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<mods:title id="67C8EF95B3E63D866AB4DDAAE2557254">A preliminary review of the fossil species of Ranina Lamarck, 1801 (Decapoda, Brachyura, Raninidae), with systematic remarks</mods:title>
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<paragraph id="8BDB32197B186B76FFE0FF118601A1C4" blockId="12.[113,576,135,192]" box="[113,576,135,162]" pageId="12" pageNumber="55">
?
<taxonomicName id="4C64499A7B186B76FFEFFF118601A1C4" ID-CoL="BTSJJ" authority="Van Straelen, 1924" authorityName="Van Straelen" authorityYear="1924" box="[126,576,135,162]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="molengraaffi">
<emphasis id="B910EE0B7B186B76FFEFFF118524A1C7" box="[126,357,135,161]" italics="true" pageId="12" pageNumber="55">Ranina molengraaffi</emphasis>
<bibRefCitation id="EFF54FE87B186B76FEFAFF118601A1C4" author="Van Straelen V." box="[363,576,135,162]" pageId="12" pageNumber="55" pagination="489 - 492" refId="ref25213" refString="Van Straelen V., 1924 - Description de Crustaces decapodes nouveaux des terrains tertiares de Borneo. Koninklijke Akademie van Wetenschappen te Amsterdam, Proceedings, 26: 489 - 492." type="journal article" year="1924">Van Straelen, 1924</bibRefCitation>
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<figureCitation id="135F2E9C7B186B76FFE0FF3F84F2A1A6" box="[113,179,166,192]" captionStart="Fig" captionStartId="12.[114,147,1914,1936]" captionTargetBox="[194,679,1171,1867]" captionTargetId="figure-728@12.[187,681,1164,1874]" captionTargetPageId="12" captionText="Fig. 9 -?Ranina molengraaffi Van Straelen, 1924; Van Straelens original figures (Fig. 1 A, B)." figureDoi="http://doi.org/10.5281/zenodo.12753823" httpUri="https://zenodo.org/record/12753823/files/figure.png" pageId="12" pageNumber="55">Fig. 9</figureCitation>
</paragraph>
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<paragraph id="8BDB32197B186B76FFE0FF7A8494A07C" blockId="12.[113,757,226,282]" pageId="12" pageNumber="55">
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<treatmentCitation id="0AC514087B186B76FFE0FF7A8685A19A" author="Van Straelen V." box="[113,708,226,252]" page="777" pageId="12" pageNumber="55" year="1924">
<taxonomicName id="4C64499A7B186B76FFE0FF7A8685A19A" ID-CoL="BTSJJ" authority="Van Straelen, 1924: 777" authorityName="Van Straelen" authorityPageNumber="777" authorityYear="1924" box="[113,708,226,252]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="molengraaffi" subGenus="Hela">
<emphasis id="B910EE0B7B186B76FFE0FF7A8483A19A" box="[113,194,227,252]" italics="true" pageId="12" pageNumber="55">Ranina</emphasis>
(
<emphasis id="B910EE0B7B186B76FF42FF7B8548A19A" box="[211,265,226,252]" italics="true" pageId="12" pageNumber="55">Hela</emphasis>
)
<emphasis id="B910EE0B7B186B76FE8AFF7B85EBA19A" box="[283,426,226,252]" italics="true" pageId="12" pageNumber="55">molengraaffi</emphasis>
<bibRefCitation id="EFF54FE87B186B76FE20FF7B8685A19A" author="Van Straelen V." box="[433,708,226,252]" pageId="12" pageNumber="55" pagination="489 - 492" refId="ref25213" refString="Van Straelen V., 1924 - Description de Crustaces decapodes nouveaux des terrains tertiares de Borneo. Koninklijke Akademie van Wetenschappen te Amsterdam, Proceedings, 26: 489 - 492." type="journal article" year="1924">Van Straelen, 1924: 777</bibRefCitation>
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, fig. 1a, b.
