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<document id="97CE6366619ED8FF655ADF546CA64C2D" ID-DOI="10.11646/zootaxa.5505.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13745912" ID-ZooBank="93F5E9F6-9EDC-4ED2-8F2B-1E6D0C18F893" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726052234810" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="03E7FF50FFEE5020FF23FB7BFC8346F6" docLanguage="en" docName="zootaxa.5505.1.1.pdf" docOrigin="Zootaxa 5505 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5505.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Anhoraeomorphus diegoi Jałoszyński 2024, comb. n." docType="treatment" docVersion="2" lastPageNumber="33" masterDocId="FFDE8728FFF05000FFB4FFCCFFC54473" masterDocTitle="The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="31" updateTime="1726053143081" updateUser="ExternalLinkService">
<mods:mods id="4AEA666C48FF9A2CAD6ECD3D3EF34D7B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="A34101E20B96D944B9648998157FE3E4">The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:date id="A54726A6DB033DE5A68414AC1B9BD3F1">2024</mods:date>
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<mods:number id="49BE7C3F12E449D86F00977F026342DC">2024-09-04</mods:number>
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<mods:number id="CBF53EADB72B739EFACBDD8FABD2D946">5505</mods:number>
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<mods:identifier id="55FD96B931498A5DD52B89186460D0DA" type="DOI">10.11646/zootaxa.5505.1.1</mods:identifier>
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<treatment id="03E7FF50FFEE5020FF23FB7BFC8346F6" ID-DOI="http://doi.org/10.5281/zenodo.13746045" ID-Zenodo-Dep="13746045" LSID="urn:lsid:plazi:treatment:03E7FF50FFEE5020FF23FB7BFC8346F6" httpUri="http://treatment.plazi.org/id/03E7FF50FFEE5020FF23FB7BFC8346F6" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
<subSubSection id="C3541DCDFFEE501EFF23FB7BFD6E40A1" box="[151,683,1207,1234]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8BF14E46FFEE501EFF23FB7BFD6E40A1" blockId="30.[151,683,1207,1270]" box="[151,683,1207,1234]" pageId="30" pageNumber="31">
<heading id="D0B9F92AFFEE501EFF23FB7BFD6E40A1" bold="true" box="[151,683,1207,1234]" fontSize="11" level="2" pageId="30" pageNumber="31" reason="4">
<emphasis id="B93A9254FFEE501EFF23FB7BFD6E40A1" bold="true" box="[151,683,1207,1234]" pageId="30" pageNumber="31">
<taxonomicName id="4C4E35C5FFEE501EFF23FB7BFDF640A2" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,563,1207,1234]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="diegoi" status="comb. nov.">
<emphasis id="B93A9254FFEE501EFF23FB7BFE0940A1" bold="true" box="[151,460,1207,1234]" italics="true" pageId="30" pageNumber="31">Anhoraeomorphus diegoi</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFEE501EFDF5FB74FD6E40A1" box="[577,683,1208,1234]" pageId="30" pageNumber="31" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEE501EFF23FB17FE854085" box="[151,320,1243,1270]" pageId="30" pageNumber="31" type="description">
<paragraph id="8BF14E46FFEE501EFF23FB17FE854085" blockId="30.[151,683,1207,1270]" box="[151,320,1243,1270]" pageId="30" pageNumber="31">
(
<figureCitation id="137552C3FFEE501EFF2BFB17FEFD4085" box="[159,312,1243,1270]" captionStart="FIGURES 111–118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111–118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Figs 111–118</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEE501EFF23FAD5FD8C4142" box="[151,585,1305,1329]" pageId="30" pageNumber="31" type="reference_group">
<paragraph id="8BF14E46FFEE501EFF23FAD5FD8C4142" blockId="30.[151,585,1305,1329]" box="[151,585,1305,1329]" pageId="30" pageNumber="31">
<treatmentCitationGroup id="AB5E6968FFEE501EFF23FAD5FD8C4142" box="[151,585,1305,1329]" pageId="30" pageNumber="31">
<treatmentCitation id="0AEF6857FFEE501EFF23FAD5FD804142" author="Franz, H." box="[151,581,1305,1329]" page="156" pageId="30" pageNumber="31" year="1986">
<taxonomicName id="4C4E35C5FFEE501EFF23FAD5FD804142" ID-CoL="6MDVK" authority="Franz, 1986 b: 156" authorityName="Franz" authorityPageNumber="156" authorityYear="1986" box="[151,581,1305,1329]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="diegoi">
<emphasis id="B93A9254FFEE501EFF23FAD5FE464142" box="[151,387,1305,1329]" italics="true" pageId="30" pageNumber="31">Horaeomorphus diegoi</emphasis>
<bibRefCitation id="EFDF33B7FFEE501EFE3DFAD6FD804142" author="Franz, H." box="[393,581,1305,1329]" pageId="30" pageNumber="31" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 156</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEE501EFF23FA95FB0742C4" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8BF14E46FFEE501EFF23FA95FD6841C8" blockId="30.[151,1437,1368,2043]" pageId="30" pageNumber="31">
<emphasis id="B93A9254FFEE501EFF23FA95FE754101" bold="true" box="[151,432,1368,1395]" pageId="30" pageNumber="31">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFEE501EFE0AFA95FD6F41C8" collectionCode="MUSEUM, PARIS, BATHIE, NHMW" country="Madagascar" determinerName="H. Franz" location="MADAGASCAR" pageId="30" pageNumber="31" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B93A9254FFEE501EFE0AFA95FD254101" bold="true" box="[446,736,1368,1395]" pageId="30" pageNumber="31">
<typeStatus id="54F5F0E4FFEE501EFE0AFA95FDEE4100" box="[446,555,1369,1395]" pageId="30" pageNumber="31" type="paratype">Paratype</typeStatus>
(Madagascar)
</emphasis>
:
<specimenCount id="9D4885CFFFEE501EFD40FA95FCCE4100" box="[756,779,1369,1395]" count="1" pageId="30" pageNumber="31" type="male">♂</specimenCount>
, three labels: “
<collectionCode id="ED5FD683FFEE501EFC70FA95FB584100" box="[964,1181,1369,1395]" pageId="30" pageNumber="31">MUSEUM PARIS</collectionCode>
/
<collectingCountry id="F3590ED6FFEE501EFB74FA95FA424100" box="[1216,1415,1369,1395]" name="Madagascar" pageId="30" pageNumber="31">MADAGASCAR</collectingCountry>
/ (PERRIER DE LA
<collectionCode id="ED5FD683FFEE501EFE30FAB1FE2941E4" box="[388,492,1405,1431]" pageId="30" pageNumber="31">BATHIE</collectionCode>
)” [bluish, printed], “
<taxonomicName id="4C4E35C5FFEE501EFD56FAB1FC5C41E4" box="[738,921,1405,1431]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ diegoi m. / det.
