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treatments-xml/data/03/EF/62/03EF6217BF32FFCC0AC0F9DD980EAAF0.xml
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<document id="D4F6E37714EC242797855127659FEC7A" ID-CLB-Dataset="20762" ID-DOI="10.11646/zootaxa.3365.1.1" ID-GBIF-Dataset="1db9bc74-46dd-4263-9784-88a59bd05d59" ID-ISSN="1175­5334" ID-Zenodo-Dep="6038900" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629787146833" checkinUser="felipe" docAuthor="Garcia, Francisco Hita &amp; Fisher, Brian L." docDate="2012" docId="03EF6217BF32FFCC0AC0F9DD980EAAF0" docLanguage="en" docName="zt03365p123.pdf" docOrigin="Zootaxa 3365" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Tetramorium artemis Hita Garcia &amp; Fisher 2012, sp. n." docType="treatment" docVersion="8" lastPageNumber="15" masterDocId="FFD61A6FBF39FFC20A57FF889C5BAB77" masterDocTitle="The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region — taxonomy of the T. bessonii, T. bonibony, T. dysalum, T. marginatum, T. tsingy, and T. weitzeckeri species groups" masterLastPageNumber="123" masterPageNumber="1" pageNumber="12" updateTime="1699142484625" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="DE19E0B54DC329B230B9CD1D65E19203" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F3D9CFE4540C907794CEB94AEA613119">
<mods:title id="488E4026BC74A871206AE74D19D122A4">The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region — taxonomy of the T. bessonii, T. bonibony, T. dysalum, T. marginatum, T. tsingy, and T. weitzeckeri species groups</mods:title>
</mods:titleInfo>
<mods:name id="6041960421AE8DE19E08A9D94FB56E06" type="personal">
<mods:role id="5B0D3F777CA0E5A0C6CAC0D5348D3610">
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</mods:role>
<mods:namePart id="A08765CA403B644AFC8F5158DAC20092">Garcia, Francisco Hita</mods:namePart>
</mods:name>
<mods:name id="D6875359E19FFA78D56FEA40B6375734" type="personal">
<mods:role id="E0FE16C41066821E8655CDB116F9D401">
<mods:roleTerm id="0528DCCCEEF38A7C548978B050515905">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6BD5EB76004E28E44FC4E98FA5D32BD1">Fisher, Brian L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="7CBA0A89A5D8FE6EA8FE8837C8946899">text</mods:typeOfResource>
<mods:relatedItem id="DB545E709E6601B2FF2AFA7591E0131A" type="host">
<mods:titleInfo id="76E909457C41FB25620A721A70A3CAFE">
<mods:title id="0668DBF560F322E7E68C1D23AC066DB6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D847DFDF8F059715EB48823FB2F37BF7">
<mods:date id="158D874F8AE771EB9729BE5B2D7DED4E">2012</mods:date>
<mods:detail id="76AFE1AB95C91F54AF45CA0D6A6D3879" type="pubDate">
<mods:number id="EFF04650461431AF9C712810105DBB7E">2012-07-04</mods:number>
</mods:detail>
<mods:detail id="0A63875DE2BE5DDE5C1AA024AF2E5AA9" type="volume">
<mods:number id="75C8E3F1EFB876F52370F3F8EBC421EF">3365</mods:number>
</mods:detail>
<mods:extent id="DEDC9F9E0B2E3A1AAEEBCE06854FF05C" unit="page">
<mods:start id="3CDE5B13D4517B5C93AF847E608DB980">1</mods:start>
<mods:end id="9851083F15E4CCAC6B65AC9D1ACCB51B">123</mods:end>
</mods:extent>
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<mods:classification id="C44E9A6BD7D1082C5E84E51E639CE966">journal article</mods:classification>
<mods:identifier id="A2EA61164C663F816426B0A18175D8B3" type="CLB-Dataset">20762</mods:identifier>
<mods:identifier id="5477528AC71D71E889DCB321F2BDBF5F" type="DOI">10.11646/zootaxa.3365.1.1</mods:identifier>
<mods:identifier id="20B2697A0BFB1326B92627EE11FBBB39" type="GBIF-Dataset">1db9bc74-46dd-4263-9784-88a59bd05d59</mods:identifier>
<mods:identifier id="00219F134FE8DDA9BB78EFBD011969DD" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="6B31F9673395A261B10328B5E50C07A5" type="Zenodo-Dep">6038900</mods:identifier>
</mods:mods>
<treatment id="03EF6217BF32FFCC0AC0F9DD980EAAF0" ID-DOI="http://doi.org/10.5281/zenodo.5253624" ID-GBIF-Taxon="193281054" ID-Zenodo-Dep="5253624" LSID="urn:lsid:plazi:treatment:03EF6217BF32FFCC0AC0F9DD980EAAF0" httpUri="http://treatment.plazi.org/id/03EF6217BF32FFCC0AC0F9DD980EAAF0" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12">
<subSubSection id="C35C808ABF32FFC90AC0F9DD9EA9AD18" box="[151,754,1621,1647]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BF9D301BF32FFC90AC0F9DD9EA9AD18" blockId="11.[151,754,1621,1680]" box="[151,754,1621,1647]" pageId="11" pageNumber="12">
