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<mods:title id="081DE65041B97D26DF9DED657057E0FA">First records of some coral reef-associated brachyuran crabs from the Nicobar archipelago, India</mods:title>
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<mods:namePart id="8ACFE334A670AA1C4F28898360ADBCB6">Cubelio, Sherine Sonia</mods:namePart>
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<mods:namePart id="1A84C92742534343CBC925F93DD8C067">Takeda, Masatsune</mods:namePart>
<mods:affiliation id="4D2DD362249E12AAD827CC961604F4BE">Department of Zoology, National Museum of Nature and Science, Tokyo. 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan.</mods:affiliation>
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<taxonomicName id="4C434D047F24FF87FF79FB2FB776D3CD" ID-CoL="5C4TN" authority="Mendoza, 2013" authorityName="Mendoza" authorityYear="2013" box="[151,575,1230,1257]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="491" phylum="Arthropoda" rank="species" species="joanneae">
<emphasis id="B937EA957F24FF87FF79FB2FB776D3CD" bold="true" box="[151,575,1230,1257]" pageId="15" pageNumber="491">
<emphasis id="B937EA957F24FF87FF79FB2FB432D3CD" bold="true" box="[151,379,1230,1257]" italics="true" pageId="15" pageNumber="491">Xanthias joanneae</emphasis>
<bibRefCitation id="EFD24B767F24FF87FE6CFB2FB776D3CD" author="Mendoza, J. C. E." box="[386,575,1230,1256]" pageId="15" pageNumber="491" pagination="374 - 384" refId="ref12717" refString="Mendoza, J. C. E. (2013) A new species of ocellated Xanthias Rathbun, 1897 (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, Philippines. Zootaxa, 3636 (2), 374 - 384. https: // doi. org / 10.11646 / zootaxa. 3636.2.8" type="journal article" year="2013">Mendoza, 2013</bibRefCitation>
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<paragraph id="8BFC36877F24FF87FF79FB13B4FED229" blockId="15.[151,575,1230,1292]" box="[151,439,1266,1292]" pageId="15" pageNumber="491">
(
<figureCitation id="13782A027F24FF87FF71FB13B44AD229" box="[159,259,1266,1292]" captionStart="FIGURE 2" captionStartId="3.[152,255,1683,1707]" captionTargetBox="[177,1410,181,1659]" captionTargetId="figure-37@3.[177,1410,181,1659]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal habitus photographs (AC, EG: fresh specimens; D: ethanol-preserved specimen): (A) Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm); (B) Urnalana angulata (Rathbun, 1911), undetermined sex (IO/SS/BRC/00184, 6.9 × 7.0 mm); (C) Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm; (D) Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm); (E) Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm); (F) Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/SS/ BRC/00194, 5.2 × 3.6 mm); (G) Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). Scale bars: AB, E, G, 2 mm, C, 10 mm, D, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210274" httpUri="https://zenodo.org/record/13210274/files/figure.png" pageId="15" pageNumber="491">Figs. 2G</figureCitation>
,
<figureCitation id="13782A027F24FF87FEE1FB13B41DD229" box="[271,340,1266,1292]" captionStart="FIGURE 9" captionStartId="16.[152,255,1914,1938]" captionTargetBox="[281,1306,597,1890]" captionTargetId="figure-220@16.[281,1306,597,1890]" captionTargetPageId="16" captionText="FIGURE 9. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm), Andaman Sea, India: colour photographs: (A) carapace, dorsal view; (B) mxp3, chelipeds, pereopods, sternum and pleon, ventral view. Scale bars: AB, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210296" httpUri="https://zenodo.org/record/13210296/files/figure.png" pageId="15" pageNumber="491">9AB</figureCitation>
,
<figureCitation id="13782A027F24FF87FE8EFB13B4E6D229" box="[352,431,1266,1292]" captionStart="FIGURE 10" captionStartId="17.[151,250,1667,1691]" captionTargetBox="[304,1294,191,1633]" captionTargetId="figure-37@17.[292,1295,181,1645]" captionTargetPageId="17" captionText="FIGURE 10. First and second gonopods: Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm): (A) entire G1, pleonal view; (B) entire G1, sternal view; (C) entire G2, pleonal view. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm): (D) entire G1, pleonal view; (E) G1 distal portion, pleonal view; (F) G1 distal portion, sternal view; (G) entire G1, sternal view; (H) entire G2, pleonal view. Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm): (I) entire G1, pleonal view; (J) entire G1, lateral view; (K) entire G1, mesial view; (L) entire G2, pleonal view. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm): (M) entire G1, pleonal view; (N) entire G1, sternal view; (O) entire G2, sternal view. Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/ SS/BRC/00194, 5.2 × 3.6 mm): (P) entire G1, pleonal view; (Q) G1 distal portion, pleonal view; (R) entire G1, sternal view. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). (S) entire G1, pleonal view; (T) G1 distal portion, pleonal view; (U) G1 distal portion, sternal view; (V) entire G1, sternal view. Scale bars: AC, PV, 0.2 mm, D, GN, 1 mm, E, F, O, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210300" httpUri="https://zenodo.org/record/13210300/files/figure.png" pageId="15" pageNumber="491">10SV</figureCitation>
)
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<taxonomicName id="4C434D047F24FF87FF79FAD6B76BD26B" ID-CoL="5C4TN" authority="Mendoza 2013: 375" authorityName="Mendoza" authorityPageNumber="375" authorityYear="2013" box="[151,546,1334,1359]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="491" phylum="Arthropoda" rank="species" species="joanneae">
<emphasis id="B937EA957F24FF87FF79FAD6B41AD26B" box="[151,339,1335,1359]" italics="true" pageId="15" pageNumber="491">Xanthias joanneae</emphasis>
<bibRefCitation id="EFD24B767F24FF87FEB6FAD6B76BD26B" author="Mendoza, J. C. E." box="[344,546,1334,1359]" pageId="15" pageNumber="491" pagination="374 - 384" refId="ref12717" refString="Mendoza, J. C. E. (2013) A new species of ocellated Xanthias Rathbun, 1897 (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, Philippines. Zootaxa, 3636 (2), 374 - 384. https: // doi. org / 10.11646 / zootaxa. 3636.2.8" type="journal article" year="2013">Mendoza 2013: 375</bibRefCitation>
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, figs. 14 (
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locality: Panglao Island,
<collectingCountry id="F35476177F24FF87FC57FAD6B167D26A" box="[953,1070,1335,1359]" name="Philippines" pageId="15" pageNumber="491">Philippines</collectingCountry>
,
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depth).—
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<bibRefCitation id="EFD24B767F24FF87FB51FAD6B0D5D26B" author="Poupin, J. &amp; Cleva, R. &amp; Bouchard, J. M. &amp; Dinhut, V. &amp; Dumas, J." box="[1215,1436,1334,1359]" pageId="15" pageNumber="491" pagination="1 - 109" refId="ref13887" refString="Poupin, J., Cleva, R., Bouchard, J. M., Dinhut, V. &amp; Dumas, J. (2018) The crabs from Mayotte Island (Crustacea, Decapoda, Brachyura). Atoll Research Bulletin, 617, 1 - 109. https: // doi. org / 10.5479 / si. 0077 - 5630.617" type="journal article" year="2018">
Poupin
<emphasis id="B937EA957F24FF87FAE0FAD6B009D26A" box="[1294,1344,1335,1359]" italics="true" pageId="15" pageNumber="491">et al.</emphasis>
2018: 65
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<paragraph id="8BFC36877F24FF87FF29FAB4B1C7D2A9" blockId="15.[151,1436,1334,1420]" pageId="15" pageNumber="491">
(remarks), 82.—
<bibRefCitation id="EFD24B767F24FF87FE81FAB4B7EBD248" author="Takeda, M. &amp; Komatsu, H." box="[367,674,1365,1389]" pageId="15" pageNumber="491" pagination="153 - 189" refId="ref15381" refString="Takeda, M. &amp; Komatsu, H. (2018) Offshore Crabs of the Family Xanthidae and Some Related Families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Science Museum, Tokyo, 52, 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu 2018: 171</bibRefCitation>
, 172 (remarks), 183 (discussion).—Takeda
<emphasis id="B937EA957F24FF87FBB0FAB7B1D8D248" box="[1118,1169,1365,1389]" italics="true" pageId="15" pageNumber="491">et al.</emphasis>
2018: 121 (discussion).
