymalus oblongus</emphasis>
|
|
</taxonomicName>
|
|
Reitter, 1889; Russia: North and Central Europea teritorries, Sweden, East Siberia (JK)
|
|
</paragraph>
|
|
<paragraph pageId="100" pageNumber="101">
|
|
<normalizedToken originalValue="Léveillé">Leveille</normalizedToken>
|
|
, A. 1910: 33. Lafer, G. Sh. 1992: 86.
|
|
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
|
|
, J. 2007a: 366. Krasutskii, B. V. 2006: 763 (biology). Nikitsky, N. B. et al. 1998: 28 (lectotype designated)
|
|
</paragraph>
|
|
<paragraph pageId="100" pageNumber="101">
|
|
<taxonomicName class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="parviceps">
|
|
<emphasis italics="true" pageId="100" pageNumber="101">Thymalus parviceps</emphasis>
|
|
</taxonomicName>
|
|
Lewis, 1894; Japan (varA)
|
|
</paragraph>
|
|
<paragraph pageId="100" pageNumber="101">
|
|
<normalizedToken originalValue="Léveillé">Leveille</normalizedToken>
|
|
, A. 1910: 33. Esaki, T. et al. 1951: 1063.
|
|
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
|
|
, J. 2007a: 366. Lafer, G. Sh. 1992: 86. Nakane, T. et al. 1963: 181
|
|
</paragraph>
|
|
<paragraph pageId="100" pageNumber="101">
|
|
<taxonomicName class="Hexapoda" family="Trogossitidae" genus="Thymalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="punctidorsum">
|
|
<emphasis italics="true" pageId="100" pageNumber="101">Thymalus punctidorsum</emphasis>
|
|
</taxonomicName>
|
|
Latreille, 1894; Japan (varA)
|
|
</paragraph>
|
|
<paragraph pageId="100" pageNumber="101">
|
|
<normalizedToken originalValue="Léveillé">Leveille</normalizedToken>
|
|
, A. 1910: 33.
|
|
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
|
|
, J. 2007a: 366. Lafer, G. Sh. 1992: 86. Nakane, T. et al. 1963: 181
|
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |