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<document id="544F1AB2013479B0AB4920427B7A211D" ID-DOI="10.11646/phytotaxa.388.2.5" ID-ISSN="1179-3163" ID-Zenodo-Dep="13725355" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725673446816" checkinUser="felipe" docAuthor="Doweld, Alexander B." docDate="2019" docId="03EA87B7095BFFE26BA9647DFC0C499F" docLanguage="en" docName="phytotaxa.388.2.5.pdf" docOrigin="Phytotaxa 388 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.388.2.5" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Ilex subrotunda Doweld 2019, sp. nov." docType="treatment" docVersion="2" lastPageNumber="184" masterDocId="FFD3FFCF095EFFE76B21675BFF8F4D70" masterDocTitle="New names for Ilex and Ilexpollenites (Aquifoliaceae), extant and fossil. Notulae Systematicae ad Palaeofloram Europaeam spectantes II. Aquifoliaceae" masterLastPageNumber="191" masterPageNumber="179" pageNumber="184" updateTime="1725684925477" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="910401AE328DD2B3926DF622D8C07291">New names for Ilex and Ilexpollenites (Aquifoliaceae), extant and fossil. Notulae Systematicae ad Palaeofloram Europaeam spectantes II. Aquifoliaceae</mods:title>
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<mods:namePart id="DF137460AA7A6755526A6EF7783FF133">Doweld, Alexander B.</mods:namePart>
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<mods:title id="B6FC7183711A5BF4292ED354F4E1CE14">Phytotaxa</mods:title>
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<mods:date id="0478020D7DAB17E8D080245488CF0856">2019</mods:date>
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<mods:identifier id="8B97D519F425121D35FBE650EC875B4D" type="DOI">10.11646/phytotaxa.388.2.5</mods:identifier>
<mods:identifier id="E5664B2582B98DA2CD225FFC5F39D74F" type="ISSN">1179-3163</mods:identifier>
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<treatment id="03EA87B7095BFFE26BA9647DFC0C499F" LSID="urn:lsid:plazi:treatment:03EA87B7095BFFE26BA9647DFC0C499F" httpUri="http://treatment.plazi.org/id/03EA87B7095BFFE26BA9647DFC0C499F" lastPageNumber="184" pageId="5" pageNumber="184">
<subSubSection id="C359652A095BFFE26BA9647DFD8C4E31" box="[136,515,806,833]" pageId="5" pageNumber="184" type="nomenclature">
<paragraph id="8BFC36A1095BFFE26BA9647DFD8C4E31" blockId="5.[136,515,806,833]" box="[136,515,806,833]" pageId="5" pageNumber="184">
<heading id="D0B481CD095BFFE26BA9647DFD8C4E31" box="[136,515,806,833]" fontSize="11" level="2" pageId="5" pageNumber="184" reason="3">
<taxonomicName id="4C434D22095BFFE26BA9647DFE2D4E31" authority="Doweld" authorityName="Doweld" authorityYear="2019" box="[136,418,806,833]" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="5" pageNumber="184" phylum="Tracheophyta" rank="species" species="subrotunda" status="sp. nov.">
<emphasis id="B937EAB3095BFFE26BA9647DFECF4E31" bold="true" box="[136,320,806,833]" italics="true" pageId="5" pageNumber="184">Ilex subrotunda</emphasis>
Doweld
</taxonomicName>
,
<emphasis id="B937EAB3095BFFE26A8F6473FD8C4E31" box="[430,515,808,833]" italics="true" pageId="5" pageNumber="184">
<taxonomicNameLabel id="A20457C8095BFFE26A8F6473FD8C4E31" box="[430,515,808,833]" pageId="5" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C359652A095BFFE26BA96415FCBE4E13" box="[136,817,846,867]" pageId="5" pageNumber="184" type="reference_group">
