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<document id="7FB6DE72C8C2C0C94AAA06BF77C621C4" ID-DOI="10.1080/002229301300323910" ID-ISSN="1464-5262" ID-Zenodo-Dep="5277129" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630022732653" checkinUser="felipe" docAuthor="Gussmann, S. M. V. &amp; Scholtz, C. H." docDate="2001" docId="DA651E6CFF87FB29FE4B3506FDA1FB75" docLanguage="en" docName="JNatHist.35.7.1013-1084.pdf" docOrigin="Journal of Natural History 35 (7)" docSource="http://www.tandfonline.com/doi/abs/10.1080/002229301300323910" docStyle="DocumentStyle:B6EF65DE5B4D2965D4D236BA736D172F.2:JNatHist.2000-2002.journal_article.2cover" docStyleId="B6EF65DE5B4D2965D4D236BA736D172F" docStyleName="JNatHist.2000-2002.journal_article.2cover" docStyleVersion="2" docTitle="Bolbaoeer guineaensis Gussmann &amp; Scholtz 2001, sp. nov." docType="treatment" docVersion="4" lastPageNumber="1054" masterDocId="265C6614FFADFB02FFDA3151FFDBFFCD" masterDocTitle="Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species" masterLastPageNumber="1084" masterPageNumber="1013" pageNumber="1053" updateTime="1699137266390" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="EF94BFF82D94471B638E792D250240F8">Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species</mods:title>
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<mods:namePart id="D6CDC412E7691ED39023E86612A48F1A">Gussmann, S. M. V.</mods:namePart>
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<mods:namePart id="25676853A502882C1E6FB10B6C8FFCC3">Scholtz, C. H.</mods:namePart>
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<mods:title id="FCBBCB27BC30005644D79196B8FB68E6">Journal of Natural History</mods:title>
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<mods:date id="B0023C475DBAD4CDDE60E37624456ECE">2001</mods:date>
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<mods:number id="AA4C46E6D8526E62683A19F8B511D88D">2001-07-31</mods:number>
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<mods:number id="BF8EEB0A5BC2ABDF69376BABFBB74C4C">35</mods:number>
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<mods:identifier id="8EF2379C81C0AB691DE591C6A4E72C82" type="ISSN">1464-5262</mods:identifier>
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<subSubSection id="1AD6FCF1FF87FB28FE4B3506FD7AFB42" pageId="42" pageNumber="1053" type="nomenclature">
<paragraph id="5273AF7AFF87FB28FE4B3506FD3AFBA2" blockId="42.[401,737,1111,1168]" box="[401,737,1111,1135]" pageId="42" pageNumber="1053">
<heading id="093B1816FF87FB28FE4B3506FD3AFBA2" box="[401,737,1111,1135]" centered="true" fontSize="10" level="2" pageId="42" pageNumber="1053" reason="3">
<taxonomicName id="95CCD4F9FF87FB28FE4B3506FDA5FBA2" authority="Gussmann &amp; Scholtz, 2001" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[401,638,1111,1135]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="1053" phylum="Arthropoda" rank="species" species="guineaensis" status="sp. nov.">
<emphasis id="60B87368FF87FB28FE4B3506FDA5FBA2" bold="true" box="[401,638,1111,1135]" italics="true" pageId="42" pageNumber="1053">BolbaOEer guineaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7B8BCE13FF87FB28FD5C3509FD3AFBA2" box="[646,737,1112,1135]" pageId="42" pageNumber="1053" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="5273AF7AFF87FB28FE0B3526FD7AFB42" blockId="42.[401,737,1111,1168]" box="[465,673,1143,1168]" pageId="42" pageNumber="1053">(®gures 3o, 15a±e)</paragraph>