<heading id="D0B1646DBF32FFC90AC0F9DD9EA9AD18" bold="true" box="[151,754,1621,1647]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="B9320F13BF32FFC90AC0F9DD9EA9AD18" bold="true" box="[151,754,1621,1647]" pageId="11" pageNumber="12">
<taxonomicName id="4C46A882BF32FFC90AC0F9DD9EF3AD18" authority="Hita Garcia &amp; Fisher" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[151,680,1621,1647]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="artemis" status="sp. nov.">
<emphasis id="B9320F13BF32FFC90AC0F9DD9DCCAD18" bold="true" box="[151,407,1621,1647]" italics="true" pageId="11" pageNumber="12">Tetramorium artemis</emphasis>
Hita Garcia &amp; Fisher
</taxonomicName>
<taxonomicNameLabel id="A201B268BF32FFC908E7F9DD9EA9AD18" box="[688,754,1621,1647]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C808ABF32FFC90AC0F9F09DDAADE7" box="[151,385,1656,1680]" pageId="11" pageNumber="12" type="description">
<paragraph id="8BF9D301BF32FFC90AC0F9F09DDAADE7" blockId="11.[151,754,1621,1680]" box="[151,385,1656,1680]" pageId="11" pageNumber="12">(figs 1, 3, 11, 12, 13)</paragraph>
</subSubSection>
<subSubSection id="C35C808ABF32FFC90AC0F948982FAC1F" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8BF9D301BF32FFC90AC0F948982FAC1F" blockId="11.[151,1436,1728,1896]" pageId="11" pageNumber="12">
<materialsCitation id="3B2ED95CBF32FFC90AC0F9489FFFAC57" ID-GBIF-Occurrence="3462706109" collectingDate="2002-02-13" collectingDateMax="2002-02-19" collectingDateMin="2002-02-13" collectionCode="CASC" collectorName="Fisher, B. L. &amp; C. Griswold" country="Madagascar" elevation="160" latitude="-25.59444" location="Reserve Speciale de Cap Sainte Marie" longLatPrecision="1" longitude="45.14683" pageId="11" pageNumber="12" specimenCode="CASENT0481732" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
<typeStatus id="54FD6DA3BF32FFC90AC0F9489CA4ADAF" box="[151,255,1728,1752]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
worker,
<collectingCountry id="F3519391BF32FFC90B31F9489E6BADAF" box="[358,560,1728,1752]" name="Madagascar" pageId="11" pageNumber="12">MADAGASCAR</collectingCountry>
,
<collectingRegion id="49821DE3BF32FFC9086AF9489ED4ADAF" box="[573,655,1728,1752]" country="Madagascar" name="Toliara" pageId="11" pageNumber="12">Toliara</collectingRegion>
,
<location id="8E9985DABF32FFC908CBF948980FADAF" LSID="urn:lsid:plazi:treatment:03EF6217BF32FFCC0AC0F9DD980EAAF0:8E9985DABF32FFC908CBF948980FADAF" box="[668,1108,1728,1752]" country="Madagascar" latitude="-25.59444" longLatPrecision="1" longitude="45.14683" name="Reserve Speciale de Cap Sainte Marie" pageId="11" pageNumber="12" stateProvince="Toliara">Réserve Spéciale de Cap Sainte Marie</location>
,
<locationDeviation id="9A4BCA23BF32FFC90E36F94899C3ADAF" box="[1121,1432,1728,1752]" location="Marovato" pageId="11" pageNumber="12">
<quantity id="4CBE7EE4BF32FFC90E36F94898E4ADAF" box="[1121,1215,1728,1752]" metricMagnitude="4" metricUnit="m" metricValue="1.49" pageId="11" pageNumber="12" unit="km" value="14.9">14.9 km</quantity>
261° W Marovato
</locationDeviation>
,
<geoCoordinate id="EE72B5C6BF32FFC90AC0F96C9D45AD8B" box="[151,286,1764,1788]" degrees="25.59444" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="1" value="-25.59444">25.59444 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF32FFC90B65F96C9DE0AD8C" box="[306,443,1764,1788]" degrees="45.14683" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="1" value="45.14683">45.14683 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF32FFC90B99F96C9E46AD8C" box="[462,541,1764,1788]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="12" unit="m" value="160.0">
<elevation id="006B3432BF32FFC90B99F96C9E46AD8C" box="[462,541,1764,1788]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="12" unit="m" value="160.0">160 m</elevation>
</quantity>
, spiny forest/thicket, sifted litter (leaf mold, rotten wood), BLF5570,
<date id="FFF8F5C1BF32FFC90F3CF96C9D50AC57" pageId="11" pageNumber="12" value="2002-02-13" valueMax="2002-02-19" valueMin="2002-02-13">
<collectingDate id="EFBC0C29BF32FFC90F3CF96C9D50AC57" pageId="11" pageNumber="12" value="2002-02-13" valueMax="2002-02-19" valueMin="2002-02-13">13.– 19.II.2002</collectingDate>
</date>
(