<taxonomicName id="4C434D047F24FF87FF79FA95B429D2A9" ID-CoL="5C4TS" authorityName="Sakai" authorityYear="1961" box="[151,352,1396,1420]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="491" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B937EA957F24FF87FF79FA95B429D2A9" box="[151,352,1396,1420]" italics="true" pageId="15" pageNumber="491">Xanthias maculatus</emphasis>
</taxonomicName>
<bibRefCitation id="EFD24B767F24FF87FE97FA95B7E5D2A9" author="Takeda, M. &amp; Komatsu, H." box="[377,684,1396,1420]" pageId="15" pageNumber="491" pagination="271 - 88" refId="ref15343" refString="Takeda, M. &amp; Komatsu, H. (2005) Collections of crabs dredged off Amami-Oshima Island, the northern Ryukyu Islands. National Science Museum Monographs, Tokyo, 29, 271 - 88." type="journal article" year="2005">Takeda &amp; Komatsu 2005: 282</bibRefCitation>
, fig. 5A [Not
<taxonomicName id="4C434D047F24FF87FCD6FA95B1CAD2A9" ID-CoL="5C4TS" authority="Sakai, 1961" authorityName="Sakai" authorityYear="1961" box="[824,1155,1396,1420]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="491" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B937EA957F24FF87FCD6FA95B148D2A9" box="[824,1025,1396,1420]" italics="true" pageId="15" pageNumber="491">Xanthias maculatus</emphasis>
<bibRefCitation id="EFD24B767F24FF87FBE9FA95B1CAD2A9" author="Sakai, T." box="[1031,1155,1396,1420]" pageId="15" pageNumber="491" pagination="131 - 150" refId="ref14171" refString="Sakai, T. (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2), 131 - 150, pls. 3 - 4. https: // doi. org / 10.1163 / 156854061 X 00635" type="journal article" year="1961">Sakai, 1961</bibRefCitation>
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].
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<emphasis id="B937EA957F24FF87FF79FA5BB4CDD2F1" bold="true" box="[151,388,1466,1492]" pageId="15" pageNumber="491">Material examined.</emphasis>
<materialsCitation id="3B2B3CDA7F24FF87FE63FA5BB074D2DD" collectingDate="2019-08-16" collectionCode="FORVSS" collectorName="V. P. Padate" country="India" elevation="56" latitude="6.65" location="Great Nicobar Island" longLatPrecision="782" longitude="93.83" municipality="Nicobars" pageId="15" pageNumber="491" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D45FD0E7F24FF87FE63FA5BB492D2F1" box="[397,475,1466,1492]" count="1" pageId="15" pageNumber="491" type="male">1 male</specimenCount>
(6.4 ×
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) (IO/SS/BRC/00197),
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stn. 38818, south of
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,
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<quantity id="4CBB9B627F24FF87FD82FA3FB7ECD2DC" box="[620,677,1502,1529]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="15" pageNumber="491" unit="m" value="56.0">56 m</quantity>
depth
</elevation>
, chain bag dredge, coll.
<collectorName id="26B653517F24FF87FC13FA3EB134D2DD" box="[1021,1149,1502,1529]" pageId="15" pageNumber="491">V.P. Padate</collectorName>
,
<date id="FFFD10477F24FF87FB69FA3FB074D2DD" box="[1159,1341,1502,1528]" pageId="15" pageNumber="491" value="2019-08-16">
<collectingDate id="EFB9E9AF7F24FF87FB69FA3FB074D2DD" box="[1159,1341,1502,1528]" pageId="15" pageNumber="491" value="2019-08-16">16 August 2019</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C359650C7F24FF98FF29F9E3B48ED7DC" lastPageId="16" lastPageNumber="492" pageId="15" pageNumber="491" type="description">
<paragraph id="8BFC36877F24FF87FF29F9E3B03DD189" blockId="15.[151,1437,1466,2033]" pageId="15" pageNumber="491">
<emphasis id="B937EA957F24FF87FF29F9E3B410D139" bold="true" box="[199,345,1538,1564]" pageId="15" pageNumber="491">Description.</emphasis>
Carapace transversely ovate, subhexagonal, 1.5 times as long as wide, dorsal surface smooth, glabrous, only orbital region and anterior portions of 2M, 3M regions indistinctly demarcated (
<figureCitation id="13782A027F24FF87FB1CF9C7B013D165" box="[1266,1370,1574,1600]" captionStart="FIGURE 2" captionStartId="3.[152,255,1683,1707]" captionTargetBox="[177,1410,181,1659]" captionTargetId="figure-37@3.[177,1410,181,1659]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal habitus photographs (AC, EG: fresh specimens; D: ethanol-preserved specimen): (A) Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm); (B) Urnalana angulata (Rathbun, 1911), undetermined sex (IO/SS/BRC/00184, 6.9 × 7.0 mm); (C) Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm; (D) Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm); (E) Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm); (F) Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/SS/ BRC/00194, 5.2 × 3.6 mm); (G) Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). Scale bars: AB, E, G, 2 mm, C, 10 mm, D, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210274" httpUri="https://zenodo.org/record/13210274/files/figure.png" pageId="15" pageNumber="491">Figs. 2G</figureCitation>
,
<figureCitation id="13782A027F24FF87FA84F9C7B0C6D165" box="[1386,1423,1574,1600]" captionStart="FIGURE 9" captionStartId="16.[152,255,1914,1938]" captionTargetBox="[281,1306,597,1890]" captionTargetId="figure-220@16.[281,1306,597,1890]" captionTargetPageId="16" captionText="FIGURE 9. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm), Andaman Sea, India: colour photographs: (A) carapace, dorsal view; (B) mxp3, chelipeds, pereopods, sternum and pleon, ventral view. Scale bars: AB, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210296" httpUri="https://zenodo.org/record/13210296/files/figure.png" pageId="15" pageNumber="491">9A</figureCitation>
). Frontal margin slightly deflexed, divided into 2 gently convex lobes by wide V-shaped notch. Anterolateral margin convex, divided into 4 lobiform teeth separated by V-shaped notches. Posterolateral margin longer than anterolateral margin, nearly straight, distinctly converging posteriorly. Posterior margin slightly concave medially (
<figureCitation id="13782A027F24FF87FAE3F973B02ED189" box="[1293,1383,1682,1708]" captionStart="FIGURE 9" captionStartId="16.[152,255,1914,1938]" captionTargetBox="[281,1306,597,1890]" captionTargetId="figure-220@16.[281,1306,597,1890]" captionTargetPageId="16" captionText="FIGURE 9. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm), Andaman Sea, India: colour photographs: (A) carapace, dorsal view; (B) mxp3, chelipeds, pereopods, sternum and pleon, ventral view. Scale bars: AB, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210296" httpUri="https://zenodo.org/record/13210296/files/figure.png" pageId="15" pageNumber="491">Fig. 9A</figureCitation>
).
</paragraph>
<paragraph id="8BFC36877F24FF87FF29F957B44CD1D1" blockId="15.[151,1437,1466,2033]" pageId="15" pageNumber="491">
Antennules folding transversely; mxp3 merus subquadrate, half as long as ischium, anterolateral angle rounded (
<figureCitation id="13782A027F24FF87FF71F93BB5B1D1D1" box="[159,248,1754,1780]" captionStart="FIGURE 9" captionStartId="16.[152,255,1914,1938]" captionTargetBox="[281,1306,597,1890]" captionTargetId="figure-220@16.[281,1306,597,1890]" captionTargetPageId="16" captionText="FIGURE 9. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm), Andaman Sea, India: colour photographs: (A) carapace, dorsal view; (B) mxp3, chelipeds, pereopods, sternum and pleon, ventral view. Scale bars: AB, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210296" httpUri="https://zenodo.org/record/13210296/files/figure.png" pageId="15" pageNumber="491">Fig. 9B</figureCitation>
).
</paragraph>
<paragraph id="8BFC36877F24FF87FF29F91FB1ADD019" blockId="15.[151,1437,1466,2033]" pageId="15" pageNumber="491">
Chelipeds massive, subequal, 1.4 times as long as CL, surfaces smooth, dactylus robust, longer than palm (
<figureCitation id="13782A027F24FF87FA81F91FB5F5D019" captionStart="FIGURE 9" captionStartId="16.[152,255,1914,1938]" captionTargetBox="[281,1306,597,1890]" captionTargetId="figure-220@16.[281,1306,597,1890]" captionTargetPageId="16" captionText="FIGURE 9. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm), Andaman Sea, India: colour photographs: (A) carapace, dorsal view; (B) mxp3, chelipeds, pereopods, sternum and pleon, ventral view. Scale bars: AB, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210296" httpUri="https://zenodo.org/record/13210296/files/figure.png" pageId="15" pageNumber="491">Fig. 9B</figureCitation>
). Pereopods 25 shorter than chelipeds, P4 longest, dactyli nearly as long as propodi (
<figureCitation id="13782A027F24FF87FB91F8C3B191D019" box="[1151,1240,1826,1852]" captionStart="FIGURE 9" captionStartId="16.[152,255,1914,1938]" captionTargetBox="[281,1306,597,1890]" captionTargetId="figure-220@16.[281,1306,597,1890]" captionTargetPageId="16" captionText="FIGURE 9. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm), Andaman Sea, India: colour photographs: (A) carapace, dorsal view; (B) mxp3, chelipeds, pereopods, sternum and pleon, ventral view. Scale bars: AB, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210296" httpUri="https://zenodo.org/record/13210296/files/figure.png" pageId="15" pageNumber="491">Fig. 9B</figureCitation>
).