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<taxonomicName id="4C64499A7B186B76FFE0FEA48518A030" ID-CoL="BTSJJ" authorityName="Van Straelen" authorityYear="1924" box="[113,345,316,342]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="molengraaffi">
<emphasis id="B910EE0B7B186B76FFE0FEA48518A030" box="[113,345,316,342]" italics="true" pageId="12" pageNumber="55">Ranina molengraaffi</emphasis>
</taxonomicName>
-
<treatmentCitation id="0AC514087B186B76FEE2FEA5861FA030" author="Glaessner M. F." box="[371,606,316,342]" page="362" pageId="12" pageNumber="55" year="1929">
<bibRefCitation id="EFF54FE87B186B76FEE2FEA5861FA030" author="Glaessner M. F." box="[371,606,316,342]" pageId="12" pageNumber="55" pagination="1 - 464" refId="ref23938" refString="Glaessner M. F., 1929 - Crustacea Decapoda. In: Fossilium Catalogus. I: Animalia. Pompeckj F. J. (ed.). W. Junk, Berlin, 41: 1 - 464." type="journal article" year="1929">Glaessner 1929: 362</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AC514087B186B76FD10FEA48512A013" author="De Angeli A. &amp; Dall'Igna G. L. &amp; Ceccon L." page="120" pageId="12" pageNumber="55" year="2009">
<bibRefCitation id="EFF54FE87B186B76FD10FEA48512A013" author="De Angeli A. &amp; Dall'Igna G. L. &amp; Ceccon L." pageId="12" pageNumber="55" pagination="119 - 122" refId="ref23670" refString="De Angeli A., Dall'Igna G. L. &amp; Ceccon L., 2009 - Segnalazione di Ranina sp. (Decapoda, Brachyura, Raninidae) dell'Eocene superiore di Monte Malo (Vicenza, Italia settentrionale). Lavori - Societa Veneziana di Scienze Naturali, 34: 119 - 122." type="journal article" year="2009">
De Angeli
<emphasis id="B910EE0B7B186B76FF06FEC2848AA012" box="[151,203,346,372]" italics="true" pageId="12" pageNumber="55">et al</emphasis>
. 2009: 120
</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AC514087B186B76FEE8FEC386ECA013" author="Schweitzer C. &amp; Feldmann R. M. &amp; Garassino A. &amp; Karasawa H. &amp; Schweigert G." box="[377,685,346,373]" page="74" pageId="12" pageNumber="55" year="2010">
<bibRefCitation id="EFF54FE87B186B76FEE8FEC386ECA013" author="Schweitzer C. &amp; Feldmann R. M. &amp; Garassino A. &amp; Karasawa H. &amp; Schweigert G." box="[377,685,346,373]" pageId="12" pageNumber="55" pagination="1 - 222" refId="ref24966" refString="Schweitzer C. E, Feldmann R. M., Garassino A., Karasawa H. &amp; Schweigert G., 2010 - Systematic list of fossil decapod crustacean species. Crustaceana Monographs, 10: 1 - 222." type="journal article" year="2010">
Schweitzer
<emphasis id="B910EE0B7B186B76FE6EFEC2867AA012" box="[511,571,346,372]" italics="true" pageId="12" pageNumber="55">et al.</emphasis>
2010: 74
</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AC514087B186B76FD42FEC28592A0F5" author="De Angeli A. &amp; Beschin C." page="13" pageId="12" pageNumber="55" year="2011">
<bibRefCitation id="EFF54FE87B186B76FD42FEC28592A0F5" author="De Angeli A. &amp; Beschin C." pageId="12" pageNumber="55" pagination="11 - 20" refId="ref23611" refString="De Angeli A. &amp; Beschin C., 2011 - Il genere Ranina Lamarck, 1801 (Crustacea, Decapoda, Raninidae) nel Terziario del Vicentino, con descrizione di due nuove specie. Studi e Ricerche - Associazione Amici del Museo - Museo Civico &quot; G. Zannato &quot;, 18: 11 - 20." type="book chapter" year="2011">De Angeli &amp; Beschin 2011: 13</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AC514087B186B76FE64FEE08485A0D7" author="Van Bakel W. M. &amp; Guinot D. &amp; Artal P. &amp; Fraaije R. H. B. &amp; Jagt J. W. M." page="209" pageId="12" pageNumber="55" year="2012">
<bibRefCitation id="EFF54FE87B186B76FE64FEE08485A0D7" author="Van Bakel W. M. &amp; Guinot D. &amp; Artal P. &amp; Fraaije R. H. B. &amp; Jagt J. W. M." pageId="12" pageNumber="55" pagination="1 - 216" refId="ref25151" refString="Van Bakel W. M., Guinot D., Artal P., Fraaije R. H. B. &amp; Jagt J. W. M., 2012 - A revision of the Palaeocorystoidea and the phylogeny of raninoidian crabs (Crustacea, Decapoda, Brachyura, Podotremata). Zootaxa, 3215: 1 - 216." type="journal article" year="2012">
Van Bakel
<emphasis id="B910EE0B7B186B76FDE3FEE086EDA0F4" box="[626,684,376,402]" italics="true" pageId="12" pageNumber="55">et al.</emphasis>