<emphasis id="B9320F13BF32FFC90B49F8809E0EAC57" box="[286,597,1800,1824]" italics="true" pageId="11" pageNumber="12">
<collectorName id="26B3B6D7BF32FFC90B49F8809DF8AC57" box="[286,419,1800,1824]" pageId="11" pageNumber="12">Fisher, B.L.</collectorName>
&amp;
<collectorName id="26B3B6D7BF32FFC90B9CF8809E0EAC57" box="[459,597,1800,1824]" pageId="11" pageNumber="12">C. Griswold</collectorName>
</emphasis>
) (
<collectionCode id="ED574BC4BF32FFC90826F8809E9BAC57" box="[625,704,1800,1824]" pageId="11" pageNumber="12">CASC</collectionCode>
:
<specimenCode id="DBE07B7ABF32FFC9089BF8809FC6AC57" box="[716,925,1800,1824]" collectionCode="CASC" pageId="11" pageNumber="12">CASENT0481732</specimenCode>
)
</materialsCitation>
<materialsCitation id="3B2ED95CBF32FFC909FAF8819E3DAC33" ID-GBIF-Occurrence="3462705893" collectingDate="2002-02-13" collectingDateMax="2002-02-19" collectingDateMin="2002-02-13" collectionCode="BMNH" collectorName="Fisher, B. L." country="Madagascar" elevation="160" latitude="-25.59444" location="Reserve Speciale de Cap Sainte Marie" longLatPrecision="1" longitude="45.14683" pageId="11" pageNumber="12" specimenCode="CASENT0481734" specimenCount="1" specimenCount-worker="1" stateProvince="Toliara" typeStatus="paratype">
<typeStatus id="54FD6DA3BF32FFC909FAF8819846AC56" box="[941,1053,1801,1825]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="9D401888BF32FFC90E7CF880989EAC57" box="[1067,1221,1800,1824]" pageId="11" pageNumber="12" type="worker">eight workers</specimenCount>
with same data as holotype (
<collectionCode id="ED574BC4BF32FFC90B4EF8A49D28AC33" box="[281,371,1836,1860]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="11" pageNumber="12">BMNH</collectionCode>
:
<specimenCode id="DBE07B7ABF32FFC90BD8F8A49E39AC33" box="[399,610,1836,1860]" collectionCode="BMNH" pageId="11" pageNumber="12">CASENT0481734</specimenCode>
;
</materialsCitation>
<materialsCitation id="3B2ED95CBF32FFC90829F8A49FE6AC33" ID-GBIF-Occurrence="3462705588" box="[638,957,1836,1860]" collectingDate="2002-02-13" collectingDateMax="2002-02-19" collectingDateMin="2002-02-13" collectionCode="CASC" collectorName="Fisher, B. L." country="Madagascar" elevation="160" latitude="-25.59444" location="Reserve Speciale de Cap Sainte Marie" longLatPrecision="1" longitude="45.14683" pageId="11" pageNumber="12" specimenCode="CASENT0481723" specimenCount="1" specimenCount-worker="1" stateProvince="Toliara" typeStatus="paratype">
<collectionCode id="ED574BC4BF32FFC90829F8A49E90AC33" box="[638,715,1836,1860]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="11" pageNumber="12">CASC</collectionCode>
:
<specimenCode id="DBE07B7ABF32FFC908BFF8A49FE2AC33" box="[744,953,1836,1860]" collectionCode="CASC" pageId="11" pageNumber="12">CASENT0481723</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B2ED95CBF32FFC90981F8A498F6AC33" ID-GBIF-Occurrence="3462705732" box="[982,1197,1836,1860]" collectingDate="2002-02-13" collectingDateMax="2002-02-19" collectingDateMin="2002-02-13" collectionCode="BMNH, CASC, MHNG" collectorName="Fisher, B. L." country="Madagascar" elevation="160" latitude="-25.59444" location="Reserve Speciale de Cap Sainte Marie" longLatPrecision="1" longitude="45.14683" pageId="11" pageNumber="12" specimenCode="CASENT0481729" specimenCount="1" specimenCount-worker="1" stateProvince="Toliara" typeStatus="paratype">
<specimenCode id="DBE07B7ABF32FFC90981F8A498F2AC33" box="[982,1193,1836,1860]" collectionCode="CASC" pageId="11" pageNumber="12">CASENT0481729</specimenCode>
;
</materialsCitation>
<materialsCitation id="3B2ED95CBF32FFC90E92F8A499C7AC33" ID-GBIF-Occurrence="3462706206" box="[1221,1436,1836,1860]" collectingDate="2002-02-13" collectingDateMax="2002-02-19" collectingDateMin="2002-02-13" collectionCode="BMNH, CASC, MHNG" collectorName="Fisher, B. L." country="Madagascar" elevation="160" latitude="-25.59444" location="Reserve Speciale de Cap Sainte Marie" longLatPrecision="1" longitude="45.14683" pageId="11" pageNumber="12" specimenCode="CASENT0481734, CASENT0481723, CASENT0481729, CASENT0481743" specimenCount="1" specimenCount-worker="1" stateProvince="Toliara" typeStatus="paratype">CASENT0481731;</materialsCitation>