<determinerName id="7E2DA282FFEE501EFBC0FAB1FB1441E4" box="[1140,1233,1405,1431]" pageId="30" pageNumber="31">H.Franz</determinerName>
/
<typeStatus id="54F5F0E4FFEE501EFB46FAB1FA5641E4" box="[1266,1427,1405,1431]" pageId="30" pageNumber="31" type="paratype">PARATYPUS</typeStatus>
” [yellowish, handwritten and printed] (
<collectionCode id="ED5FD683FFEE501EFD8BFA6DFD6741C8" box="[575,674,1441,1467]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="30" pageNumber="31" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF14E46FFEE501EFF73FA08FB0742C4" blockId="30.[151,1437,1368,2043]" pageId="30" pageNumber="31">
<emphasis id="B93A9254FFEE501EFF73FA08FE5B41AD" bold="true" box="[199,414,1476,1502]" pageId="30" pageNumber="31">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFEE501EFE58FA09FD8341AC" box="[492,582,1477,1503]" captionStart="FIGURES 111–118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111–118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Fig.111</figureCitation>
) slender; head (
<figureCitation id="137552C3FFEE501EFD45FA09FC4D41AC" box="[753,904,1477,1503]" captionStart="FIGURES 111–118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111–118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Figs 112–113</figureCitation>
) round, with vertex posteriorly broadly rounded; pronotum (
<figureCitation id="137552C3FFEE501EFEACFA25FEBC4270" box="[280,377,1513,1539]" captionStart="FIGURES 111–118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111–118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Fig. 111</figureCitation>
) bell-shaped, broadest between middle and anterior third, with one pair of large and deep antebasal pits; antennomeres 3–6 (
<figureCitation id="137552C3FFEE501EFDA8F9C1FDBE4254" box="[540,635,1549,1575]" captionStart="FIGURES 111–118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111–118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Fig. 114</figureCitation>
) each about as long as broad, 7–10 each slightly transverse; profemora slightly thicker than meso- and metafemora; hind tibiae in male modified, strongly thickening to distal third and then narrowing towards apex; aedeagus (
<figureCitation id="137552C3FFEE501EFDD4F999FD3E4203" box="[608,763,1621,1648]" captionStart="FIGURES 111–118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111–118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Figs 115–116</figureCitation>
) in ventral view broadest in sub-basal third, apical region of dorsal wall in lateral view weakly curved dorsally, distolateral endophallic sclerites broadly separated, each elongate and with curved subtriangular apex, each paramere with minute 2–3 apical and subapical setae.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFEE5020FF73F90CFE5B4532" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31" type="description">
<paragraph id="8BF14E46FFEE501EFF73F90CFF0F428C" blockId="30.[151,1437,1368,2043]" pageId="30" pageNumber="31">
<emphasis id="B93A9254FFEE501EFF73F90CFEB042A9" bold="true" box="[199,373,1728,1754]" pageId="30" pageNumber="31">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFEE501EFD96F90DFDBA42A8" box="[546,639,1729,1755]" captionStart="FIGURES 111–118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111–118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Fig. 111</figureCitation>
) flattened, moderately uniformly light brown, setae yellowish; BL
<quantity id="4CB6E3A3FFEE501EFADFF90DFF03428C" metricMagnitude="-3" metricUnit="m" metricValue="1.79" pageId="30" pageNumber="31" unit="mm" value="1.79">1.79 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFEE501EFF73F8C5FA844388" blockId="30.[151,1437,1368,2043]" pageId="30" pageNumber="31">
Head (
<figureCitation id="137552C3FFEE501EFEA0F8C5FEB14350" box="[276,372,1801,1827]" captionStart="FIGURES 111–118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111–118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Fig. 112</figureCitation>
) in anterodorsal view round, broadest at eyes, HL
<quantity id="4CB6E3A3FFEE501EFC0AF8C5FBE34350" box="[958,1062,1801,1827]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="31" unit="mm" value="0.3">0.30 mm</quantity>
, HW
<quantity id="4CB6E3A3FFEE501EFBDFF8C5FB114350" box="[1131,1236,1801,1827]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="31" unit="mm" value="0.3">0.30 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFEE501EFF6AF8E1FEFB4334" box="[222,318,1837,1863]" captionStart="FIGURES 111–118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111–118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Fig. 113</figureCitation>
) subequal to eye; vertex and frons confluent, together weakly convex and about as long as broad; vertex posteriorly broadly rounded; each eye moderately large and weakly convex, oval, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFEE501EFC95F855FC4543C0" box="[801,896,1945,1971]" captionStart="FIGURES 111–118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111–118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="30" pageNumber="31">Fig. 114</figureCitation>
) slender but short, indistinctly shorter than half BL, AnL
<quantity id="4CB6E3A3FFEE501EFEB0F871FEAE43A4" box="[260,363,1981,2007]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="30" pageNumber="31" unit="mm" value="0.8">0.80 mm</quantity>
, antennomeres 1–2 each weakly elongate, 3–6 each about as long as broad, 7–10 each weakly transverse, 11 indistinctly shorter than 9 and 10 combined, as wide as 10, about 1.8 times as long as broad.