</subSubSection>
<subSubSection id="1AD6FCF1FF87FB29FF863590FDA1FB75" lastPageId="43" lastPageNumber="1054" pageId="42" pageNumber="1053" type="description">
<paragraph id="5273AF7AFF87FB28FF863590FEA4FB15" blockId="42.[92,1047,1216,1686]" box="[92,383,1216,1240]" pageId="42" pageNumber="1053">
<emphasis id="60B87368FF87FB28FF863590FEA4FB15" box="[92,383,1216,1240]" italics="true" pageId="42" pageNumber="1053">
Description male
<typeStatus id="8D7711D8FF87FB28FEFB3591FEA4FB15" box="[289,383,1216,1240]" pageId="42" pageNumber="1053" type="holotype">holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="5273AF7AFF87FB29FF5E35B1FDC1FEB6" blockId="42.[92,1047,1216,1686]" lastBlockId="43.[92,1047,132,1208]" lastPageId="43" lastPageNumber="1054" pageId="42" pageNumber="1053">
<emphasis id="60B87368FF87FB28FF5E35B1FED1FB35" box="[132,266,1248,1272]" italics="true" pageId="42" pageNumber="1053">Body length</emphasis>
<quantity id="9534029FFF87FB28FECE35B1FEACFB35" box="[276,375,1248,1272]" metricMagnitude="-2" metricUnit="m" metricValue="2.16" pageId="42" pageNumber="1053" unit="mm" value="21.6">21.6 mm</quantity>
.
<emphasis id="60B87368FF87FB28FE5E35B1FE14FB35" box="[388,463,1248,1272]" italics="true" pageId="42" pageNumber="1053">Colour</emphasis>
mid-brown.
<emphasis id="60B87368FF87FB28FDB035B1FC4BFB35" box="[618,912,1248,1272]" italics="true" pageId="42" pageNumber="1053">Outer margin of mandible</emphasis>
very feebly sinuate.
<emphasis id="60B87368FF87FB28FF663451FECFFAD5" box="[188,276,1280,1304]" italics="true" pageId="42" pageNumber="1053">Clypeus</emphasis>
with bituberculate, moderately arcuate transverse posterior carina; tubercles in line with clypeo-frontal transition; posterior carina slightly shorter than anterior carina.
<emphasis id="60B87368FF87FB28FEC13410FE82FA95" box="[283,345,1345,1368]" italics="true" pageId="42" pageNumber="1053">Frons</emphasis>
moderately depressed posterior to posterior clypeal carina.
<emphasis id="60B87368FF87FB28FF863431FF28FAB5" box="[92,243,1376,1400]" italics="true" pageId="42" pageNumber="1053">Antennal club</emphasis>
with glabrous area of basal segment about one-half of exposed surface.
<emphasis id="60B87368FF87FB28FF8634D0FF12FA55" box="[92,201,1409,1432]" italics="true" pageId="42" pageNumber="1053">Pronotum</emphasis>
with anterior margin in dorsal view medially very feebly bisinuate and not projecting over frons; with one pair of outer, short, slightly inward inclined horns near posterior margin (®gure 3o); disc with both ®ne and large, well-spaced punctures, these increasingly densely spaced towards sides and along anterior margin.
<taxonomicName id="95CCD4F9FF87FB28FF8634AEFF13F9DA" box="[92,200,1535,1559]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="42" pageNumber="1053" phylum="Arthropoda" rank="genus">
<emphasis id="60B87368FF87FB28FF8634AEFF13F9DA" box="[92,200,1535,1559]" italics="true" pageId="42" pageNumber="1053">Scutellum</emphasis>
</taxonomicName>
with surface moderately punctate.
<emphasis id="60B87368FF87FB28FD8634AEFD74F9DA" box="[604,687,1535,1559]" italics="true" pageId="42" pageNumber="1053">Elytron</emphasis>
with strial punctures separated by four to six puncture diameters; intervals with approximately ®ve to six punctures between two striae.
<emphasis id="60B87368FF87FB28FE9B376EFE47F99A" box="[321,412,1599,1623]" italics="true" pageId="42" pageNumber="1053">Protibia</emphasis>
®ve-dentate.