<materialsCitation id="3B2ED95CBF32FFC90AC0F8D89D36AC1F" ID-GBIF-Occurrence="3462706082" box="[151,365,1872,1896]" collectingDate="2002-02-13" collectingDateMax="2002-02-19" collectingDateMin="2002-02-13" collectionCode="BMNH, CASC, MHNG" collectorName="Fisher, B. L." country="Madagascar" elevation="160" latitude="-25.59444" location="Reserve Speciale de Cap Sainte Marie" longLatPrecision="1" longitude="45.14683" pageId="11" pageNumber="12" specimenCode="CASENT0481734, CASENT0481723, CASENT0481729, CASENT0481743" specimenCount="1" specimenCount-worker="1" stateProvince="Toliara" typeStatus="paratype">CASENT0481750;</materialsCitation>
<materialsCitation id="3B2ED95CBF32FFC90B22F8D89E10AC1F" ID-GBIF-Occurrence="3462705589" box="[373,587,1872,1896]" collectingDate="2002-02-13" collectingDateMax="2002-02-19" collectingDateMin="2002-02-13" collectionCode="BMNH, CASC, MHNG" collectorName="Fisher, B. L." country="Madagascar" elevation="160" latitude="-25.59444" location="Reserve Speciale de Cap Sainte Marie" longLatPrecision="1" longitude="45.14683" pageId="11" pageNumber="12" specimenCode="CASENT0481734, CASENT0481723, CASENT0481729, CASENT0481743" specimenCount="1" specimenCount-worker="1" stateProvince="Toliara" typeStatus="paratype">CASENT0481752;</materialsCitation>
<materialsCitation id="3B2ED95CBF32FFC90805F8D89F72AC1F" ID-GBIF-Occurrence="3462706139" box="[594,809,1872,1896]" collectingDate="2002-02-13" collectingDateMax="2002-02-19" collectingDateMin="2002-02-13" collectionCode="BMNH, CASC, MHNG" collectorName="Fisher, B. L." country="Madagascar" elevation="160" latitude="-25.59444" location="Reserve Speciale de Cap Sainte Marie" longLatPrecision="1" longitude="45.14683" pageId="11" pageNumber="12" specimenCode="CASENT0481734, CASENT0481723, CASENT0481729, CASENT0481743" specimenCount="1" specimenCount-worker="1" stateProvince="Toliara" typeStatus="paratype">CASENT0481756;</materialsCitation>
<materialsCitation id="3B2ED95CBF32FFC90967F8D8982BAC1F" ID-GBIF-Occurrence="3462705923" box="[816,1136,1872,1896]" collectingDate="2002-02-13" collectingDateMax="2002-02-19" collectingDateMin="2002-02-13" collectionCode="MHNG" collectorName="Fisher, B. L." country="Madagascar" elevation="160" latitude="-25.59444" location="Reserve Speciale de Cap Sainte Marie" longLatPrecision="1" longitude="45.14683" pageId="11" pageNumber="12" specimenCode="CASENT0481743" specimenCount="1" specimenCount-worker="1" stateProvince="Toliara" typeStatus="paratype">
<collectionCode id="ED574BC4BF32FFC90967F8D89FD1AC1F" box="[816,906,1872,1896]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="11" pageNumber="12" type="Museum">MHNG</collectionCode>
:
<specimenCode id="DBE07B7ABF32FFC909C3F8D89833AC1F" box="[916,1128,1872,1896]" collectionCode="MHNG" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="11" pageNumber="12" type="Museum">CASENT0481743</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSection id="E2C9CAEABF32FFCE0AC0F81F9DE3AA6B" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="multiple">
<subSubSection id="C35C808ABF32FFCE0AC0F81F9DE3AA6B" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="8BF9D301BF32FFC90AC0F81F9D51ACC7" blockId="11.[151,1435,1943,2005]" box="[151,266,1943,1968]" pageId="11" pageNumber="12">
<heading id="D0B1646DBF32FFC90AC0F81F9D51ACC7" bold="true" box="[151,266,1943,1968]" fontSize="10" level="2" pageId="11" pageNumber="12" reason="2">
<emphasis id="B9320F13BF32FFC90AC0F81F9D51ACC7" bold="true" box="[151,266,1943,1968]" pageId="11" pageNumber="12">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF32FFCE0AC0F8349DE3AA6B" blockId="11.[151,1435,1943,2005]" lastBlockId="12.[151,1436,151,284]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
The following character combination separates
<taxonomicName id="4C46A882BF32FFC908FAF8359F46ACA3" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[685,797,1981,2004]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="artemis">
<emphasis id="B9320F13BF32FFC908FAF8359F46ACA3" box="[685,797,1981,2004]" italics="true" pageId="11" pageNumber="12">T. artemis</emphasis>
</taxonomicName>
from the other species of the group: head approximately as long as wide (CI 99–101); petiolar node squamiform and strongly anteroposteriorly compressed with anterodorsal angle situated higher than posterodorsal, dorsum tapering backwards posteriorly, in dorsal view strongly transverse (DPeI 232–250); and reduced cephalic sculpture between the frontal carinae with just one median longitudinal ruga.