<paragraph id="8BFC36A1095BFFE26BA96415FCBE4E13" blockId="5.[136,1452,846,1263]" box="[136,817,846,867]" pageId="5" pageNumber="184">
<treatmentCitationGroup id="AB53118F095BFFE26BA96415FCBE4E13" box="[136,817,846,867]" pageId="5" pageNumber="184">
<treatmentCitation id="0AE210B0095BFFE26BBD6415FE534E13" author="Huzioka, K." box="[156,476,846,867]" page=")" pageId="5" pageNumber="184" year="1963">
<taxonomicName id="4C434D22095BFFE26BBD6415FE534E13" authority="Huzioka (1963: 208)" authorityName="Huzioka" authorityPageNumber="208" authorityYear="1963" box="[156,476,846,867]" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="5" pageNumber="184" phylum="Tracheophyta" rank="species" species="ohashii">
<emphasis id="B937EAB3095BFFE26BBD6415FE824E13" box="[156,269,846,867]" italics="true" pageId="5" pageNumber="184">Ilex ohashii</emphasis>
<bibRefCitation id="EFD24B50095BFFE26A336415FE534E13" author="Huzioka, K." box="[274,476,846,867]" pageId="5" pageNumber="184" pagination="153 - 216" refId="ref8915" refString="Huzioka, K. (1963) The Utto flora. In: Tertiary Floras of Japan. Miocene Floras. The Collaborating Association to Commemorate the 80 th Anniversary of the Geological Survey of Japan, [Tokyo], pp. 153 - 216." type="book chapter" year="1963">Huzioka (1963: 208)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<emphasis id="B937EAB3095BFFE26AC76415FDC34E13" box="[486,588,846,867]" italics="true" pageId="5" pageNumber="184">nom. inval</emphasis>
. (Art. 40.1 of the ICN).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C359652A095BFFE26BA96429FB014EDB" pageId="5" pageNumber="184" type="materials_examined">
<paragraph id="8BFC36A1095BFFE26BA96429FB014EDB" blockId="5.[136,1452,846,1263]" pageId="5" pageNumber="184">
<emphasis id="B937EAB3095BFFE26BA96429FF354EF7" bold="true" box="[136,186,882,903]" pageId="5" pageNumber="184">Type</emphasis>
:—[fossil leaves] Yamakayakusa, Ani-machi, Kita-Akita-gun
<collectingRegion id="4987F843095BFFE2682F6429FC3F4EF7" box="[782,944,882,903]" country="Japan" name="Akita" pageId="5" pageNumber="184">Akita Prefecture</collectingRegion>
,
<collectingCountry id="F3547631095BFFE2689A6429FC7D4EF7" box="[955,1010,882,903]" name="Japan" pageId="5" pageNumber="184">Japan</collectingCountry>
(
<typeStatus id="54F88803095BFFE268DE6429FBD84EF7" box="[1023,1111,882,903]" pageId="5" pageNumber="184" type="holotype">holotype</typeStatus>
, AKMG-3132, Institute of Applied Geology, Mining College,
<collectingRegion id="4987F843095BFFE26A9C64CDFE7D4EDB" box="[445,498,918,939]" country="Japan" name="Akita" pageId="5" pageNumber="184">Akita</collectingRegion>
University,
<collectingRegion id="4987F843095BFFE2694764CDFD114EDB" box="[614,670,918,939]" country="Japan" name="Akita" pageId="5" pageNumber="184">Akita</collectingRegion>
, Japan-illustrated by
<bibRefCitation id="EFD24B50095BFFE2684B64CDFC794EDB" author="Huzioka, K." box="[874,1014,918,939]" pageId="5" pageNumber="184" pagination="153 - 216" refId="ref8915" refString="Huzioka, K. (1963) The Utto flora. In: Tertiary Floras of Japan. Miocene Floras. The Collaborating Association to Commemorate the 80 th Anniversary of the Geological Survey of Japan, [Tokyo], pp. 153 - 216." type="book chapter" year="1963">Huzioka 1963</bibRefCitation>
: pl. 35, fig. 10).