2012: 209
</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AC514087B186B76FF75FE0E85BFA0D7" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M. &amp; Luque J." box="[228,510,406,433]" page="45" pageId="12" pageNumber="55" year="2014">
<bibRefCitation id="EFF54FE87B186B76FF75FE0E85BFA0D7" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M. &amp; Luque J." box="[228,510,406,433]" pageId="12" pageNumber="55" pagination="216 - 272" refId="ref24222" refString="Karasawa H., Schweitzer C. E., Feldmann R. M. &amp; Luque J., 2014 - Phylogeny and classification of Raninoida (Decapoda: Brachyura). Journal of Crustacean Biology, 34 (2): 216 - 272." type="journal article" year="2014">
Karasawa
<emphasis id="B910EE0B7B186B76FECBFE0E85D3A0D7" box="[346,402,407,433]" italics="true" pageId="12" pageNumber="55">et al.</emphasis>
2014: 45
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</heading>
</paragraph>
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<treatmentCitationGroup id="AB7415377B186B76FFE0FE4A854EA36D" pageId="12" pageNumber="55">
<taxonomicName id="4C64499A7B186B76FFE0FE4A8500A08B" authority="Lőrenthey in Lőrenthey &amp; Beurlen 1929: 109" box="[113,321,467,493]" class="Malacostraca" family="Unciolidae" genus="Hela" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="molengraaffi">
<emphasis id="B910EE0B7B186B76FFE0FE4A8500A08B" box="[113,321,467,493]" italics="true" pageId="12" pageNumber="55">Hela molengraaffi</emphasis>
</taxonomicName>
- Lőrenthey
<emphasis id="B910EE0B7B186B76FE4CFE4D85B2A08B" box="[477,499,468,493]" italics="true" pageId="12" pageNumber="55">in</emphasis>
<treatmentCitation id="0AC514087B186B76FE6CFE4A854AA36D" author="Lorenthey E. &amp; Beurlen K." page="109" pageId="12" pageNumber="55" year="1929">
<bibRefCitation id="EFF54FE87B186B76FE6CFE4A854AA36D" author="Lorenthey E. &amp; Beurlen K." pageId="12" pageNumber="55" pagination="1 - 421" refId="ref24319" refString="Lorenthey E. &amp; Beurlen K., 1929 - Die Fossilen Deka- poden der Lander der Ungarischen Krone. Geologica Hungarica (Palaeontologica), 3: 1 - 421." type="journal article" year="1929">Lőrenthey &amp; Beurlen 1929: 109</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C37E61927B186B76FF06FDB486A4A2A9" pageId="12" pageNumber="55" type="materials_examined">
<paragraph id="8BDB32197B186B76FF06FDB48513A213" blockId="12.[113,756,557,1095]" pageId="12" pageNumber="55">
<emphasis id="B910EE0B7B186B76FF06FDB48504A321" bold="true" box="[151,325,557,583]" pageId="12" pageNumber="55">
<typeStatus id="54DF8CBB7B186B76FF06FDB48493A321" box="[151,210,557,583]" pageId="12" pageNumber="55">Type</typeStatus>
material
</emphasis>
: the specimen N° 6 of the sample K.A. 6491 was housed in the Geological Museum of the
<location id="8EBB64C27B186B76FFE0FDF385D2A3E2" LSID="urn:lsid:plazi:treatment:03CD830F7B186B76FFE0FF11872FA413:8EBB64C27B186B76FFE0FDF385D2A3E2" box="[113,403,618,644]" country="Netherlands" municipality="Technology in Delft" name="University of Technology" pageId="12" pageNumber="55">
University of
<collectingMunicipality id="6BBFA8637B186B76FE9EFDF385B5A3E2" box="[271,500,618,644]" pageId="12" pageNumber="55">Technology in Delft</collectingMunicipality>
</location>
(
<collectingCountry id="F37372897B186B76FD95FDF386FDA3E3" box="[516,700,618,645]" name="Netherlands" pageId="12" pageNumber="55">the Netherlands</collectingCountry>
), but nowadays the KA collection is housed in Leiden in the Naturalis Biodiversity Center (
<collectorName id="269157CF7B186B76FE42FD3F86F5A3A6" box="[467,692,678,704]" pageId="12" pageNumber="55">Maaike van Tooren</collectorName>
pers. comm., 2016); the fossil crustacean collection of the Naturalis Biodiversity Center wont be available until 2018 and unfortunately
<collectorName id="269157CF7B186B76FED5FC9885A6A27C" box="[324,487,768,795]" pageId="12" pageNumber="55">Van Straelens</collectorName>
specimen does not appear in the digital records of the invertebrate fossil collection because it is not digitized yet (
<collectorName id="269157CF7B186B76FD8DFCA586B5A230" box="[540,756,828,854]" pageId="12" pageNumber="55">Natasja den Ouden</collectorName>
pers. comm., 2016).