</paragraph>
</subSubSection>
</subSection>
<subSubSection id="C35C808ABF35FFCE0AC0FEC39FCAAD5C" pageId="12" pageNumber="13" type="description">
<subSection id="E2C9CAEABF35FFCE0AC0FEC39EE9AEEB" pageId="12" pageNumber="13" type="multiple">
<paragraph id="8BF9D301BF35FFCE0AC0FEC39D7AAA13" blockId="12.[151,1437,331,1436]" box="[151,289,331,356]" pageId="12" pageNumber="13">
<heading id="D0B1646DBF35FFCE0AC0FEC39D7AAA13" bold="true" box="[151,289,331,356]" fontSize="10" level="2" pageId="12" pageNumber="13" reason="2">
<emphasis id="B9320F13BF35FFCE0AC0FEC39D7AAA13" bold="true" box="[151,289,331,356]" pageId="12" pageNumber="13">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF35FFCE0AC0FEF8992BA960" blockId="12.[151,1437,331,1436]" pageId="12" pageNumber="13">
HL 0.67–0.72 (0.69); HW 0.66–0.71 (0.69); SL 0.47–0.51 (0.49); EL 0.14–0.16 (0.15);
<collectingCountry id="F3519391BF35FFCE0E2CFEF898C4AAF0" box="[1147,1183,368,391]" name="Philippines" pageId="12" pageNumber="13">PH</collectingCountry>
0.37–0.43 (0.40); PW 0.48–0.52 (0.50); WL 0.82–0.89 (0.86); PSL 0.18–0.20 (0.19); PTL 0.10–0.11 (0.10); PTH 0.27–0.31 (0.29); PTW 0.24–0.26 (0.24); PPL 0.21–0.23 (0.22); PPH 0.27–0.32 (0.29); PPW 0.30–0.32 (0.31); CI 99–101 (100); SI 69–73 (71); OI 20–23 (21); LMI 44–48 (46); DMI 58–61 (59); PSLI 27–29 (28); PeNI 47–50 (48); LPeI 32–38 (35); DPeI 232–250 (240); PpNI 60–63 (62); LPpI 70–79 (75); DPpI 136–143 (140); PPI 122–131 (127) (nine measured).
</paragraph>
<paragraph id="8BF9D301BF35FFCE0A90FDAC9EE9AEEB" blockId="12.[151,1437,331,1436]" pageId="12" pageNumber="13">Head approximately as long as wide (CI 99–101). Anterior clypeal margin with distinct median impression. Frontal carinae well-developed, ending shortly behind level of posterior eye margin. Antennal scrobes faint to absent, posterior and ventral margins never differentiated. Antennal scapes comparatively short, not reaching posterior head margin (SI 69–73). Eyes comparatively small to moderate (OI 20–23). Mesosomal outline in profile convex, dorsum transversely rounded, dorsolateral margin only weakly developed, promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 44–48). Propodeal spines elongatetriangular, long, and acute (PSLI 27–29). Propodeal lobes reduced, very small, and bluntly triangular. Petiolar node strongly squamiform and anteroposteriorly compressed, anterior and posterior faces not parallel, anterodorsal margin higher situated and better developed than weaker posterodorsal margin, dorsum tapering backwards posteriorly; node in dorsal view transverse, between 2.3 to 3.5 times wider than long (DPeI 232–250), in lateral view distinctly more than 2.5 times higher than long (LPeI 32–38). Postpetiole in profile rounded and weakly anteroposteriorly compressed, approximately 1.3 to 1.4 times higher than long (LPpI 70–79), in dorsal view approximately 1.3 to 1.4 times wider than long (DPpI 136–143). Postpetiole in profile more voluminous than petiolar node, in dorsal view approximately 1.2 to 1.3 times wider than petiolar node (PPI 122–131). Mandibles distinctly longitudinally striate; clypeus always with strong median longitudinal ruga and one weaker ruga at each side; remainder of head mostly unsculptured, cephalic dorsum between frontal carinae only with one welldeveloped longitudinal median ruga, median ruga diverging approximately at eye level into two rugae running to posterior clypeal margin, remainder unsculptured, median ruga of same length as frontal carinae; head laterally and ventrally mostly unsculptured, anteriorly approximately at malar area irregularly rugose. Head with faint punctate ground sculpture. Mesosoma, waist segments, and gaster unsculptured and smooth, only moderately shining due to faint ground sculpture. Head generally with several fine, long, erect hairs, and more numerous appressed to decumbent widely spaced pubescence; mesosoma usually with one pair of standing hairs on promesonotum and very sparse, short, and inconspicuous pubescence; waist segments and first gastral tergite without any pilosity, only with very short, widely spaced, strongly appressed pubescence. Head, antennae, legs, and gaster of brown colour, mesosoma and waist segments of lighter brown.</paragraph>
</subSection>