</paragraph>
<paragraph id="8BFC36877F24FF87FF29F8A7B4B4D08D" blockId="15.[151,1437,1466,2033]" pageId="15" pageNumber="491">
Male pleon subtriangular, somites 35 fused; telson subtriangular, 0.7 times as long as wide (
<figureCitation id="13782A027F24FF87FB22F8A7B06DD045" box="[1228,1316,1862,1888]" captionStart="FIGURE 9" captionStartId="16.[152,255,1914,1938]" captionTargetBox="[281,1306,597,1890]" captionTargetId="figure-220@16.[281,1306,597,1890]" captionTargetPageId="16" captionText="FIGURE 9. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm), Andaman Sea, India: colour photographs: (A) carapace, dorsal view; (B) mxp3, chelipeds, pereopods, sternum and pleon, ventral view. Scale bars: AB, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210296" httpUri="https://zenodo.org/record/13210296/files/figure.png" pageId="15" pageNumber="491">Fig. 9B</figureCitation>
). G1 stout, distal half curving laterally; distal tip with narrow apical lobe, subdistal portion with patch of spinules and several subterminal setae (
<figureCitation id="13782A027F24FF87FE84F86FB4B9D08D" box="[362,496,1934,1960]" captionStart="FIGURE 10" captionStartId="17.[151,250,1667,1691]" captionTargetBox="[304,1294,191,1633]" captionTargetId="figure-37@17.[292,1295,181,1645]" captionTargetPageId="17" captionText="FIGURE 10. First and second gonopods: Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm): (A) entire G1, pleonal view; (B) entire G1, sternal view; (C) entire G2, pleonal view. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm): (D) entire G1, pleonal view; (E) G1 distal portion, pleonal view; (F) G1 distal portion, sternal view; (G) entire G1, sternal view; (H) entire G2, pleonal view. Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm): (I) entire G1, pleonal view; (J) entire G1, lateral view; (K) entire G1, mesial view; (L) entire G2, pleonal view. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm): (M) entire G1, pleonal view; (N) entire G1, sternal view; (O) entire G2, sternal view. Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/ SS/BRC/00194, 5.2 × 3.6 mm): (P) entire G1, pleonal view; (Q) G1 distal portion, pleonal view; (R) entire G1, sternal view. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). (S) entire G1, pleonal view; (T) G1 distal portion, pleonal view; (U) G1 distal portion, sternal view; (V) entire G1, sternal view. Scale bars: AC, PV, 0.2 mm, D, GN, 1 mm, E, F, O, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210300" httpUri="https://zenodo.org/record/13210300/files/figure.png" pageId="15" pageNumber="491">Fig. 10SV</figureCitation>
).
</paragraph>
<paragraph id="8BFC36877F24FF98FF29F853B48ED7DC" blockId="15.[151,1437,1466,2033]" lastBlockId="16.[151,1437,151,573]" lastPageId="16" lastPageNumber="492" pageId="15" pageNumber="491">
<emphasis id="B937EA957F24FF87FF29F853B416D0E9" bold="true" box="[199,351,1970,1996]" pageId="15" pageNumber="491">Colouration.</emphasis>
Fresh specimen: Carapace, chelipeds and pereopods light yellowish orange with randomly arranged ocelli or bluish-white spots fringed with dark orange. Carapace with 48 ocelli (
<figureCitation id="13782A027F24FF87FB9EF837B183D0D5" box="[1136,1226,2006,2032]" captionStart="FIGURE 2" captionStartId="3.[152,255,1683,1707]" captionTargetBox="[177,1410,181,1659]" captionTargetId="figure-37@3.[177,1410,181,1659]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal habitus photographs (AC, EG: fresh specimens; D: ethanol-preserved specimen): (A) Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm); (B) Urnalana angulata (Rathbun, 1911), undetermined sex (IO/SS/BRC/00184, 6.9 × 7.0 mm); (C) Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm; (D) Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm); (E) Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm); (F) Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/SS/ BRC/00194, 5.2 × 3.6 mm); (G) Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). Scale bars: AB, E, G, 2 mm, C, 10 mm, D, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210274" httpUri="https://zenodo.org/record/13210274/files/figure.png" pageId="15" pageNumber="491">Fig. 2G</figureCitation>
). Lower distal half of cheliped palm with saddle-shaped dark olive patch extending to proximal four-fifths of fingers, finger tips whitish (
<figureCitation id="13782A027F3BFF98FF71FF5AB44DD7F0" box="[159,260,187,213]" captionStart="FIGURE 2" captionStartId="3.[152,255,1683,1707]" captionTargetBox="[177,1410,181,1659]" captionTargetId="figure-37@3.[177,1410,181,1659]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal habitus photographs (AC, EG: fresh specimens; D: ethanol-preserved specimen): (A) Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm); (B) Urnalana angulata (Rathbun, 1911), undetermined sex (IO/SS/BRC/00184, 6.9 × 7.0 mm); (C) Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm; (D) Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm); (E) Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm); (F) Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/SS/ BRC/00194, 5.2 × 3.6 mm); (G) Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). Scale bars: AB, E, G, 2 mm, C, 10 mm, D, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210274" httpUri="https://zenodo.org/record/13210274/files/figure.png" pageId="16" pageNumber="492">Figs. 2G</figureCitation>
,
<figureCitation id="13782A027F3BFF98FEFEFF5AB47DD7F0" box="[272,308,187,213]" captionStart="FIGURE 9" captionStartId="16.[152,255,1914,1938]" captionTargetBox="[281,1306,597,1890]" captionTargetId="figure-220@16.[281,1306,597,1890]" captionTargetPageId="16" captionText="FIGURE 9. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm), Andaman Sea, India: colour photographs: (A) carapace, dorsal view; (B) mxp3, chelipeds, pereopods, sternum and pleon, ventral view. Scale bars: AB, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210296" httpUri="https://zenodo.org/record/13210296/files/figure.png" pageId="16" pageNumber="492">9B</figureCitation>
). Pereopods with 24 ocelli on the dorsal surfaces of meri, carpi and propodi and white bands at the articulation (
<figureCitation id="13782A027F3BFF98FEC8FF3EB4C3D7DC" box="[294,394,223,249]" captionStart="FIGURE 2" captionStartId="3.[152,255,1683,1707]" captionTargetBox="[177,1410,181,1659]" captionTargetId="figure-37@3.[177,1410,181,1659]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal habitus photographs (AC, EG: fresh specimens; D: ethanol-preserved specimen): (A) Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm); (B) Urnalana angulata (Rathbun, 1911), undetermined sex (IO/SS/BRC/00184, 6.9 × 7.0 mm); (C) Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm; (D) Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm); (E) Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm); (F) Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/SS/ BRC/00194, 5.2 × 3.6 mm); (G) Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). Scale bars: AB, E, G, 2 mm, C, 10 mm, D, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210274" httpUri="https://zenodo.org/record/13210274/files/figure.png" pageId="16" pageNumber="492">Figs. 2G</figureCitation>
,
<figureCitation id="13782A027F3BFF98FE78FF3EB4F3D7DC" box="[406,442,223,249]" captionStart="FIGURE 9" captionStartId="16.[152,255,1914,1938]" captionTargetBox="[281,1306,597,1890]" captionTargetId="figure-220@16.[281,1306,597,1890]" captionTargetPageId="16" captionText="FIGURE 9. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm), Andaman Sea, India: colour photographs: (A) carapace, dorsal view; (B) mxp3, chelipeds, pereopods, sternum and pleon, ventral view. Scale bars: AB, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210296" httpUri="https://zenodo.org/record/13210296/files/figure.png" pageId="16" pageNumber="492">9B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C359650C7F3BFF98FF29FEE3B7A5D640" pageId="16" pageNumber="492" type="biology_ecology">
<paragraph id="8BFC36877F3BFF98FF29FEE3B7A5D640" blockId="16.[151,1437,151,573]" pageId="16" pageNumber="492">