</paragraph>
</subSubSection>
<subSubSection id="C359652A095BFFE26BA964E1FC0C499F" pageId="5" pageNumber="184" type="description">
<paragraph id="8BFC36A1095BFFE26BA964E1FC834967" blockId="5.[136,1452,846,1263]" pageId="5" pageNumber="184">
<emphasis id="B937EAB3095BFFE26BA964E1FF704EBF" bold="true" box="[136,255,954,975]" pageId="5" pageNumber="184">Description</emphasis>
:—Leaves elliptical,
<quantity id="4CBB9B44095BFFE26AEF64E1FDA44EBF" box="[462,555,954,975]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="5" pageNumber="184" unit="cm" value="4.75" valueMax="5.0" valueMin="4.5">4.55 cm</quantity>
long,
<quantity id="4CBB9B44095BFFE2694F64E1FD524EBF" box="[622,733,954,975]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.5" metricValueMin="1.9" pageId="5" pageNumber="184" unit="cm" value="2.2" valueMax="2.5" valueMin="1.9">1.92.5 cm</quantity>
wide, acute at both basis and apex, entire, long-petiolate (
<quantity id="4CBB9B44095BFFE26E0F64E1FAE64EBF" box="[1326,1385,954,975]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="184" unit="mm" value="6.0">6 mm</quantity>
long); venation of 7 pairs of secondary veinlets leave the midrib at angles of 4540, camptodrome, tertiary venation reticulate or percurrent in the inter-secondary spaces (after
<bibRefCitation id="EFD24B50095BFFE269326359FD134967" author="Huzioka, K." box="[531,668,1026,1047]" pageId="5" pageNumber="184" pagination="153 - 216" refId="ref8915" refString="Huzioka, K. (1963) The Utto flora. In: Tertiary Floras of Japan. Miocene Floras. The Collaborating Association to Commemorate the 80 th Anniversary of the Geological Survey of Japan, [Tokyo], pp. 153 - 216." type="book chapter" year="1963">Huzioka 1963</bibRefCitation>
, modified).
</paragraph>
<paragraph id="8BFC36A1095BFFE26BA9637DFC24494B" blockId="5.[136,1452,846,1263]" box="[136,939,1062,1083]" pageId="5" pageNumber="184">
<emphasis id="B937EAB3095BFFE26BA9637DFF70494B" bold="true" box="[136,255,1062,1083]" pageId="5" pageNumber="184">Occurrence</emphasis>
:—Middle Miocene (Utto Formation); Eurasia (Japanese Archipelago).
</paragraph>
<paragraph id="8BFC36A1095BFFE26BA96311FCE4492F" blockId="5.[136,1452,846,1263]" box="[136,875,1098,1119]" pageId="5" pageNumber="184">
<emphasis id="B937EAB3095BFFE26BA96311FF64492F" bold="true" box="[136,235,1098,1119]" pageId="5" pageNumber="184">Eponymy</emphasis>
:—Similar to extant
<taxonomicName id="4C434D22095BFFE26A8E6311FD08492F" authority="Thunberg" authorityName="Thunberg" box="[431,647,1098,1119]" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="5" pageNumber="184" phylum="Tracheophyta" rank="species" species="rotunda">
<emphasis id="B937EAB3095BFFE26A8E6311FDAB492F" box="[431,548,1098,1119]" italics="true" pageId="5" pageNumber="184">Ilex rotunda</emphasis>
Thunberg
</taxonomicName>
(in
<bibRefCitation id="EFD24B50095BFFE2698F6311FCD0492F" author="Murray, J. A." box="[686,863,1098,1119]" pageId="5" pageNumber="184" refId="ref9945" refString="Murray, J. A. (1784) Caroli a Linne equitis Systema vegetabilium secundum classes ordines genera species cum characteribus et differentiis. Editio decima quarta praecedente longe auctior et correctior curante Jo. Andrea Murray. Typis et impensis Jo. Christ. Dieterich, Gottingae, 1004 pp. https: // doi. org / 10.5962 / bhl. title. 549" type="book" year="1784">Murray 1784: 168</bibRefCitation>
).