</paragraph>
<caption id="DF311ECEFFEF501FFF23FAA3FE4941BC" ID-DOI="http://doi.org/10.5281/zenodo.13745961" ID-Zenodo-Dep="13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,264,1391,1415]" targetBox="[151,1422,181,1357]" targetPageId="31" targetType="figure">
<paragraph id="8BF14E46FFEF501FFF23FAA3FE4941BC" blockId="31.[151,1437,1391,1487]" pageId="31" pageNumber="32">
<emphasis id="B93A9254FFEF501FFF23FAA3FEAD41F4" bold="true" box="[151,360,1391,1415]" pageId="31" pageNumber="32">FIGURES 111–118.</emphasis>
<taxonomicName id="4C4E35C5FFEF501FFEDAFAA3FD0741F4" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[366,706,1391,1415]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="diegoi">
<emphasis id="B93A9254FFEF501FFEDAFAA3FDAA41F4" box="[366,623,1391,1415]" italics="true" pageId="31" pageNumber="32">Anhoraeomorphus diegoi</emphasis>
(Franz)
</taxonomicName>
, paratype. Habitus in dorsal view (
<emphasis id="B93A9254FFEF501FFBABFAA3FB8741F4" bold="true" box="[1055,1090,1391,1415]" pageId="31" pageNumber="32">111</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFEF501FFADCFAA3FA4841F4" bold="true" box="[1384,1421,1391,1415]" pageId="31" pageNumber="32">112</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFEF501FFCD1FA5FFC4F41D8" bold="true" box="[869,906,1427,1451]" pageId="31" pageNumber="32">113</emphasis>
); antenna (
<emphasis id="B93A9254FFEF501FFBB4FA5FFBE041D8" bold="true" box="[1024,1061,1427,1451]" pageId="31" pageNumber="32">114</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFEF501FFAA5FA5FFAAD41D8" bold="true" box="[1297,1384,1427,1451]" pageId="31" pageNumber="32">115, 117</emphasis>
) and lateral (
<emphasis id="B93A9254FFEF501FFF51FA7BFEF941BC" bold="true" box="[229,316,1463,1487]" pageId="31" pageNumber="32">116, 118</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFEF501FFF73F9EEFF3542B8" blockId="31.[151,1436,1569,2028]" pageId="31" pageNumber="32">
Pronotum bell-shaped, broadest between middle and anterior third; PL
<quantity id="4CB6E3A3FFEF501FFC6FF9EEFB85424F" box="[987,1088,1570,1596]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="31" pageNumber="32" unit="mm" value="0.48">0.48 mm</quantity>
, PW
<quantity id="4CB6E3A3FFEF501FFBCEF9EEFB25424F" box="[1146,1248,1570,1596]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="31" pageNumber="32" unit="mm" value="0.4">0.40 mm</quantity>
. Anterior margin strongly rounded, lateral margins rounded in anterior half, strongly sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of large and deep pits, sublateral carinae indiscernible. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFEF501FFF73F919FA944344" blockId="31.[151,1436,1569,2028]" pageId="31" pageNumber="32">
Elytra together oval, broadest distinctly in front of middle; EL
<quantity id="4CB6E3A3FFEF501FFC39F91AFC314283" box="[909,1012,1750,1776]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="31" pageNumber="32" unit="mm" value="0.93">0.93 mm</quantity>
, EW
<quantity id="4CB6E3A3FFEF501FFB87F919FB5F4283" box="[1075,1178,1749,1776]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="31" pageNumber="32" unit="mm" value="0.6">0.60 mm</quantity>
, EI 1.54; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures very dense but extremely shallow and diffuse, so that surface of elytra appears slightly less glossy than pronotum; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFEF501FFF73F88EFED743D0" blockId="31.[151,1436,1569,2028]" pageId="31" pageNumber="32">Legs moderately long, slender, unmodified except all tibiae, which are thick and slightly recurved, fore tibiae with large distomesal concave patch of dense adhesive setae, metatibiae strongly broadening to distal third and then narrowing.</paragraph>