<emphasis id="B937EA957F3BFF98FF29FEE3B46ED639" bold="true" box="[199,295,258,284]" pageId="16" pageNumber="492">Biology.</emphasis>
The present specimen was collected from rocky substrate in the vicinity of crinoids, at
<quantity id="4CBB9B627F3BFF98FAFAFEE3B006D638" box="[1300,1359,258,285]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="16" pageNumber="492" unit="m" value="56.0">56 m</quantity>
depth. Known to occur in coral rubble on sand and gravel, subtidal coral platform, reef slopes and deep waters from
<quantity id="4CBB9B627F3BFF98FAB8FEC6B5E4D640" metricMagnitude="2" metricUnit="m" metricValue="1.465" metricValueMax="2.9" metricValueMin="0.03" pageId="16" pageNumber="492" unit="m" value="146.5" valueMax="290.0" valueMin="3.0">3290 m</quantity>
depths (
<bibRefCitation id="EFD24B767F3BFF98FEE2FEAAB4F1D640" author="Mendoza, J. C. E." box="[268,440,331,357]" pageId="16" pageNumber="492" pagination="374 - 384" refId="ref12717" refString="Mendoza, J. C. E. (2013) A new species of ocellated Xanthias Rathbun, 1897 (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, Philippines. Zootaxa, 3636 (2), 374 - 384. https: // doi. org / 10.11646 / zootaxa. 3636.2.8" type="journal article" year="2013">Mendoza 2013</bibRefCitation>
;
<bibRefCitation id="EFD24B767F3BFF98FE2DFEAAB7A9D640" author="Takeda, M. &amp; Komatsu, H." box="[451,736,331,357]" pageId="16" pageNumber="492" pagination="153 - 189" refId="ref15381" refString="Takeda, M. &amp; Komatsu, H. (2018) Offshore Crabs of the Family Xanthidae and Some Related Families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Science Museum, Tokyo, 52, 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C359650C7F3BFF9AFF29FE8EB622D5D4" lastPageId="18" lastPageNumber="494" pageId="16" pageNumber="492" type="discussion">
<paragraph id="8BFC36877F3BFF9AFF29FE8EB12AD5A0" blockId="16.[151,1437,151,573]" lastBlockId="18.[151,1437,150,753]" lastPageId="18" lastPageNumber="494" pageId="16" pageNumber="492">
<emphasis id="B937EA957F3BFF98FF29FE8EB472D6AC" bold="true" box="[199,315,367,393]" pageId="16" pageNumber="492">Remarks.</emphasis>
<bibRefCitation id="EFD24B767F3BFF98FEABFE8EB74AD6AC" author="Mendoza, J. C. E." box="[325,515,367,393]" pageId="16" pageNumber="492" pagination="374 - 384" refId="ref12717" refString="Mendoza, J. C. E. (2013) A new species of ocellated Xanthias Rathbun, 1897 (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, Philippines. Zootaxa, 3636 (2), 374 - 384. https: // doi. org / 10.11646 / zootaxa. 3636.2.8" type="journal article" year="2013">Mendoza (2013)</bibRefCitation>
described
<taxonomicName id="4C434D047F3BFF98FD6DFE8EB611D6AD" authorityName="Mendoza" authorityYear="2013" box="[643,856,367,393]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="492" phylum="Arthropoda" rank="species" species="joanneae">
<emphasis id="B937EA957F3BFF98FD6DFE8EB611D6AD" box="[643,856,367,393]" italics="true" pageId="16" pageNumber="492">Xanthias joanneae</emphasis>
</taxonomicName>
from
<specimenCount id="9D45FD0E7F3BFF98FC4DFE8EB6B5D6AC" box="[931,1020,367,393]" count="6" pageId="16" pageNumber="492" type="male">6 males</specimenCount>
and
<specimenCount id="9D45FD0E7F3BFF98FBD4FE8EB1E0D6AC" box="[1082,1193,367,393]" count="8" pageId="16" pageNumber="492" type="female">8 females</specimenCount>
collected from coral reef microhabitats at
<quantity id="4CBB9B627F3BFF98FE6BFE72B4AED688" box="[389,487,403,429]" metricMagnitude="1" metricUnit="m" metricValue="5.6" metricValueMax="11.0" metricValueMin="0.2" pageId="16" pageNumber="492" unit="m" value="56.0" valueMax="110.0" valueMin="2.0">2110 m</quantity>
depths in the
<collectingRegion id="4987F8657F3BFF98FD6AFE72B781D688" box="[644,712,403,429]" country="Philippines" name="Bohol" pageId="16" pageNumber="492">Bohol</collectingRegion>
Sea,
<collectingCountry id="F35476177F3BFF98FCE8FE72B6C1D688" box="[774,904,403,429]" name="Philippines" pageId="16" pageNumber="492">Philippines</collectingCountry>
.
<taxonomicName id="4C434D047F3BFF98FC7DFE72B12DD689" authorityName="Mendoza" authorityYear="2013" box="[915,1124,403,429]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="492" phylum="Arthropoda" rank="species" species="joanneae">
<emphasis id="B937EA957F3BFF98FC7DFE72B12DD689" box="[915,1124,403,429]" italics="true" pageId="16" pageNumber="492">Xanthias joanneae</emphasis>
</taxonomicName>
closely resembles
<taxonomicName id="4C434D047F3BFF98FAD7FE72B4DCD6F4" authority="Sakai, 1961" authorityName="Sakai" authorityYear="1961" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="492" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B937EA957F3BFF98FAD7FE72B443D6F4" italics="true" pageId="16" pageNumber="492">Xanthias maculatus</emphasis>
<bibRefCitation id="EFD24B767F3BFF98FEFFFE56B4DCD6F4" author="Sakai, T." box="[273,405,439,465]" pageId="16" pageNumber="492" pagination="131 - 150" refId="ref14171" refString="Sakai, T. (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2), 131 - 150, pls. 3 - 4. https: // doi. org / 10.1163 / 156854061 X 00635" type="journal article" year="1961">Sakai, 1961</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C434D047F3BFF98FE25FE56B603D6F4" authority="Mendoza, 2023" authorityName="Mendoza" authorityYear="2023" box="[459,842,439,465]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="492" phylum="Arthropoda" rank="species" species="claudiae">
<emphasis id="B937EA957F3BFF98FE25FE56B7DCD6F4" box="[459,661,439,465]" italics="true" pageId="16" pageNumber="492">Xanthias claudiae</emphasis>
<bibRefCitation id="EFD24B767F3BFF98FD72FE56B603D6F4" author="Mendoza, J. C. E." box="[668,842,439,465]" pageId="16" pageNumber="492" pagination="119 - 200" refId="ref12776" refString="Mendoza, J. C. E. (2023) The xanthid crabs (Crustacea: Brachyura: Xanthoidea: Xanthidae) of Madagascar and adjacent southwestern Indian Ocean regions from recent marine biodiversity expeditions, 2009 - 2017. Deep-Sea Crustaceans from South-West Indian Ocean. Tropical Deep-Sea Benthos 33. Memoires du Museum national d'Histoire naturelle, 217, 119 - 200." type="journal article" year="2023">
Mendoza,
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in the smooth texture of the carapace and pereopods, basal width of male telson exceeding that of anterior margin of pleonal somite 6, and the presence of ocelli on the body and appendages. However, it differs from the other two species in the higher number of small ocelli (&gt; 40) on the carapace (
<figureCitation id="13782A027F3BFF98FE6BFDC2B4A0D518" box="[389,489,547,573]" captionStart="FIGURE 2" captionStartId="3.[152,255,1683,1707]" captionTargetBox="[177,1410,181,1659]" captionTargetId="figure-37@3.[177,1410,181,1659]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal habitus photographs (AC, EG: fresh specimens; D: ethanol-preserved specimen): (A) Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm); (B) Urnalana angulata (Rathbun, 1911), undetermined sex (IO/SS/BRC/00184, 6.9 × 7.0 mm); (C) Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm; (D) Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm); (E) Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm); (F) Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/SS/ BRC/00194, 5.2 × 3.6 mm); (G) Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). Scale bars: AB, E, G, 2 mm, C, 10 mm, D, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210274" httpUri="https://zenodo.org/record/13210274/files/figure.png" pageId="16" pageNumber="492">Figs. 2G</figureCitation>
,
<figureCitation id="13782A027F3BFF98FE18FDC2B753D518" box="[502,538,547,573]" captionStart="FIGURE 9" captionStartId="16.[152,255,1914,1938]" captionTargetBox="[281,1306,597,1890]" captionTargetId="figure-220@16.[281,1306,597,1890]" captionTargetPageId="16" captionText="FIGURE 9. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm), Andaman Sea, India: colour photographs: (A) carapace, dorsal view; (B) mxp3, chelipeds, pereopods, sternum and pleon, ventral view. Scale bars: AB, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210296" httpUri="https://zenodo.org/record/13210296/files/figure.png" pageId="16" pageNumber="492">9A</figureCitation>