<subSection id="E2C9CAEABF35FFCE0AC0FA449FCAAD5C" pageId="12" pageNumber="13" type="multiple">
<paragraph id="8BF9D301BF35FFCE0AC0FA449C81AE93" blockId="12.[151,1438,1484,2012]" box="[151,218,1484,1508]" pageId="12" pageNumber="13">
<heading id="D0B1646DBF35FFCE0AC0FA449C81AE93" bold="true" box="[151,218,1484,1508]" fontSize="10" level="2" pageId="12" pageNumber="13" reason="2">
<emphasis id="B9320F13BF35FFCE0AC0FA449C81AE93" bold="true" box="[151,218,1484,1508]" pageId="12" pageNumber="13">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF35FFCE0AC0FA679FCAAD5C" blockId="12.[151,1438,1484,2012]" pageId="12" pageNumber="13">
At present, the new species is only known from the
<typeStatus id="54FD6DA3BF35FFCE08B6FA789F4AAD7F" box="[737,785,1520,1544]" pageId="12" pageNumber="13">type</typeStatus>
locality, the Cap Saint Marie, where it was sampled from sifted litter in a spiny forest/thicket habitat at an elevation of
<specimenCount id="9D401888BF35FFCE0914F99C9FCAAD5C" box="[835,913,1556,1580]" pageId="12" pageNumber="13" type="generic">
<quantity id="4CBE7EE4BF35FFCE0914F99C9FD6AD5C" box="[835,909,1556,1580]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="13" unit="m" value="160.0">160 m</quantity>
.
</specimenCount>
</paragraph>
</subSection>
</subSubSection>
<subSubSection id="C35C808ABF35FFCE0A90F9BF983CACAB" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8BF9D301BF35FFCE0A90F9BF983CACAB" blockId="12.[151,1438,1484,2012]" pageId="12" pageNumber="13">
Within the species group, the identification of
<taxonomicName id="4C46A882BF35FFCE0899F9B09FE1AD38" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[718,954,1592,1615]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="artemis">
<emphasis id="B9320F13BF35FFCE0899F9B09FE1AD38" box="[718,954,1592,1615]" italics="true" pageId="12" pageNumber="13">Tetramorium artemis</emphasis>
</taxonomicName>
is straightforward and comparatively easy. It can be well separated from
<taxonomicName id="4C46A882BF35FFCE0BB3F9D59E32AD03" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[484,617,1628,1652]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="malagasy">
<emphasis id="B9320F13BF35FFCE0BB3F9D59E32AD03" box="[484,617,1628,1652]" italics="true" pageId="12" pageNumber="13">T. malagasy</emphasis>
</taxonomicName>
,
<taxonomicName id="4C46A882BF35FFCE0820F9D59EA3AD03" authorityName="Forel" authorityYear="1895" box="[631,760,1628,1652]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="orientale">
<emphasis id="B9320F13BF35FFCE0820F9D59EA3AD03" box="[631,760,1628,1652]" italics="true" pageId="12" pageNumber="13">T. orientale</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C46A882BF35FFCE0962F9D59FB0AD03" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[821,1003,1628,1652]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ryanphelanae">
<emphasis id="B9320F13BF35FFCE0962F9D59FB0AD03" box="[821,1003,1628,1652]" italics="true" pageId="12" pageNumber="13">T. ryanphelanae</emphasis>
</taxonomicName>
since the latter three all display a high nodiform and much less anteroposteriorly compressed petiolar node in comparison to the squamiform node of
<taxonomicName id="4C46A882BF35FFCE0FD0F9099CB0ADCB" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="artemis">
<emphasis id="B9320F13BF35FFCE0FD0F9099CB0ADCB" italics="true" pageId="12" pageNumber="13">T. artemis</emphasis>
</taxonomicName>
. The remaining two species,
<taxonomicName id="4C46A882BF35FFCE081DF92D9E9DADCB" authorityName="Forel" authorityYear="1891" box="[586,710,1700,1724]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bessonii">
<emphasis id="B9320F13BF35FFCE081DF92D9E9DADCB" box="[586,710,1700,1724]" italics="true" pageId="12" pageNumber="13">T. bessonii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C46A882BF35FFCE0952F92D9F3EADCB" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[773,869,1700,1724]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="wardi">
<emphasis id="B9320F13BF35FFCE0952F92D9F3EADCB" box="[773,869,1700,1724]" italics="true" pageId="12" pageNumber="13">T. wardi</emphasis>
</taxonomicName>
, have approximately the same node shape as
<taxonomicName id="4C46A882BF35FFCE0FD0F92D9CB0ADA8" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="artemis">
