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<document id="19A3BD15565CE2CA3694CF0AA9D42841" ID-DOI="10.4081/nhs.2017.310" ID-ISSN="2385-0922" ID-Zenodo-Dep="12753805" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1721184724886" checkinUser="felipe" docAuthor="Pasini, Giovanni &amp; Garassino, Alessandro" docDate="2017" docId="03CD830F7B0E6B60FFE0FE688079A2A9" docLanguage="en" docName="NatHistSci.4.1.43-72.pdf" docOrigin="Natural History Sciences 4 (1)" docSource="http://dx.doi.org/10.4081/nhs.2017.310" docStyle="DocumentStyle:926963A4B76E17DFFD817EF231EB8DA9.3:NatHistSci.2014-.journal_article" docStyleId="926963A4B76E17DFFD817EF231EB8DA9" docStyleName="NatHistSci.2014-.journal_article" docStyleVersion="3" docTitle="Ranina burleighensis Holland &amp; Cvancara 1958" docType="treatment" docVersion="1" lastPageNumber="69" masterDocId="FFF4FB777B146B7AFF91FF998441A166" masterDocTitle="A preliminary review of the fossil species of Ranina Lamarck, 1801 (Decapoda, Brachyura, Raninidae), with systematic remarks" masterLastPageNumber="72" masterPageNumber="43" pageNumber="69" updateTime="1726747683517" updateUser="guilherme" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="2C5C22FB93CE04AE638B2B14CF8B2530">A preliminary review of the fossil species of Ranina Lamarck, 1801 (Decapoda, Brachyura, Raninidae), with systematic remarks</mods:title>
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<treatment id="03CD830F7B0E6B60FFE0FE688079A2A9" LSID="urn:lsid:plazi:treatment:03CD830F7B0E6B60FFE0FE688079A2A9" httpUri="http://treatment.plazi.org/id/03CD830F7B0E6B60FFE0FE688079A2A9" lastPageNumber="69" pageId="26" pageNumber="69">
<subSubSection id="C37E61927B0E6B60FFE0FE6884E8A34C" pageId="26" pageNumber="69" type="nomenclature">
<paragraph id="8BDB32197B0E6B60FFE0FE6884E8A34C" blockId="26.[113,756,497,583]" pageId="26" pageNumber="69">
“
<taxonomicName id="4C64499A7B0E6B60FFEFFE6B853BA36D" ID-CoL="9ZPLH" authority="Holland in Holland &amp; Cvancara, 1958" authorityName="Holland &amp; Cvancara" authorityYear="1958" box="[126,378,497,523]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="69" phylum="Arthropoda" rank="species" species="burleighensis">
<emphasis id="B910EE0B7B0E6B60FFEFFE6B848EA36D" box="[126,207,498,523]" italics="true" pageId="26" pageNumber="69">Ranina</emphasis>
”
<emphasis id="B910EE0B7B0E6B60FF72FE68853BA36D" box="[227,378,497,523]" italics="true" pageId="26" pageNumber="69">burleighensis</emphasis>
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<collectingCountry id="F37372897B0E6B60FE13FE68859DA36D" box="[386,476,497,523]" name="Netherlands" pageId="26" pageNumber="69">Holland</collectingCountry>
<emphasis id="B910EE0B7B0E6B60FE74FE6B85BAA36D" box="[485,507,498,523]" italics="true" pageId="26" pageNumber="69">in</emphasis>
<bibRefCitation id="EFF54FE87B0E6B60FD92FE6884E8A34C" author="Holland F. D. Jr. &amp; Cvancara A. M." pageId="26" pageNumber="69" pagination="495 - 5" refId="ref24095" refString="Holland F. D. Jr. &amp; Cvancara A. M., 1958 - Crabs from the Cannonbal Formation (Paleocene) of North Dakota. Journal of Paleontology, 32 (3): 495 - 5." type="journal article" year="1958">
<collectingCountry id="F37372897B0E6B60FD92FE68861CA36D" box="[515,605,497,523]" name="Netherlands" pageId="26" pageNumber="69">Holland</collectingCountry>
&amp; Cvancara, 1958
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</paragraph>
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<figureCitation id="135F2E9C7B0E6B60FFE0FDB48480A321" box="[113,193,557,583]" captionStart="Fig" captionStartId="26.[113,146,1917,1939]" captionTargetBox="[116,752,1454,1884]" captionTargetId="figure-805@26.[113,756,1451,1887]" captionTargetPageId="26" captionText="Fig. 20 - Ranina burleighensis Holland in Holland &amp; Cvancara, 1958; holotype, USNM 562094 (x 2.2)." figureDoi="http://doi.org/10.5281/zenodo.12753843" httpUri="https://zenodo.org/record/12753843/files/figure.png" pageId="26" pageNumber="69">Fig. 20</figureCitation>
</paragraph>
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<paragraph id="8BDB32197B0E6B60FFE0FDF38601A3C5" blockId="26.[113,756,618,675]" pageId="26" pageNumber="69">
<heading id="D09385757B0E6B60FFE0FDF38601A3C5" centered="true" fontSize="10" level="2" pageId="26" pageNumber="69" reason="3">
<treatmentCitationGroup id="AB7415377B0E6B60FFE0FDF38601A3C5" pageId="26" pageNumber="69">
<taxonomicName id="4C64499A7B0E6B60FFE0FDF38531A3E2" ID-CoL="9ZPLH" authorityName="Holland &amp; Cvancara" authorityYear="1958" box="[113,368,618,644]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="69" phylum="Arthropoda" rank="species" species="burleighensis">
<emphasis id="B910EE0B7B0E6B60FFE0FDF38483A3E5" box="[113,194,618,643]" italics="true" pageId="26" pageNumber="69">Ranina</emphasis>
?