) (versus 1013 large ocelli in
<taxonomicName id="4C434D047F3BFF98FC9AFDC2B6BDD518" authorityName="Mendoza" authorityYear="2023" box="[884,1012,547,573]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="492" phylum="Arthropoda" rank="species" species="claudiae">
<emphasis id="B937EA957F3BFF98FC9AFDC2B6BDD518" box="[884,1012,547,573]" italics="true" pageId="16" pageNumber="492">X. claudiae</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD24B767F3BFF98FBEAFDC2B1FBD518" author="Mendoza, J. C. E." box="[1028,1202,547,573]" pageId="16" pageNumber="492" pagination="119 - 200" refId="ref12776" refString="Mendoza, J. C. E. (2023) The xanthid crabs (Crustacea: Brachyura: Xanthoidea: Xanthidae) of Madagascar and adjacent southwestern Indian Ocean regions from recent marine biodiversity expeditions, 2009 - 2017. Deep-Sea Crustaceans from South-West Indian Ocean. Tropical Deep-Sea Benthos 33. Memoires du Museum national d'Histoire naturelle, 217, 119 - 200." type="journal article" year="2023">Mendoza 2023</bibRefCitation>
: fig. 26A, B); 1420 moderately large ocelli in
<taxonomicName id="4C434D047F39FF9AFE28FF76B712D794" authorityName="Sakai" authorityYear="1961" box="[454,603,151,177]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="494" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B937EA957F39FF9AFE28FF76B712D794" box="[454,603,151,177]" italics="true" pageId="18" pageNumber="494">X. maculatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD24B767F39FF9AFD80FF76B6DBD794" author="Takeda, M. &amp; Komatsu, H." box="[622,914,151,177]" pageId="18" pageNumber="494" pagination="153 - 189" refId="ref15381" refString="Takeda, M. &amp; Komatsu, H. (2018) Offshore Crabs of the Family Xanthidae and Some Related Families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Science Museum, Tokyo, 52, 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu 2018</bibRefCitation>
: fig. 5B)); ill-defined 2M, 3M regions on the carapace (
<figureCitation id="13782A027F39FF9AFEE5FF5AB42CD7F0" box="[267,357,187,213]" captionStart="FIGURE 9" captionStartId="16.[152,255,1914,1938]" captionTargetBox="[281,1306,597,1890]" captionTargetId="figure-220@16.[281,1306,597,1890]" captionTargetPageId="16" captionText="FIGURE 9. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm), Andaman Sea, India: colour photographs: (A) carapace, dorsal view; (B) mxp3, chelipeds, pereopods, sternum and pleon, ventral view. Scale bars: AB, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210296" httpUri="https://zenodo.org/record/13210296/files/figure.png" pageId="18" pageNumber="494">Fig. 9A</figureCitation>
) (versus feebly indicated in
<taxonomicName id="4C434D047F39FF9AFD5EFF5AB678D7F0" authorityName="Mendoza" authorityYear="2023" box="[688,817,187,213]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="494" phylum="Arthropoda" rank="species" species="claudiae">
<emphasis id="B937EA957F39FF9AFD5EFF5AB678D7F0" box="[688,817,187,213]" italics="true" pageId="18" pageNumber="494">X. claudiae</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD24B767F39FF9AFCADFF5AB6BBD7F0" author="Mendoza, J. C. E." box="[835,1010,187,213]" pageId="18" pageNumber="494" pagination="119 - 200" refId="ref12776" refString="Mendoza, J. C. E. (2023) The xanthid crabs (Crustacea: Brachyura: Xanthoidea: Xanthidae) of Madagascar and adjacent southwestern Indian Ocean regions from recent marine biodiversity expeditions, 2009 - 2017. Deep-Sea Crustaceans from South-West Indian Ocean. Tropical Deep-Sea Benthos 33. Memoires du Museum national d'Histoire naturelle, 217, 119 - 200." type="journal article" year="2023">Mendoza 2023</bibRefCitation>
: figs. 27A, 28A); discernible regions in
<taxonomicName id="4C434D047F39FF9AFF5BFF3EB401D7DC" authorityName="Sakai" authorityYear="1961" box="[181,328,223,249]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="494" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B937EA957F39FF9AFF5BFF3EB401D7DC" box="[181,328,223,249]" italics="true" pageId="18" pageNumber="494">X. maculatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD24B767F39FF9AFEB6FF3EB74FD7DC" author="Mendoza, J. C. E." box="[344,518,223,249]" pageId="18" pageNumber="494" pagination="374 - 384" refId="ref12717" refString="Mendoza, J. C. E. (2013) A new species of ocellated Xanthias Rathbun, 1897 (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, Philippines. Zootaxa, 3636 (2), 374 - 384. https: // doi. org / 10.11646 / zootaxa. 3636.2.8" type="journal article" year="2013">Mendoza 2013</bibRefCitation>
: fig. 5A)); moderately lobiform anterolateral teeth separated by narrow V-shaped notches (
<figureCitation id="13782A027F39FF9AFF12FEE2B41DD638" box="[252,340,259,285]" captionStart="FIGURE 9" captionStartId="16.[152,255,1914,1938]" captionTargetBox="[281,1306,597,1890]" captionTargetId="figure-220@16.[281,1306,597,1890]" captionTargetPageId="16" captionText="FIGURE 9. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm), Andaman Sea, India: colour photographs: (A) carapace, dorsal view; (B) mxp3, chelipeds, pereopods, sternum and pleon, ventral view. Scale bars: AB, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210296" httpUri="https://zenodo.org/record/13210296/files/figure.png" pageId="18" pageNumber="494">Fig. 9A</figureCitation>
) (versus indistinctly lobiform teeth in
<taxonomicName id="4C434D047F39FF9AFCEDFEE2B6CBD638" authorityName="Mendoza" authorityYear="2023" box="[771,898,259,285]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="494" phylum="Arthropoda" rank="species" species="claudiae">
<emphasis id="B937EA957F39FF9AFCEDFEE2B6CBD638" box="[771,898,259,285]" italics="true" pageId="18" pageNumber="494">X. claudiae</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD24B767F39FF9AFC7FFEE2B177D638" author="Mendoza, J. C. E." box="[913,1086,259,285]" pageId="18" pageNumber="494" pagination="119 - 200" refId="ref12776" refString="Mendoza, J. C. E. (2023) The xanthid crabs (Crustacea: Brachyura: Xanthoidea: Xanthidae) of Madagascar and adjacent southwestern Indian Ocean regions from recent marine biodiversity expeditions, 2009 - 2017. Deep-Sea Crustaceans from South-West Indian Ocean. Tropical Deep-Sea Benthos 33. Memoires du Museum national d'Histoire naturelle, 217, 119 - 200." type="journal article" year="2023">Mendoza 2023</bibRefCitation>
: figs. 27A, 28A); distinct acute teeth separated by wide troughs in
<taxonomicName id="4C434D047F39FF9AFDCAFEC6B7FED664" authorityName="Sakai" authorityYear="1961" box="[548,695,295,321]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="494" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B937EA957F39FF9AFDCAFEC6B7FED664" box="[548,695,295,321]" italics="true" pageId="18" pageNumber="494">X. maculatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD24B767F39FF9AFD29FEC6B63CD664" author="Mendoza, J. C. E." box="[711,885,295,321]" pageId="18" pageNumber="494" pagination="374 - 384" refId="ref12717" refString="Mendoza, J. C. E. (2013) A new species of ocellated Xanthias Rathbun, 1897 (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, Philippines. Zootaxa, 3636 (2), 374 - 384. https: // doi. org / 10.11646 / zootaxa. 3636.2.8" type="journal article" year="2013">Mendoza 2013</bibRefCitation>
: fig. 5A)); a relatively narrow thoracic sternite 4 with straight lateral margins (
<figureCitation id="13782A027F39FF9AFE32FEAAB779D640" box="[476,560,331,357]" captionStart="FIGURE 9" captionStartId="16.[152,255,1914,1938]" captionTargetBox="[281,1306,597,1890]" captionTargetId="figure-220@16.[281,1306,597,1890]" captionTargetPageId="16" captionText="FIGURE 9. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm), Andaman Sea, India: colour photographs: (A) carapace, dorsal view; (B) mxp3, chelipeds, pereopods, sternum and pleon, ventral view. Scale bars: AB, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210296" httpUri="https://zenodo.org/record/13210296/files/figure.png" pageId="18" pageNumber="494">Fig. 9B</figureCitation>
) (versus relatively wider sternite 4 with convex lateral margins in