<emphasis id="B9320F13BF35FFCE0FD0F92D9CB0ADA8" italics="true" pageId="12" pageNumber="13">T. artemis</emphasis>
</taxonomicName>
, but can be easily distinguished from the latter. The head of
<taxonomicName id="4C46A882BF35FFCE09F5F9419840AD97" authorityName="Forel" authorityYear="1891" box="[930,1051,1736,1760]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bessonii">
<emphasis id="B9320F13BF35FFCE09F5F9419840AD97" box="[930,1051,1736,1760]" italics="true" pageId="12" pageNumber="13">T. bessonii</emphasis>
</taxonomicName>
is distinctly longer than wide (CI 92–96) while the head of
<taxonomicName id="4C46A882BF35FFCE0B91F9659E63AC73" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[454,568,1773,1796]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="artemis">
<emphasis id="B9320F13BF35FFCE0B91F9659E63AC73" box="[454,568,1773,1796]" italics="true" pageId="12" pageNumber="13">T. artemis</emphasis>
</taxonomicName>
is approximately as long as wide (99–101). In addition,
<taxonomicName id="4C46A882BF35FFCE0E86F9659917AC73" authorityName="Forel" authorityYear="1891" box="[1233,1356,1772,1796]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bessonii">
<emphasis id="B9320F13BF35FFCE0E86F9659917AC73" box="[1233,1356,1772,1796]" italics="true" pageId="12" pageNumber="13">T. bessonii</emphasis>
</taxonomicName>
shows comparatively long appressed to decumbent pubescence on the first gastral tergite, which contrasts with the very short, strongly appressed pubescence of
<taxonomicName id="4C46A882BF35FFCE080AF8BD9E95AC3B" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[605,718,1845,1868]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="artemis">
<emphasis id="B9320F13BF35FFCE080AF8BD9E95AC3B" box="[605,718,1845,1868]" italics="true" pageId="12" pageNumber="13">T. artemis</emphasis>
</taxonomicName>
.
<taxonomicName id="4C46A882BF35FFCE088BF8BD9FEFAC3B" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[732,948,1844,1868]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="wardi">
<emphasis id="B9320F13BF35FFCE088BF8BD9FEFAC3B" box="[732,948,1844,1868]" italics="true" pageId="12" pageNumber="13">Tetramorium wardi</emphasis>
</taxonomicName>
shares most characters with
<taxonomicName id="4C46A882BF35FFCE0EADF8BD9931AC3B" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[1274,1386,1845,1868]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="artemis">
<emphasis id="B9320F13BF35FFCE0EADF8BD9931AC3B" box="[1274,1386,1845,1868]" italics="true" pageId="12" pageNumber="13">T. artemis</emphasis>
</taxonomicName>
, but differs in cephalic sculpture and petiolar node shape. In
<taxonomicName id="4C46A882BF35FFCE095BF8D19F33AC07" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[780,872,1880,1904]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="wardi">
<emphasis id="B9320F13BF35FFCE095BF8D19F33AC07" box="[780,872,1880,1904]" italics="true" pageId="12" pageNumber="13">T. wardi</emphasis>
</taxonomicName>
the cephalic sculpture between the frontal carinae consists of a strong median ruga and several irregular rugae between the median ruga and the frontal carinae. In
<taxonomicName id="4C46A882BF35FFCE0FD0F8F59CB0ACCF" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="artemis">
<emphasis id="B9320F13BF35FFCE0FD0F8F59CB0ACCF" italics="true" pageId="12" pageNumber="13">T. artemis</emphasis>
</taxonomicName>
though, only one median ruga is developed between the frontal carinae. Also, the petiolar node of
<taxonomicName id="4C46A882BF35FFCE0F17F82999C7ACCF" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[1344,1436,1952,1976]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="wardi">
<emphasis id="B9320F13BF35FFCE0F17F82999C7ACCF" box="[1344,1436,1952,1976]" italics="true" pageId="12" pageNumber="13">T. wardi</emphasis>
</taxonomicName>
(DPeI 267–333) is more compressed and transverse than in
<taxonomicName id="4C46A882BF35FFCE0963F84D9FFFACAB" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[820,932,1989,2012]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="artemis">
<emphasis id="B9320F13BF35FFCE0963F84D9FFFACAB" box="[820,932,1989,2012]" italics="true" pageId="12" pageNumber="13">T. artemis</emphasis>
</taxonomicName>
(DPeI 232–250).