<emphasis id="60B87368FF87FB28FDE2376EFD09F99A" box="[568,722,1599,1623]" italics="true" pageId="42" pageNumber="1053">Protibial spur</emphasis>
longer than ®fth tarsomere, acuminate.
<emphasis id="60B87368FF87FB28FF39370EFE54F9BA" box="[227,399,1631,1655]" italics="true" pageId="42" pageNumber="1053">Metatrochanter</emphasis>
of unmodi®ed shape; surface in basal third and along posterior margin with long and fairly dense setation, remaining surface asetose.
<emphasis id="60B87368FF86FB29FF8631D5FF0CFF51" box="[92,215,132,156]" italics="true" pageId="43" pageNumber="1054">Metafemur</emphasis>
in ventral view in posterior third with line of densely spaced moderately long and long setae; with fairly dense, long setation in anterior third and in apical area; with fairly dense, long setae along posterior margin; remaining areas with some isolated setae.
<emphasis id="60B87368FF86FB29FE9431B5FE60FF31" box="[334,443,228,252]" italics="true" pageId="43" pageNumber="1054">Metatibia</emphasis>
in lateral view with subapical carina bilobed; with unmodi®ed spurs.
<emphasis id="60B87368FF86FB29FEF63055FE41FED1" box="[300,410,260,284]" italics="true" pageId="43" pageNumber="1054">Underside</emphasis>
with sternites 4 and 5 unmodi®ed (®gure 15b); posterior margin of sternite 6 very feebly concave (®gure 15b); pygidium unmodi®ed (®gure 15a).
<emphasis id="60B87368FF86FB29FF353015FE8FFE91" box="[239,340,324,348]" italics="true" pageId="43" pageNumber="1054">Genitalia</emphasis>
with aedeagus very small, as in ®gure 15c±e; genital capsule apically with moderately long setation.
</paragraph>
<paragraph id="5273AF7AFF86FB29FF5E30D2FE96FE56" blockId="43.[92,1047,132,1208]" box="[132,333,387,411]" pageId="43" pageNumber="1054">
<emphasis id="60B87368FF86FB29FF5E30D2FF09FE56" box="[132,210,387,411]" italics="true" pageId="43" pageNumber="1054">Female</emphasis>
unknown.
</paragraph>
<paragraph id="5273AF7AFF86FB29FF5E30F2FC41FE36" blockId="43.[92,1047,132,1208]" pageId="43" pageNumber="1054">
<emphasis id="60B87368FF86FB29FF5E30F2FED3FE76" box="[132,264,419,443]" italics="true" pageId="43" pageNumber="1054">Distribution</emphasis>
. The only label data`Guinea is presumed to refer to the West African country`Guinea and not to`Guinea-Bissau or`Equatorial Guinea. Since proper locality data are missing, the species distribution is not mapped.
</paragraph>
<paragraph id="5273AF7AFF86FB29FF5E3355FCC6FDD6" blockId="43.[92,1047,132,1208]" box="[132,797,515,539]" pageId="43" pageNumber="1054">
<emphasis id="60B87368FF86FB29FF5E3355FE49FDD6" box="[132,402,515,539]" italics="true" pageId="43" pageNumber="1054">Type material examined</emphasis>
.
<materialsCitation id="E2A4A527FF86FB29FE793355FCC6FDD6" box="[419,797,515,539]" collectionCode="ZMHB" pageId="43" pageNumber="1054" specimenCount="1" typeStatus="holotype">
<typeStatus id="8D7711D8FF86FB29FE793355FDCEFDD6" box="[419,533,515,539]" pageId="43" pageNumber="1054" type="holotype">HOLOTYPE</typeStatus>
:`Guinea (
<collectionCode id="34DD37BFFF86FB29FD623352FCD5FDD6" box="[696,782,515,539]" pageId="43" pageNumber="1054">ZMHB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="5273AF7AFF86FB29FF5E3372FD46FDF6" blockId="43.[92,1047,132,1208]" box="[132,669,547,572]" pageId="43" pageNumber="1054">
<emphasis id="60B87368FF86FB29FF5E3372FF20FDF6" box="[132,251,547,571]" italics="true" pageId="43" pageNumber="1054">Etymology</emphasis>
. Named after the country of origin.