<taxonomicName id="4C434D047F39FF9AFAE5FEAAB0D5D640" authorityName="Sakai" authorityYear="1961" box="[1291,1436,331,357]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="494" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B937EA957F39FF9AFAE5FEAAB0D5D640" box="[1291,1436,331,357]" italics="true" pageId="18" pageNumber="494">X. maculatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD24B767F39FF9AFF71FE8EB405D6AC" author="Mendoza, J. C. E." box="[159,332,367,393]" pageId="18" pageNumber="494" pagination="374 - 384" refId="ref12717" refString="Mendoza, J. C. E. (2013) A new species of ocellated Xanthias Rathbun, 1897 (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, Philippines. Zootaxa, 3636 (2), 374 - 384. https: // doi. org / 10.11646 / zootaxa. 3636.2.8" type="journal article" year="2013">Mendoza 2013</bibRefCitation>
: fig. 5B)); smooth external surface of the cheliped palm (
<figureCitation id="13782A027F39FF9AFC22FE8EB168D6AC" box="[972,1057,367,393]" captionStart="FIGURE 9" captionStartId="16.[152,255,1914,1938]" captionTargetBox="[281,1306,597,1890]" captionTargetId="figure-220@16.[281,1306,597,1890]" captionTargetPageId="16" captionText="FIGURE 9. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm), Andaman Sea, India: colour photographs: (A) carapace, dorsal view; (B) mxp3, chelipeds, pereopods, sternum and pleon, ventral view. Scale bars: AB, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210296" httpUri="https://zenodo.org/record/13210296/files/figure.png" pageId="18" pageNumber="494">Fig. 9B</figureCitation>
) (versus palm with 2 longitudinal ridges on lower outer surface in
<taxonomicName id="4C434D047F39FF9AFE14FE72B731D688" authorityName="Mendoza" authorityYear="2023" box="[506,632,403,429]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="494" phylum="Arthropoda" rank="species" species="claudiae">
<emphasis id="B937EA957F39FF9AFE14FE72B731D688" box="[506,632,403,429]" italics="true" pageId="18" pageNumber="494">X. claudiae</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD24B767F39FF9AFD68FE72B67BD688" author="Mendoza, J. C. E." box="[646,818,403,429]" pageId="18" pageNumber="494" pagination="119 - 200" refId="ref12776" refString="Mendoza, J. C. E. (2023) The xanthid crabs (Crustacea: Brachyura: Xanthoidea: Xanthidae) of Madagascar and adjacent southwestern Indian Ocean regions from recent marine biodiversity expeditions, 2009 - 2017. Deep-Sea Crustaceans from South-West Indian Ocean. Tropical Deep-Sea Benthos 33. Memoires du Museum national d'Histoire naturelle, 217, 119 - 200." type="journal article" year="2023">Mendoza 2023</bibRefCitation>
: fig. 28I, J) and
<taxonomicName id="4C434D047F39FF9AFC0CFE72B13BD688" authorityName="Sakai" authorityYear="1961" box="[994,1138,403,429]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="494" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B937EA957F39FF9AFC0CFE72B13BD688" box="[994,1138,403,429]" italics="true" pageId="18" pageNumber="494">X. maculatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD24B767F39FF9AFB6EFE72B065D688" author="Mendoza, J. C. E." box="[1152,1324,403,429]" pageId="18" pageNumber="494" pagination="374 - 384" refId="ref12717" refString="Mendoza, J. C. E. (2013) A new species of ocellated Xanthias Rathbun, 1897 (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, Philippines. Zootaxa, 3636 (2), 374 - 384. https: // doi. org / 10.11646 / zootaxa. 3636.2.8" type="journal article" year="2013">Mendoza 2013</bibRefCitation>
: fig. 5B)); relatively broader pereopod meri (
<figureCitation id="13782A027F39FF9AFDCDFE56B734D6F4" box="[547,637,439,465]" captionStart="FIGURE 2" captionStartId="3.[152,255,1683,1707]" captionTargetBox="[177,1410,181,1659]" captionTargetId="figure-37@3.[177,1410,181,1659]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal habitus photographs (AC, EG: fresh specimens; D: ethanol-preserved specimen): (A) Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm); (B) Urnalana angulata (Rathbun, 1911), undetermined sex (IO/SS/BRC/00184, 6.9 × 7.0 mm); (C) Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm; (D) Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm); (E) Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm); (F) Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/SS/ BRC/00194, 5.2 × 3.6 mm); (G) Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). Scale bars: AB, E, G, 2 mm, C, 10 mm, D, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210274" httpUri="https://zenodo.org/record/13210274/files/figure.png" pageId="18" pageNumber="494">Fig. 2G</figureCitation>
) (versus relatively narrower pereopod meri in
<taxonomicName id="4C434D047F39FF9AFB70FE56B069D6F4" authorityName="Mendoza" authorityYear="2023" box="[1182,1312,439,465]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="494" phylum="Arthropoda" rank="species" species="claudiae">
<emphasis id="B937EA957F39FF9AFB70FE56B069D6F4" box="[1182,1312,439,465]" italics="true" pageId="18" pageNumber="494">X. claudiae</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD24B767F39FF9AFADCFE56B59AD6D0" author="Mendoza, J. C. E." pageId="18" pageNumber="494" pagination="119 - 200" refId="ref12776" refString="Mendoza, J. C. E. (2023) The xanthid crabs (Crustacea: Brachyura: Xanthoidea: Xanthidae) of Madagascar and adjacent southwestern Indian Ocean regions from recent marine biodiversity expeditions, 2009 - 2017. Deep-Sea Crustaceans from South-West Indian Ocean. Tropical Deep-Sea Benthos 33. Memoires du Museum national d'Histoire naturelle, 217, 119 - 200." type="journal article" year="2023">Mendoza 2023</bibRefCitation>
: fig. 27G) and
<taxonomicName id="4C434D047F39FF9AFE6FFE3AB75DD6D0" authorityName="Sakai" authorityYear="1961" box="[385,532,475,501]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="494" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B937EA957F39FF9AFE6FFE3AB75DD6D0" box="[385,532,475,501]" italics="true" pageId="18" pageNumber="494">X. maculatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD24B767F39FF9AFDCAFE3AB79AD6D0" author="Mendoza, J. C. E." box="[548,723,475,501]" pageId="18" pageNumber="494" pagination="374 - 384" refId="ref12717" refString="Mendoza, J. C. E. (2013) A new species of ocellated Xanthias Rathbun, 1897 (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, Philippines. Zootaxa, 3636 (2), 374 - 384. https: // doi. org / 10.11646 / zootaxa. 3636.2.8" type="journal article" year="2013">Mendoza 2013</bibRefCitation>
: fig. 5A)); and a relatively stouter G1 with a wider apical lobe (
<figureCitation id="13782A027F39FF9AFF71FE1EB46BD53C" box="[159,290,511,537]" captionStart="FIGURE 10" captionStartId="17.[151,250,1667,1691]" captionTargetBox="[304,1294,191,1633]" captionTargetId="figure-37@17.[292,1295,181,1645]" captionTargetPageId="17" captionText="FIGURE 10. First and second gonopods: Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm): (A) entire G1, pleonal view; (B) entire G1, sternal view; (C) entire G2, pleonal view. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm): (D) entire G1, pleonal view; (E) G1 distal portion, pleonal view; (F) G1 distal portion, sternal view; (G) entire G1, sternal view; (H) entire G2, pleonal view. Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm): (I) entire G1, pleonal view; (J) entire G1, lateral view; (K) entire G1, mesial view; (L) entire G2, pleonal view. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm): (M) entire G1, pleonal view; (N) entire G1, sternal view; (O) entire G2, sternal view. Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/ SS/BRC/00194, 5.2 × 3.6 mm): (P) entire G1, pleonal view; (Q) G1 distal portion, pleonal view; (R) entire G1, sternal view. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). (S) entire G1, pleonal view; (T) G1 distal portion, pleonal view; (U) G1 distal portion, sternal view; (V) entire G1, sternal view. Scale bars: AC, PV, 0.2 mm, D, GN, 1 mm, E, F, O, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210300" httpUri="https://zenodo.org/record/13210300/files/figure.png" pageId="18" pageNumber="494">Fig. 10SV</figureCitation>
) (versus a relatively slender G1 with a narrower apical lobe in