</paragraph>
<paragraph id="8BDB32197B186B76FF06FCE086E4A2F5" blockId="12.[113,756,557,1095]" box="[151,677,888,915]" pageId="12" pageNumber="55">
<materialsCitation id="3B0C38447B186B76FF06FCE086E0A2F5" box="[151,673,888,915]" location="Toehoep River Valley" pageId="12" pageNumber="55" specimenCount="1" typeStatus="holotype">
<emphasis id="B910EE0B7B186B76FF06FCE08571A2F4" bold="true" box="[151,304,888,915]" pageId="12" pageNumber="55">
<typeStatus id="54DF8CBB7B186B76FF06FCE08493A2F5" box="[151,210,889,915]" pageId="12" pageNumber="55">Type</typeStatus>
locality
</emphasis>
:
<location id="8EBB64C27B186B76FEAFFCE08673A2F5" LSID="urn:lsid:plazi:treatment:03CD830F7B186B76FFE0FF11872FA413:8EBB64C27B186B76FEAFFCE08673A2F5" box="[318,562,889,915]" name="Toehoep River Valley" pageId="12" pageNumber="55">Toehoep River Valley</location>
(Borneo)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDB32197B186B76FF06FC0E85FCA2D7" blockId="12.[113,756,557,1095]" box="[151,445,919,945]" pageId="12" pageNumber="55">
<emphasis id="B910EE0B7B186B76FF06FC0E8505A2D7" bold="true" box="[151,324,919,945]" pageId="12" pageNumber="55">Geological age</emphasis>
: Miocene.
</paragraph>
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<emphasis id="B910EE0B7B186B76FF06FC2C853CA2A9" bold="true" box="[151,381,949,975]" pageId="12" pageNumber="55">Examined material</emphasis>
: Van Straelens original figures.
</paragraph>
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<paragraph id="8BDB32197B186B76FF06FC4A8031A39A" blockId="12.[113,756,557,1095]" lastBlockId="12.[832,1475,135,1397]" pageId="12" pageNumber="55">
<emphasis id="B910EE0B7B186B76FF06FC4A8672A28B" bold="true" box="[151,563,979,1005]" pageId="12" pageNumber="55">
Description by
<bibRefCitation id="EFF54FE87B186B76FEDBFC4A8672A28B" author="Van Straelen V." box="[330,563,979,1005]" pageId="12" pageNumber="55" pagination="489 - 492" refId="ref25213" refString="Van Straelen V., 1924 - Description de Crustaces decapodes nouveaux des terrains tertiares de Borneo. Koninklijke Akademie van Wetenschappen te Amsterdam, Proceedings, 26: 489 - 492." type="journal article" year="1924">Van Straelen (1924)</bibRefCitation>
</emphasis>
: “
<emphasis id="B910EE0B7B186B76FDDCFC4D86B5A521" italics="true" pageId="12" pageNumber="55">Le céphalothorax dont la longueur dépasse la largeur denviron 1/5 selargit de larrière vers lavant; sa largeur mesurée au niveau de linsertion des deux dents marginales et celle</emphasis>
<emphasis id="B910EE0B7B186B76FCD1FF118025A39A" italics="true" pageId="12" pageNumber="55">measurée au bord posterieur, sont dans le rapport de 3 à 2; le céphalothorax est bombé, la courbure saccentuant dans la region médiane, au point de constituer une crête surbaissée; la region frontale sincurve vers le haut, de sorte quelle semble précedée par une faible depression; une autre depression plus forte que la précédente, exsite dans la region médiane du cephalothorax, au tiers postérieur; la region cardiaque est indiquée par une paire de sillons an arc de cerle, à concavité ouverte vers les bords latéraux; le bord frontal sensiblement rectiligne est occupé par un rostre triangulaire, large et long, bordé par des échancrures oculaires limitées chacune latéralement par un lobe triangulaire à base très large; au delà de ces lobes, se trouvent deux petites épines et enfin une forte dent affilée et incurvée extérieurement, constituant le prologement des bords latéraux; ceux-ci sont un peu incurves et à angle droit avec le bord postérieur; ce dernier est à peu près rectiligne et bordé par un etroit sillon; le test paraissant lisse, est garni de fines granulations, légèrement acuminées, disposées sans ordre apparent. La face sternale nest pas connue.</emphasis>
</paragraph>