</paragraph>
<paragraph id="8BFC36A1095BFFE26BA96335FD5949F3" blockId="5.[136,1452,846,1263]" box="[136,726,1134,1155]" pageId="5" pageNumber="184">
<emphasis id="B937EAB3095BFFE26BA96335FF4649F3" bold="true" box="[136,201,1134,1155]" pageId="5" pageNumber="184">IFPNI</emphasis>
:—
<uuid id="FFE50C74095BFFE26BC66335FD5D49F3" box="[231,722,1134,1155]" pageId="5" pageNumber="184">9DFB6C2F-7C4D-0AAB-DDD6-71C69F0A13C4</uuid>
.
</paragraph>
<paragraph id="8BFC36A1095BFFE26BA963C9FC0C499F" blockId="5.[136,1452,846,1263]" pageId="5" pageNumber="184">
<emphasis id="B937EAB3095BFFE26BA963C9FF4E49D7" bold="true" box="[136,193,1170,1191]" pageId="5" pageNumber="184">Notes</emphasis>
:—The fossil-species
<taxonomicName id="4C434D22095BFFE26AB463C9FD5549D7" authority="Huzioka (1963: 208)" authorityName="Huzioka" authorityPageNumber="208" authorityYear="1963" box="[405,730,1170,1191]" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="5" pageNumber="184" phylum="Tracheophyta" rank="species" species="ohashii">
<emphasis id="B937EAB3095BFFE26AB463C9FD8749D7" box="[405,520,1170,1191]" italics="true" pageId="5" pageNumber="184">Ilex ohashii</emphasis>
<bibRefCitation id="EFD24B50095BFFE2692E63C9FD5549D7" author="Huzioka, K." box="[527,730,1170,1191]" pageId="5" pageNumber="184" pagination="153 - 216" refId="ref8915" refString="Huzioka, K. (1963) The Utto flora. In: Tertiary Floras of Japan. Miocene Floras. The Collaborating Association to Commemorate the 80 th Anniversary of the Geological Survey of Japan, [Tokyo], pp. 153 - 216." type="book chapter" year="1963">Huzioka (1963: 208)</bibRefCitation>
</taxonomicName>
was not validly published since the type was not designated (Art. 40.1 of the ICN) [
<specimenCount id="9D45FD28095BFFE26A3F63EDFE2149BB" box="[286,430,1206,1227]" count="2" pageId="5" pageNumber="184" type="generic">two specimens</specimenCount>
from two locations were incorrectly cited as “
<typeStatus id="54F88803095BFFE2684963EDFC4149BB" box="[872,974,1206,1227]" pageId="5" pageNumber="184" type="holotype">holotypes</typeStatus>
”].
<bibRefCitation id="EFD24B50095BFFE268C763EDFBF249BB" author="Huzioka, K." box="[998,1149,1206,1227]" pageId="5" pageNumber="184" pagination="153 - 216" refId="ref8915" refString="Huzioka, K. (1963) The Utto flora. In: Tertiary Floras of Japan. Miocene Floras. The Collaborating Association to Commemorate the 80 th Anniversary of the Geological Survey of Japan, [Tokyo], pp. 153 - 216." type="book chapter" year="1963">Huzioka (1963)</bibRefCitation>
related it to extant
<taxonomicName id="4C434D22095BFFE26E1763EDFE93499F" authority="Thunberg" authorityName="Thunberg" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="5" pageNumber="184" phylum="Tracheophyta" rank="species" species="rotunda">
<emphasis id="B937EAB3095BFFE26E1763EDFA2449BB" box="[1334,1451,1206,1227]" italics="true" pageId="5" pageNumber="184">Ilex rotunda</emphasis>
Thunberg
</taxonomicName>
, therefore, a new name is proposed to replace invalid
<taxonomicName id="4C434D22095BFFE268036381FCF2499F" authorityName="Huzioka" authorityYear="1963" box="[802,893,1242,1263]" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="5" pageNumber="184" phylum="Tracheophyta" rank="species" species="ohashii">
<emphasis id="B937EAB3095BFFE268036381FCF2499F" box="[802,893,1242,1263]" italics="true" pageId="5" pageNumber="184">I. ohashii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>