<taxonomicName id="4C434D047F39FF9AFC3BFE1EB11AD53C" authorityName="Mendoza" authorityYear="2023" box="[981,1107,511,537]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="494" phylum="Arthropoda" rank="species" species="claudiae">
<emphasis id="B937EA957F39FF9AFC3BFE1EB11AD53C" box="[981,1107,511,537]" italics="true" pageId="18" pageNumber="494">X. claudiae</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD24B767F39FF9AFB8FFE1EB044D53C" author="Mendoza, J. C. E." box="[1121,1293,511,537]" pageId="18" pageNumber="494" pagination="119 - 200" refId="ref12776" refString="Mendoza, J. C. E. (2023) The xanthid crabs (Crustacea: Brachyura: Xanthoidea: Xanthidae) of Madagascar and adjacent southwestern Indian Ocean regions from recent marine biodiversity expeditions, 2009 - 2017. Deep-Sea Crustaceans from South-West Indian Ocean. Tropical Deep-Sea Benthos 33. Memoires du Museum national d'Histoire naturelle, 217, 119 - 200." type="journal article" year="2023">Mendoza 2023</bibRefCitation>
: fig. 29AC) and
<taxonomicName id="4C434D047F39FF9AFF29FDC2B410D518" authorityName="Sakai" authorityYear="1961" box="[199,345,547,573]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="494" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B937EA957F39FF9AFF29FDC2B410D518" box="[199,345,547,573]" italics="true" pageId="18" pageNumber="494">X. maculatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD24B767F39FF9AFE89FDC2B4A3D518" author="Sakai, T." box="[359,490,547,573]" pageId="18" pageNumber="494" pagination="131 - 150" refId="ref14171" refString="Sakai, T. (1961) New species of Japanese crabs from the collection of His Majesty the Emperor of Japan. Crustaceana, 3 (2), 131 - 150, pls. 3 - 4. https: // doi. org / 10.1163 / 156854061 X 00635" type="journal article" year="1961">Sakai 1961</bibRefCitation>
: fig. 1f, g)). The Indian specimen conforms to the description and illustrations of the
<typeStatus id="54F888257F39FF9AFF79FDA6B5B0D544" box="[151,249,583,609]" pageId="18" pageNumber="494" type="holotype">holotype</typeStatus>
provided by
<bibRefCitation id="EFD24B767F39FF9AFE65FDA6B70FD544" author="Mendoza, J. C. E." box="[395,582,583,609]" pageId="18" pageNumber="494" pagination="374 - 384" refId="ref12717" refString="Mendoza, J. C. E. (2013) A new species of ocellated Xanthias Rathbun, 1897 (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, Philippines. Zootaxa, 3636 (2), 374 - 384. https: // doi. org / 10.11646 / zootaxa. 3636.2.8" type="journal article" year="2013">Mendoza (2013)</bibRefCitation>
, however differs in having a long patch of spinules (
<figureCitation id="13782A027F39FF9AFB67FDA6B045D544" box="[1161,1292,583,609]" captionStart="FIGURE 10" captionStartId="17.[151,250,1667,1691]" captionTargetBox="[304,1294,191,1633]" captionTargetId="figure-37@17.[292,1295,181,1645]" captionTargetPageId="17" captionText="FIGURE 10. First and second gonopods: Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm): (A) entire G1, pleonal view; (B) entire G1, sternal view; (C) entire G2, pleonal view. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm): (D) entire G1, pleonal view; (E) G1 distal portion, pleonal view; (F) G1 distal portion, sternal view; (G) entire G1, sternal view; (H) entire G2, pleonal view. Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm): (I) entire G1, pleonal view; (J) entire G1, lateral view; (K) entire G1, mesial view; (L) entire G2, pleonal view. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm): (M) entire G1, pleonal view; (N) entire G1, sternal view; (O) entire G2, sternal view. Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/ SS/BRC/00194, 5.2 × 3.6 mm): (P) entire G1, pleonal view; (Q) G1 distal portion, pleonal view; (R) entire G1, sternal view. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). (S) entire G1, pleonal view; (T) G1 distal portion, pleonal view; (U) G1 distal portion, sternal view; (V) entire G1, sternal view. Scale bars: AC, PV, 0.2 mm, D, GN, 1 mm, E, F, O, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210300" httpUri="https://zenodo.org/record/13210300/files/figure.png" pageId="18" pageNumber="494">Fig. 10TU</figureCitation>
) (versus few spiniform granules;
<bibRefCitation id="EFD24B767F39FF9AFE97FD8AB76DD5A0" author="Mendoza, J. C. E." box="[377,548,619,645]" pageId="18" pageNumber="494" pagination="374 - 384" refId="ref12717" refString="Mendoza, J. C. E. (2013) A new species of ocellated Xanthias Rathbun, 1897 (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, Philippines. Zootaxa, 3636 (2), 374 - 384. https: // doi. org / 10.11646 / zootaxa. 3636.2.8" type="journal article" year="2013">Mendoza 2013</bibRefCitation>
: fig. 4IJ) immediately proximal to the apical lobe.
</paragraph>
<caption id="DF3C660F7F3BFF98FF76F89BB58FD0FC" ID-DOI="http://doi.org/10.5281/zenodo.13210296" ID-Zenodo-Dep="13210296" httpUri="https://zenodo.org/record/13210296/files/figure.png" pageId="16" pageNumber="492" startId="16.[152,255,1914,1938]" targetBox="[281,1306,597,1890]" targetPageId="16" targetType="figure">
<paragraph id="8BFC36877F3BFF98FF76F89BB58FD0FC" blockId="16.[151,1436,1914,2009]" pageId="16" pageNumber="492">
<emphasis id="B937EA957F3BFF98FF76F89BB451D0B7" bold="true" box="[152,280,1914,1938]" pageId="16" pageNumber="492">FIGURE 9</emphasis>
.
<taxonomicName id="4C434D047F3BFF98FEC7F89BB7D5D0B7" authority="Mendoza, 2013" authorityName="Mendoza" authorityYear="2013" box="[297,668,1914,1938]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="492" phylum="Arthropoda" rank="species" species="joanneae">
<emphasis id="B937EA957F3BFF98FEC7F89BB4A5D0B7" box="[297,492,1914,1938]" italics="true" pageId="16" pageNumber="492">Xanthias joanneae</emphasis>
<bibRefCitation id="EFD24B767F3BFF98FE19F89BB7D5D0B7" author="Mendoza, J. C. E." box="[503,668,1914,1938]" pageId="16" pageNumber="492" pagination="374 - 384" refId="ref12717" refString="Mendoza, J. C. E. (2013) A new species of ocellated Xanthias Rathbun, 1897 (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, Philippines. Zootaxa, 3636 (2), 374 - 384. https: // doi. org / 10.11646 / zootaxa. 3636.2.8" type="journal article" year="2013">Mendoza, 2013</bibRefCitation>
</taxonomicName>
, male (IO/SS/BRC/00197, 6.4 × 4.2 mm), Andaman Sea, India: colour photographs: (A) carapace, dorsal view; (B) mxp3, chelipeds, pereopods, sternum and pleon, ventral view. Scale bars: AB, 2 mm.
</paragraph>
</caption>
<caption id="DF3C660F7F3AFF99FF79F962B409D0DF" ID-DOI="http://doi.org/10.5281/zenodo.13210300" ID-Zenodo-Dep="13210300" httpUri="https://zenodo.org/record/13210300/files/figure.png" pageId="17" pageNumber="493" startId="17.[151,250,1667,1691]" targetBox="[304,1294,191,1633]" targetPageId="17" targetType="figure">
<paragraph id="8BFC36877F3AFF99FF79F962B409D0DF" blockId="17.[151,1437,1667,2042]" pageId="17" pageNumber="493">
<emphasis id="B937EA957F3AFF99FF79F962B452D1BE" bold="true" box="[151,283,1667,1691]" pageId="17" pageNumber="493">FIGURE 10</emphasis>
. First and second gonopods:
<taxonomicName id="4C434D047F3AFF99FDBEF965B6ABD1B9" authority="(Serene, 1971)" baseAuthorityName="Serene" baseAuthorityYear="1971" box="[592,994,1668,1692]" class="Malacostraca" family="Goneplacidae" genus="Singhaplax" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="493" phylum="Arthropoda" rank="species" species="ockelmanni">
<emphasis id="B937EA957F3AFF99FDBEF965B609D1B9" box="[592,832,1668,1692]" italics="true" pageId="17" pageNumber="493">Singhaplax ockelmanni</emphasis>
(
<bibRefCitation id="EFD24B767F3AFF99FCA1F965B693D1B9" author="Serene, R." box="[847,986,1668,1692]" pageId="17" pageNumber="493" pagination="903 - 918" refId="ref14745" refString="Serene, R. (1971) Observations preliminaires sur des brachyoures nouveaux ou mal connus du sud-est asiatique (Crustacea Decapoda). Bulletin du Museum national d'Histoire naturelle, Paris, 2 eme Serie, 42, 903 - 918, 6 pls." type="journal article" year="1971">Serène, 1971</bibRefCitation>
)
</taxonomicName>
, male (IO/SS/BRC/00360, 4.8 × 2.5 mm): (A) entire G1, pleonal view; (B) entire G1, sternal view; (C) entire G2, pleonal view.