<paragraph id="8BF14E46FFEF5020FF73F861FA40456E" blockId="31.[151,1436,1569,2028]" lastBlockId="32.[151,1437,151,645]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
Aedeagus (
<figureCitation id="137552C3FFEF501FFEFCF861FE2343BB" box="[328,486,1965,1992]" captionStart="FIGURES 111–118" captionStartId="31.[151,264,1391,1415]" captionTargetBox="[151,1422,181,1357]" captionTargetId="figure-16@31.[151,1435,181,1367]" captionTargetPageId="31" captionText="FIGURES 111–118. Anhoraeomorphus diegoi (Franz), paratype. Habitus in dorsal view (111); head in anterodorsal view (112); head and prothorax in lateral view (only bristles on head are shown) (113); antenna (114); aedeagus in ventral (115, 117) and lateral (116, 118) views." figureDoi="http://doi.org/10.5281/zenodo.13745961" httpUri="https://zenodo.org/record/13745961/files/figure.png" pageId="31" pageNumber="32">Figs 115–118</figureCitation>
) slender, AeL
<quantity id="4CB6E3A3FFEF501FFD27F862FD3B43BB" box="[659,766,1966,1992]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="31" pageNumber="32" unit="mm" value="0.28">0.28 mm</quantity>
; in ventral view broadest in sub-basal third, drop-shaped, distally gradually narrowing to broadly subtriangular apex with weakly marked obtusely angulate tip; in lateral view apical region of dorsal wall weakly curved dorsally; endophallus with nearly symmetrical and conspicuously large distolateral sclerites and with strongly asymmetrical assemblage of median sclerotized structures, distolateral sclerites broadly separated, each with long subtriangular and curved apical region; parameres broad and their apices not projecting beyond aedeagal apex, each paramere with apical-subapical group of 2–3 barely discernible setae.
</paragraph>
<paragraph id="8BF14E46FFD05020FF73FEEBFE5B4532" blockId="32.[151,1437,151,645]" box="[199,414,295,321]" pageId="32" pageNumber="33">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD05020FF73FE86FC884516" box="[199,845,330,357]" pageId="32" pageNumber="33" type="distribution">
<paragraph id="8BF14E46FFD05020FF73FE86FC884516" blockId="32.[151,1437,151,645]" box="[199,845,330,357]" pageId="32" pageNumber="33">
<emphasis id="B93A9254FFD05020FF73FE86FEA44517" bold="true" box="[199,353,330,356]" pageId="32" pageNumber="33">Distribution.</emphasis>
Northernmost
<collectingCountry id="F3590ED6FFD05020FDBEFE87FD544516" box="[522,657,331,357]" name="Madagascar" pageId="32" pageNumber="33">Madagascar</collectingCountry>
(Diana Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD05020FF73FEA3FC8346F6" pageId="32" pageNumber="33" type="discussion">
<paragraph id="8BF14E46FFD05020FF73FEA3FC7345A2" blockId="32.[151,1437,151,645]" pageId="32" pageNumber="33">
<emphasis id="B93A9254FFD05020FF73FEA3FEFE45FA" bold="true" box="[199,315,367,393]" pageId="32" pageNumber="33">Remarks.</emphasis>
Males of
<taxonomicName id="4C4E35C5FFD05020FE1FFEA3FD0D45FA" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[427,712,367,393]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="diegoi">
<emphasis id="B93A9254FFD05020FE1FFEA3FD0D45FA" box="[427,712,367,393]" italics="true" pageId="32" pageNumber="33">Anhoraeomorphus diegoi</emphasis>
</taxonomicName>
have the hind tibiae similarly modified as those in
<taxonomicName id="4C4E35C5FFD05020FAB0FEA3FA5945FB" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[1284,1436,367,393]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="anosyensis">
<emphasis id="B93A9254FFD05020FAB0FEA3FA5945FB" box="[1284,1436,367,393]" italics="true" pageId="32" pageNumber="33">A. anosyensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4E35C5FFD05020FF73FE5FFEB245DF" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[199,375,403,429]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="roussettensis">