</paragraph>
<paragraph id="5273AF7AFF86FB29FF5E3313FE34FB55" blockId="43.[92,1047,132,1208]" pageId="43" pageNumber="1054">
<emphasis id="60B87368FF86FB29FF5E3313FF23FD97" box="[132,248,578,602]" italics="true" pageId="43" pageNumber="1054">Comments</emphasis>
. The presence of only one pair of horns along the posterior pronotal margin makes the single male specimen of this new species similar in appearance to males of
<taxonomicName id="95CCD4F9FF86FB29FF1D33D2FE80FD57" box="[199,347,642,666]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="1054" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="60B87368FF86FB29FF1D33D2FE80FD57" box="[199,347,642,666]" italics="true" pageId="43" pageNumber="1054">B. splendidus</emphasis>
</taxonomicName>
. There are, however, signi®cant diOEerences between the two species: both metatibial spurs are unmodi®ed in
<taxonomicName id="95CCD4F9FF86FB29FD4C33F2FCECFD77" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[662,823,675,698]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="1054" phylum="Arthropoda" rank="species" species="guineaensis">
<emphasis id="60B87368FF86FB29FD4C33F2FCECFD77" box="[662,823,675,698]" italics="true" pageId="43" pageNumber="1054">B. guineaensis</emphasis>
</taxonomicName>
but one metatibial spur is hooked in
<taxonomicName id="95CCD4F9FF86FB29FEF03392FE66FD17" box="[298,445,706,730]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="1054" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="60B87368FF86FB29FEF03392FE66FD17" box="[298,445,706,730]" italics="true" pageId="43" pageNumber="1054">B. splendidus</emphasis>
</taxonomicName>
; modi®cations on abdominal sternites and pygidium are virtually absent in
<taxonomicName id="95CCD4F9FF86FB29FEB633B2FDD7FD37" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[364,524,739,762]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="1054" phylum="Arthropoda" rank="species" species="guineaensis">
<emphasis id="60B87368FF86FB29FEB633B2FDD7FD37" box="[364,524,739,762]" italics="true" pageId="43" pageNumber="1054">B. guineaensis</emphasis>
</taxonomicName>
but are strong and distinct in
<taxonomicName id="95CCD4F9FF86FB29FC5833B2FBCCFD37" box="[898,1047,738,762]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="1054" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="60B87368FF86FB29FC5833B2FBCCFD37" box="[898,1047,738,762]" italics="true" pageId="43" pageNumber="1054">B. splendidus</emphasis>
</taxonomicName>
(compare ®gure 15a, b with 7f±h). Lastly, the aedeagi of both species bear no similarity in shape at all (compare ®gure 15c±e with 8d±g). The small size of the aedeagus of
<taxonomicName id="95CCD4F9FF86FB29FF323212FE5EFC97" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[232,389,835,858]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="1054" phylum="Arthropoda" rank="species" species="guineaensis">
<emphasis id="60B87368FF86FB29FF323212FE5EFC97" box="[232,389,835,858]" italics="true" pageId="43" pageNumber="1054">B. guineaensis</emphasis>
</taxonomicName>
is unusual as well. The presence of only one pair of horns and the general lack of other morphological modi®cations make
<taxonomicName id="95CCD4F9FF86FB29FC843232FC24FCB7" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[862,1023,867,890]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="1054" phylum="Arthropoda" rank="species" species="guineaensis">
<emphasis id="60B87368FF86FB29FC843232FC24FCB7" box="[862,1023,867,890]" italics="true" pageId="43" pageNumber="1054">B. guineaensis</emphasis>
</taxonomicName>
a unique species, easy to identify. Judging from the position and size of the pronotal horns and through comparison with males of
<taxonomicName id="95CCD4F9FF86FB29FD5D32F2FCC6FC77" box="[647,797,930,954]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="1054" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="60B87368FF86FB29FD5D32F2FCC6FC77" box="[647,797,930,954]" italics="true" pageId="43" pageNumber="1054">B. splendidus</emphasis>
</taxonomicName>
, we assume that the
<typeStatus id="8D7711D8FF86FB29FF863290FF65FC14" box="[92,190,961,985]" pageId="43" pageNumber="1054" type="holotype">holotype</typeStatus>
of
<taxonomicName id="95CCD4F9FF86FB29FF323293FE5DFC14" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[232,390,962,985]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="1054" phylum="Arthropoda" rank="species" species="guineaensis">
<emphasis id="60B87368FF86FB29FF323293FE5DFC14" box="[232,390,962,985]" italics="true" pageId="43" pageNumber="1054">B. guineaensis</emphasis>
</taxonomicName>
represents either a minor or intermediate male.