<taxonomicName id="4C434D047F3AFF99FBFBF946B59DD1C7" authority="(Calman, 1900)" baseAuthorityName="Calman" baseAuthorityYear="1900" class="Malacostraca" family="Parthenopidae" genus="Pseudolambrus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="493" phylum="Arthropoda" rank="species" species="confragosus">
<emphasis id="B937EA957F3AFF99FBFBF946B07ED19A" box="[1045,1335,1703,1727]" italics="true" pageId="17" pageNumber="493">Pseudolambrus confragosus</emphasis>
(
<bibRefCitation id="EFD24B767F3AFF99FAA8F946B584D1C7" author="Calman, W. T." pageId="17" pageNumber="493" pagination="1 - 50" refId="ref10828" refString="Calman, W. T. (1900) On a collection of Brachyura from Torres Straits. Transactions of the Linnean Society of London, Series 2, Zoology, 8, 1 - 50, pls. 1 - 3. https: // doi. org / 10.1111 / j. 1096 - 3642.1900. tb 00307. x" type="journal article" year="1900">Calman, 1900</bibRefCitation>
)
</taxonomicName>
, male (IO/SS/BRC/00323, 22.2 × 18.0 mm): (D) entire G1, pleonal view; (E) G1 distal portion, pleonal view; (F) G1 distal portion, sternal view; (G) entire G1, sternal view; (H) entire G2, pleonal view.
<taxonomicName id="4C434D047F3AFF99FC00F90CB028D021" authority="(Laurie, 1906)" baseAuthorityName="Laurie" baseAuthorityYear="1906" box="[1006,1377,1772,1797]" class="Malacostraca" family="Portunidae" genus="Cavoportunus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="493" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B937EA957F3AFF99FC00F90CB18ED021" box="[1006,1223,1772,1796]" italics="true" pageId="17" pageNumber="493">Cavoportunus dubius</emphasis>
(
<bibRefCitation id="EFD24B767F3AFF99FB3BF90DB013D021" author="Laurie, R. D." box="[1237,1370,1772,1796]" pageId="17" pageNumber="493" pagination="349 - 432" refId="ref12375" refString="Laurie, R. D. (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.), Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with Supplementary Reports Upon the Marine Biology of Ceylon by Other Naturalists. Part 5. The Royal Society, London, pp. 349 - 432." type="book chapter" year="1906">Laurie, 1906</bibRefCitation>
)
</taxonomicName>
, male (IO/SS/BRC/00191, 26.9 × 20.4 mm): (I) entire G1, pleonal view; (J) entire G1, lateral view; (K) entire G1, mesial view; (L) entire G2, pleonal view.
<taxonomicName id="4C434D047F3AFF99FE76F8D2B60AD06F" authority="(Rathbun, 1906)" baseAuthorityName="Rathbun" baseAuthorityYear="1906" box="[408,835,1842,1866]" class="Malacostraca" family="Xanthidae" genus="Cyrtocarcinus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="493" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B937EA957F3AFF99FE76F8D2B7D8D06F" box="[408,657,1843,1866]" italics="true" pageId="17" pageNumber="493">Cyrtocarcinus truncatus</emphasis>
(
<bibRefCitation id="EFD24B767F3AFF99FD4FF8D3B675D06F" author="Rathbun, M. J." box="[673,828,1842,1866]" pageId="17" pageNumber="493" pagination="827 - 930" refId="ref14062" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. 1 - 24." type="journal article" year="1906">Rathbun, 1906</bibRefCitation>
)
</taxonomicName>
, male (IO/SS/BRC/00193, 8.6 × 6.1 mm): (M) entire G1, pleonal view; (N) entire G1, sternal view; (O) entire G2, sternal view.
<taxonomicName id="4C434D047F3AFF99FC80F8B7B067D048" authority="Mendoza &amp; Ng, 2008" authorityName="Mendoza &amp; Ng" authorityYear="2008" box="[878,1326,1877,1902]" class="Malacostraca" family="Xanthidae" genus="Visayax" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="493" phylum="Arthropoda" rank="species" species="osteodictyon">
<emphasis id="B937EA957F3AFF99FC80F8B7B10AD048" box="[878,1091,1877,1901]" italics="true" pageId="17" pageNumber="493">Visayax osteodictyon</emphasis>
<bibRefCitation id="EFD24B767F3AFF99FBA5F8B4B067D048" author="Mendoza, J. C. E. &amp; Ng, P. K. L." box="[1099,1326,1877,1902]" pageId="17" pageNumber="493" pagination="385 - 404" refId="ref12842" refString="Mendoza, J. C. E. &amp; Ng, P. K. L. (2008) New genera and species of euxanthine crabs (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, the Philippines. Raffles Bulletin of Zoology, 56 (2), 385 - 404." type="journal article" year="2008">Mendoza &amp; Ng, 2008</bibRefCitation>
</taxonomicName>
, male (IO/ SS/BRC/00194, 5.2 × 3.6 mm): (P) entire G1, pleonal view; (Q) G1 distal portion, pleonal view; (R) entire G1, sternal view.
<taxonomicName id="4C434D047F3AFF99FF79F87AB4B0D096" authority="Mendoza, 2013" authorityName="Mendoza" authorityYear="2013" box="[151,505,1947,1971]" class="Malacostraca" family="Xanthidae" genus="Xanthias" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="493" phylum="Arthropoda" rank="species" species="joanneae">
<emphasis id="B937EA957F3AFF99FF79F87AB41DD096" box="[151,340,1947,1971]" italics="true" pageId="17" pageNumber="493">Xanthias joanneae</emphasis>
<bibRefCitation id="EFD24B767F3AFF99FEB4F87AB4B0D096" author="Mendoza, J. C. E." box="[346,505,1947,1971]" pageId="17" pageNumber="493" pagination="374 - 384" refId="ref12717" refString="Mendoza, J. C. E. (2013) A new species of ocellated Xanthias Rathbun, 1897 (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, Philippines. Zootaxa, 3636 (2), 374 - 384. https: // doi. org / 10.11646 / zootaxa. 3636.2.8" type="journal article" year="2013">Mendoza, 2013</bibRefCitation>
</taxonomicName>
, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). (S) entire G1, pleonal view; (T) G1 distal portion, pleonal view; (U) G1 distal portion, sternal view; (V) entire G1, sternal view. Scale bars: AC, PV, 0.2 mm, D, GN, 1 mm, E, F, O, 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BFC36877F39FF9AFF29FD6FB622D5D4" blockId="18.[151,1437,150,753]" pageId="18" pageNumber="494">
<emphasis id="B937EA957F39FF9AFF29FD6FB74DD58D" bold="true" box="[199,516,654,680]" pageId="18" pageNumber="494">Geographical distribution.</emphasis>
<collectingRegion id="4987F8657F39FF9AFDE4FD6EB707D58C" box="[522,590,655,681]" country="Philippines" name="Bohol" pageId="18" pageNumber="494">Bohol</collectingRegion>
Sea,
<collectingCountry id="F35476177F39FF9AFD65FD6EB640D58C" box="[651,777,655,681]" name="Philippines" pageId="18" pageNumber="494">Philippines</collectingCountry>
(
<bibRefCitation id="EFD24B767F39FF9AFCF8FD6EB68AD58C" author="Mendoza, J. C. E." box="[790,963,655,681]" pageId="18" pageNumber="494" pagination="374 - 384" refId="ref12717" refString="Mendoza, J. C. E. (2013) A new species of ocellated Xanthias Rathbun, 1897 (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, Philippines. Zootaxa, 3636 (2), 374 - 384. https: // doi. org / 10.11646 / zootaxa. 3636.2.8" type="journal article" year="2013">Mendoza 2013</bibRefCitation>
), Amami-Oshima Island,
<collectingCountry id="F35476177F39FF9AFB0EFD6EB069D58C" box="[1248,1312,655,681]" name="Japan" pageId="18" pageNumber="494">Japan</collectingCountry>
(
<bibRefCitation id="EFD24B767F39FF9AFADEFD6EB409D5E8" author="Takeda, M. &amp; Komatsu, H." pageId="18" pageNumber="494" pagination="153 - 189" refId="ref15381" refString="Takeda, M. &amp; Komatsu, H. (2018) Offshore Crabs of the Family Xanthidae and Some Related Families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Science Museum, Tokyo, 52, 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu 2018</bibRefCitation>
), and Nicobars,
<collectingCountry id="F35476177F39FF9AFE18FD52B779D5E8" box="[502,560,691,717]" name="India" pageId="18" pageNumber="494">India</collectingCountry>
(present study). The present observation is the first record from Indian Ocean indicating westward extension of the known geographical range.
</paragraph>
</subSubSection>
</treatment>
</document>