<emphasis id="B93A9254FFD05020FF73FE5FFEB245DF" box="[199,375,403,429]" italics="true" pageId="32" pageNumber="33">A. roussettensis</emphasis>
</taxonomicName>
. However, the body of
<taxonomicName id="4C4E35C5FFD05020FD35FE5FFD2245DE" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[641,743,403,429]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="diegoi">
<emphasis id="B93A9254FFD05020FD35FE5FFD2245DE" box="[641,743,403,429]" italics="true" pageId="32" pageNumber="33">A. diegoi</emphasis>
</taxonomicName>
is slenderer and the aedeagus has the distolateral endophallic sclerites much broader than those in
<taxonomicName id="4C4E35C5FFD05020FD86FE7BFD0F45A3" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[562,714,439,465]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="anosyensis">
<emphasis id="B93A9254FFD05020FD86FE7BFD0F45A3" box="[562,714,439,465]" italics="true" pageId="32" pageNumber="33">A. anosyensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4E35C5FFD05020FCB5FE7BFC7545A3" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[769,944,439,465]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="roussettensis">
<emphasis id="B93A9254FFD05020FCB5FE7BFC7545A3" box="[769,944,439,465]" italics="true" pageId="32" pageNumber="33">A. roussettensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF14E46FFD05020FF73FE17FC8346F6" blockId="32.[151,1437,151,645]" pageId="32" pageNumber="33">
<materialsCitation id="3B26441BFFD05020FF73FE17FC8746F6" collectionCode="NHMW" collectorName="Charles Alluaud" location="Antsiranana" pageId="32" pageNumber="33" specimenCount="6" stateProvince="Antsiranana" typeStatus="holotype">
<bibRefCitation id="EFDF33B7FFD05020FF73FE17FEA04586" author="Franz, H." box="[199,357,475,501]" pageId="32" pageNumber="33" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
mentions
<specimenCount id="9D4885CFFFD05020FE62FE17FD494586" box="[470,652,475,501]" count="3" pageId="32" pageNumber="33" type="generic">three specimens</specimenCount>
, all males, included in the type series. The
<typeStatus id="54F5F0E4FFD05020FBEAFE17FB054586" box="[1118,1216,475,501]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
comes from Diego-Suarez (currently
<collectingRegion id="498A80A4FFD05020FED0FE33FE28466A" box="[356,493,511,537]" country="Madagascar" name="Antsiranana" pageId="32" pageNumber="33">Antsiranana</collectingRegion>
) and was collected by
<collectorName id="26BB2B90FFD05020FD4FFE33FC73466A" box="[763,950,511,537]" pageId="32" pageNumber="33">Charles Alluaud</collectorName>
,
<specimenCount id="9D4885CFFFD05020FC70FE33FBA6466A" box="[964,1123,511,537]" count="2" pageId="32" pageNumber="33" type="generic" typeStatus="paratype">two paratypes</specimenCount>
are from Léon Fairmaire’s collection and their collecting site is unknown. The
<typeStatus id="54F5F0E4FFD05020FD57FDEFFC80464E" box="[739,837,547,573]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D4885CFFFD05020FCCAFDEFFBCB464E" box="[894,1038,547,573]" count="1" pageId="32" pageNumber="33" type="generic" typeStatus="paratype">one paratype</specimenCount>
are deposited in Muséum National d'Histoire Naturelle, Paris and were not available for my study. The examined
<typeStatus id="54F5F0E4FFD05020FB9CFD8BFB4D4612" box="[1064,1160,583,609]" pageId="32" pageNumber="33" type="paratype">paratype</typeStatus>
male (
<collectionCode id="ED5FD683FFD05020FB6EFD8BFAFC4612" box="[1242,1337,583,609]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="32" pageNumber="33" type="Museum">NHMW</collectionCode>
) has the aedeagus matching the illustration in the original description
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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