<taxonomicName id="95CCD4F9FF86FB29FC6B3290FF0CFC34" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="1054" phylum="Arthropoda" rank="species" species="guineaensis">
<emphasis id="60B87368FF86FB29FC6B3290FF0CFC34" italics="true" pageId="43" pageNumber="1054">BolbaOEer guineaensis</emphasis>
</taxonomicName>
cannot be confused with males of the allopatric
<taxonomicName id="95CCD4F9FF86FB29FCD632B3FCBDFC34" box="[780,870,994,1017]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="1054" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="60B87368FF86FB29FCD632B3FCBDFC34" box="[780,870,994,1017]" italics="true" pageId="43" pageNumber="1054">B. gigas</emphasis>
</taxonomicName>
,
<taxonomicName id="95CCD4F9FF86FB29FCAC32B3FC38FC34" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[886,995,993,1017]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="1054" phylum="Arthropoda" rank="species" species="sasakii">
<emphasis id="60B87368FF86FB29FCAC32B3FC38FC34" box="[886,995,993,1017]" italics="true" pageId="43" pageNumber="1054">B. sasakii</emphasis>
</taxonomicName>
and
<taxonomicName id="95CCD4F9FF86FB29FF863553FF20FBD4" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[92,251,1025,1049]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="1054" phylum="Arthropoda" rank="species" species="namibiensis">
<emphasis id="60B87368FF86FB29FF863553FF20FBD4" box="[92,251,1025,1049]" italics="true" pageId="43" pageNumber="1054">B. namibiensis</emphasis>
</taxonomicName>
, the remaining other species that possess only one pair of horns along the posterior pronotal margin: the horns of these three species, even in minor males, are much further apart from one another than in
<taxonomicName id="95CCD4F9FF86FB29FD403513FCE3FB94" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[666,824,1090,1113]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="1054" phylum="Arthropoda" rank="species" species="guineaensis">
<emphasis id="60B87368FF86FB29FD403513FCE3FB94" box="[666,824,1090,1113]" italics="true" pageId="43" pageNumber="1054">B. guineaensis</emphasis>
</taxonomicName>
(compare ®gure 3o with 4i, l, s) and sternal and pygidial modi®cations diOEer signi®cantly as well (compare descriptions and ®gures).
</paragraph>
<paragraph id="5273AF7AFF86FB29FF5E35F1FDA1FB75" blockId="43.[92,1047,132,1208]" box="[132,634,1184,1208]" pageId="43" pageNumber="1054">
Females of
<taxonomicName id="95CCD4F9FF86FB29FED035F0FE72FB75" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[266,425,1185,1208]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="1054" phylum="Arthropoda" rank="species" species="guineaensis">
<emphasis id="60B87368FF86FB29FED035F0FE72FB75" box="[266,425,1185,1208]" italics="true" pageId="43" pageNumber="1054">B. guineaensis</emphasis>
</taxonomicName>
remain unknown.
</paragraph>
</subSubSection>
</treatment>
</document>