<emphasis id="B910EE0B7B0E6B60FF48FDF38531A3E2" box="[217,368,618,644]" italics="true" pageId="26" pageNumber="69">burleighensis</emphasis>
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<collectingCountry id="F37372897B0E6B60FEEBFDF38595A3E2" box="[378,468,618,644]" name="Netherlands" pageId="26" pageNumber="69">Holland</collectingCountry>
<emphasis id="B910EE0B7B0E6B60FE4EFDF385B4A3E5" box="[479,501,618,643]" italics="true" pageId="26" pageNumber="69">in</emphasis>
<treatmentCitation id="0AC514087B0E6B60FE6EFDF3854AA3C5" author="Holland F. D. Jr. &amp; Cvancara A. M." page="504" pageId="26" pageNumber="69" year="1958">
<bibRefCitation id="EFF54FE87B0E6B60FE6EFDF3854AA3C5" author="Holland F. D. Jr. &amp; Cvancara A. M." pageId="26" pageNumber="69" pagination="495 - 5" refId="ref24095" refString="Holland F. D. Jr. &amp; Cvancara A. M., 1958 - Crabs from the Cannonbal Formation (Paleocene) of North Dakota. Journal of Paleontology, 32 (3): 495 - 5." type="journal article" year="1958">
<collectingCountry id="F37372897B0E6B60FE6EFDF38618A3E2" box="[511,601,618,644]" name="Netherlands" pageId="26" pageNumber="69">Holland</collectingCountry>
&amp; Cvancara, 1958: 504
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, 505, fig. 3c, Pl. 74, fig. 15.
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</heading>
</paragraph>
<paragraph id="8BDB32197B0E6B60FFE0FD5C8488A27C" blockId="26.[113,757,708,794]" pageId="26" pageNumber="69">
<heading id="D09385757B0E6B60FFE0FD5C8488A27C" centered="true" fontSize="10" level="2" pageId="26" pageNumber="69" reason="3">
<treatmentCitationGroup id="AB7415377B0E6B60FFE0FD5C8488A27C" pageId="26" pageNumber="69">
<taxonomicName id="4C64499A7B0E6B60FFE0FD5C8523A3B8" ID-CoL="9ZPLH" authorityName="Holland &amp; Cvancara" authorityYear="1958" box="[113,354,708,734]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="69" phylum="Arthropoda" rank="species" species="burleighensis">
<emphasis id="B910EE0B7B0E6B60FFE0FD5C8523A3B8" box="[113,354,708,734]" italics="true" pageId="26" pageNumber="69">Ranina burleighensis</emphasis>
</taxonomicName>
-
<treatmentCitation id="0AC514087B0E6B60FEEFFD5D86EEA3B8" author="Schweitzer C. &amp; Feldmann R. M. &amp; Garassino A. &amp; Karasawa H. &amp; Schweigert G." box="[382,687,708,734]" page="74" pageId="26" pageNumber="69" year="2010">
<bibRefCitation id="EFF54FE87B0E6B60FEEFFD5D86EEA3B8" author="Schweitzer C. &amp; Feldmann R. M. &amp; Garassino A. &amp; Karasawa H. &amp; Schweigert G." box="[382,687,708,734]" pageId="26" pageNumber="69" pagination="1 - 222" refId="ref24966" refString="Schweitzer C. E, Feldmann R. M., Garassino A., Karasawa H. &amp; Schweigert G., 2010 - Systematic list of fossil decapod crustacean species. Crustaceana Monographs, 10: 1 - 222." type="journal article" year="2010">
Schweitzer
<emphasis id="B910EE0B7B0E6B60FD95FD5C867FA3B8" box="[516,574,708,734]" italics="true" pageId="26" pageNumber="69">et al.</emphasis>
2010: 74
</bibRefCitation>
</treatmentCitation>
. –
<treatmentCitation id="0AC514087B0E6B60FD42FD5C8594A39A" author="De Angeli A. &amp; Beschin C." page="13" pageId="26" pageNumber="69" year="2011">