</paragraph>
</subSubSection>
<caption id="DF398389BF34FFCF0AC0F8319DFEAC85" ID-DOI="http://doi.org/10.5281/zenodo.6038918" ID-Zenodo-Dep="6038918" httpUri="https://zenodo.org/record/6038918/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,264,1977,2000]" targetBox="[151,1436,193,1968]" targetPageId="13">
<paragraph id="8BF9D301BF34FFCF0AC0F8319DFEAC85" blockId="13.[151,1437,1977,2034]" pageId="13" pageNumber="14">
<emphasis id="B9320F13BF34FFCF0AC0F8319D08ACA7" bold="true" box="[151,339,1977,2000]" pageId="13" pageNumber="14">FIGURES 11–13.</emphasis>
<taxonomicName id="4C46A882BF34FFCF0B0CF8329D9AACB8" authorityName="Hita Garcia &amp; Fisher" authorityYear="2012" box="[347,449,1978,1999]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="artemis">
<emphasis id="B9320F13BF34FFCF0B0CF8329D9AACB8" box="[347,449,1978,1999]" italics="true" pageId="13" pageNumber="14">T. artemis</emphasis>
</taxonomicName>
, holotype—CASENT0481732 (Erin Prado 2010). 11. Body in profile. 12. Body in dorsal view. 13. Head in full-face view.
</paragraph>
</caption>
<subSection id="E2C9CAEABF37FFCC0AC0FF109818AB80" pageId="14" pageNumber="15" type="multiple">
<subSubSection id="C35C808ABF37FFCC0AC0FF109818AB80" pageId="14" pageNumber="15" type="etymology">
<paragraph id="8BF9D301BF37FFCC0AC0FF109D43ABC7" blockId="14.[151,1436,152,247]" box="[151,280,152,176]" pageId="14" pageNumber="15">
<heading id="D0B1646DBF37FFCC0AC0FF109D43ABC7" bold="true" box="[151,280,152,176]" fontSize="10" level="2" pageId="14" pageNumber="15" reason="2">
<emphasis id="B9320F13BF37FFCC0AC0FF109D43ABC7" bold="true" box="[151,280,152,176]" pageId="14" pageNumber="15">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF37FFCC0AC0FF349818AB80" blockId="14.[151,1436,152,247]" pageId="14" pageNumber="15">This new species is named after the goddess &quot;Artemis&quot; from Old Greek mythology, which was the goddess of the hunt, wilderness, and animals. The name is a noun in apposition, and thus invariant.</paragraph>
</subSubSection>
</subSection>
<subSection id="E2C9CAEABF37FFCC0AC0FEAF980EAAF0" pageId="14" pageNumber="15" type="multiple">
<subSubSection id="C35C808ABF37FFCC0AC0FEAF980EAAF0" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BF9D301BF37FFCC0AC0FEAF9D22AA37" blockId="14.[151,1435,295,392]" box="[151,377,295,320]" pageId="14" pageNumber="15">
<heading id="D0B1646DBF37FFCC0AC0FEAF9D22AA37" bold="true" box="[151,377,295,320]" fontSize="10" level="2" pageId="14" pageNumber="15" reason="2">
<emphasis id="B9320F13BF37FFCC0AC0FEAF9D22AA37" bold="true" box="[151,377,295,320]" pageId="14" pageNumber="15">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF37FFCC0AC0FEC3980EAAF0" blockId="14.[151,1435,295,392]" pageId="14" pageNumber="15">
<materialsCitation id="3B2ED95CBF37FFCC0AC0FEC3980EAAF0" ID-GBIF-Occurrence="3462706292" pageId="14" pageNumber="15">
<emphasis id="B9320F13BF37FFCC0AC0FEC39D3FAA13" bold="true" box="[151,356,331,356]" pageId="14" pageNumber="15">
<collectingCountry id="F3519391BF37FFCC0AC0FEC39D3FAA13" box="[151,356,331,356]" name="Madagascar" pageId="14" pageNumber="15">MADAGASCAR</collectingCountry>
</emphasis>
:
<collectingRegion id="49821DE3BF37FFCC0B2EFEC49D90AA13" box="[377,459,332,356]" country="Madagascar" name="Toliara" pageId="14" pageNumber="15">Toliara</collectingRegion>
, Réserve Spéciale de Cap Sainte Marie,
<quantity id="4CBE7EE4BF37FFCC09EFFEC49842AA13" box="[952,1049,332,356]" metricMagnitude="4" metricUnit="m" metricValue="1.49" pageId="14" pageNumber="15" unit="km" value="14.9">14.9 km</quantity>
261° W Marovato,
<geoCoordinate id="EE72B5C6BF37FFCC0F47FEC499CDAA13" box="[1296,1430,332,356]" degrees="25.59444" direction="south" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="-25.59444">25.59444 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF37FFCC0AC0FEE79D41AAF0" box="[151,282,367,391]" degrees="45.14683" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="45.14683">45.14683 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF37FFCC0B71FEE79D34AAF0" box="[294,367,367,391]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="15" unit="m" value="160.0">160 m</quantity>
, spiny forest/thicket,
<date id="FFF8F5C1BF37FFCC0837FEE79F5EAAF0" box="[608,773,367,391]" pageId="14" pageNumber="15" value="2002-02-13" valueMax="2002-02-19" valueMin="2002-02-13">13.–19.II.2002</date>
(
<emphasis id="B9320F13BF37FFCC0941FEF8981EAAFF" box="[790,1093,368,392]" italics="true" pageId="14" pageNumber="15">Fisher, B.L. &amp; C. Griswold</emphasis>
).
</materialsCitation>
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>
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