<paragraph id="8BDB32197B186B76FCF4FC9980B5A521" blockId="12.[832,1475,135,1397]" pageId="12" pageNumber="55">Literal translation: carapace longer than wide; carapace convex, mainly in median region, forming a ridge; frontal region curved upward, forming a weak depression; strong depression in median region of carapace; cardiac region marked by two grooves; frontal region with a triangular rostrum wide and long; orbital incisions limited by a triangular lobe laterally; lateral margins with two small spines and one strong outward-directed spine; lateral margin almost straight; posterior margin straight and rimmed; dorsal surface of carapace with granules, slightly pointed, arranged randomly; sternum unknown.</paragraph>
</subSubSection>
<subSubSection id="C37E61927B186B76FCF4FBD2872FA413" pageId="12" pageNumber="55" type="discussion">
<paragraph id="8BDB32197B186B76FCF4FBD2872FA413" blockId="12.[832,1475,135,1397]" pageId="12" pageNumber="55">
<emphasis id="B910EE0B7B186B76FCF4FBD287A2A503" bold="true" box="[869,995,1099,1125]" pageId="12" pageNumber="55">Discussion</emphasis>
. Based upon the description of
<bibRefCitation id="EFF54FE87B186B76FADDFBD587F4A5E2" author="Van Straelen V." pageId="12" pageNumber="55" pagination="489 - 492" refId="ref25213" refString="Van Straelen V., 1924 - Description de Crustaces decapodes nouveaux des terrains tertiares de Borneo. Koninklijke Akademie van Wetenschappen te Amsterdam, Proceedings, 26: 489 - 492." type="journal article" year="1924">Van Straelen (1924)</bibRefCitation>
and relative figures, the species cannot be assigned to
<emphasis id="B910EE0B7B186B76FC3CFB108045A5C4" box="[941,1028,1161,1186]" italics="true" pageId="12" pageNumber="55">
<taxonomicName id="4C64499A7B186B76FC3CFB108040A5C4" box="[941,1025,1161,1186]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="55" phylum="Arthropoda" rank="genus">Ranina</taxonomicName>
,
</emphasis>
lacking the diagnostic characters of the carapace of this genus, as remarked by Van Straelen who pointed out also some general affinities with
<taxonomicName id="4C64499A7B186B76FAADFB5C87FBA59A" authority="De Haan 1841" authorityName="De Haan" authorityYear="1841" class="Malacostraca" family="Raninidae" genus="Notopus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B910EE0B7B186B76FAADFB5C81D8A5B8" box="[1340,1433,1221,1246]" italics="true" pageId="12" pageNumber="55">Notopus</emphasis>
De Haan 1841
</taxonomicName>
, “…
<emphasis id="B910EE0B7B186B76FC60FB7B8734A47C" italics="true" pageId="12" pageNumber="55">for shape and ornamentation of the carapace</emphasis>
…” (
<bibRefCitation id="EFF54FE87B186B76FC3AFA9980FEA47C" author="Van Straelen V." box="[939,1215,1280,1306]" pageId="12" pageNumber="55" pagination="489 - 492" refId="ref25213" refString="Van Straelen V., 1924 - Description de Crustaces decapodes nouveaux des terrains tertiares de Borneo. Koninklijke Akademie van Wetenschappen te Amsterdam, Proceedings, 26: 489 - 492." type="journal article" year="1924">Van Straelen, 1924: 778</bibRefCitation>
). In conclusion, the inability to recognize the diagnostic characters of
<taxonomicName id="4C64499A7B186B76FAE0FA868183A45E" box="[1393,1474,1311,1336]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B910EE0B7B186B76FAE0FA868183A45E" box="[1393,1474,1311,1336]" italics="true" pageId="12" pageNumber="55">Ranina</emphasis>
</taxonomicName>
questions the placement of
<taxonomicName id="4C64499A7B186B76FB11FAA5814EA430" box="[1152,1295,1340,1366]" class="Malacostraca" family="Unciolidae" genus="Hela" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="molengraaffi">
<emphasis id="B910EE0B7B186B76FB11FAA5814EA430" box="[1152,1295,1340,1366]" italics="true" pageId="12" pageNumber="55">molengraaffi</emphasis>
</taxonomicName>
within this genus.
</paragraph>
</subSubSection>
</treatment>
</document>