<bibRefCitation id="EFF54FE87B0E6B60FD42FD5C8594A39A" author="De Angeli A. &amp; Beschin C." pageId="26" pageNumber="69" pagination="11 - 20" refId="ref23611" refString="De Angeli A. &amp; Beschin C., 2011 - Il genere Ranina Lamarck, 1801 (Crustacea, Decapoda, Raninidae) nel Terziario del Vicentino, con descrizione di due nuove specie. Studi e Ricerche - Associazione Amici del Museo - Museo Civico &quot; G. Zannato &quot;, 18: 11 - 20." type="book chapter" year="2011">De Angeli &amp; Beschin 2011: 13</bibRefCitation>
</treatmentCitation>
. –
<treatmentCitation id="0AC514087B0E6B60FE6BFD7A8484A27C" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M. &amp; Luque J." page="260" pageId="26" pageNumber="69" year="2014">
<bibRefCitation id="EFF54FE87B0E6B60FE6BFD7A8484A27C" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M. &amp; Luque J." pageId="26" pageNumber="69" pagination="216 - 272" refId="ref24222" refString="Karasawa H., Schweitzer C. E., Feldmann R. M. &amp; Luque J., 2014 - Phylogeny and classification of Raninoida (Decapoda: Brachyura). Journal of Crustacean Biology, 34 (2): 216 - 272." type="journal article" year="2014">
Karasawa
<emphasis id="B910EE0B7B0E6B60FDE0FD7A86EAA39A" box="[625,683,738,764]" italics="true" pageId="26" pageNumber="69">et al.</emphasis>
2014: 260
</bibRefCitation>
</treatmentCitation>
.
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37E61927B0E6B60FF06FCA485B4A2A9" pageId="26" pageNumber="69" type="materials_examined">
<paragraph id="8BDB32197B0E6B60FF06FCA48631A231" blockId="26.[113,756,828,1397]" box="[151,624,828,855]" pageId="26" pageNumber="69">
<emphasis id="B910EE0B7B0E6B60FF06FCA4857FA230" bold="true" box="[151,318,828,855]" pageId="26" pageNumber="69">Type material</emphasis>
:
<materialsCitation id="3B0C38447B0E6B60FEDCFCA5862CA231" box="[333,621,828,855]" collectionCode="USNM" pageId="26" pageNumber="69" specimenCode="USNM 562094" specimenCount="1" typeStatus="holotype">
<typeStatus id="54DF8CBB7B0E6B60FEDCFCA585F3A230" box="[333,434,828,854]" pageId="26" pageNumber="69" type="holotype">holotype</typeStatus>
,
<specimenCode id="DBC29A627B0E6B60FE2CFCA4862CA231" box="[445,621,828,855]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="69" type="Museum">USNM 562094</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDB32197B0E6B60FF06FCC28519A2F5" blockId="26.[113,756,828,1397]" pageId="26" pageNumber="69">
<materialsCitation id="3B0C38447B0E6B60FF06FCC28514A2F5" country="United States of America" county="Burleigh County" location="South of Moffit" pageId="26" pageNumber="69" specimenCount="1" stateProvince="North Dakota" typeStatus="holotype">
<emphasis id="B910EE0B7B0E6B60FF06FCC28578A212" bold="true" box="[151,313,858,885]" pageId="26" pageNumber="69">
<typeStatus id="54DF8CBB7B0E6B60FF06FCC28493A213" box="[151,210,859,885]" pageId="26" pageNumber="69">Type</typeStatus>
locality
</emphasis>
:
<location id="8EBB64C27B0E6B60FEC0FCC28657A213" LSID="urn:lsid:plazi:treatment:03CD830F7B0E6B60FFE0FE688079A2A9:8EBB64C27B0E6B60FEC0FCC28657A213" box="[337,534,859,885]" country="United States of America" county="Burleigh County" name="South of Moffit" pageId="26" pageNumber="69" stateProvince="North Dakota">South of Moffit</location>
,
<collectingCounty id="62BA4A957B0E6B60FDBBFCC286B1A213" box="[554,752,859,885]" pageId="26" pageNumber="69">Burleigh County</collectingCounty>
,
<collectingRegion id="49A0FCFB7B0E6B60FFE0FCE0854FA2F5" box="[113,270,889,915]" country="United States of America" name="North Dakota" pageId="26" pageNumber="69">North Dakota</collectingRegion>
,
<collectingCountry id="F37372897B0E6B60FE88FCE08514A2F5" box="[281,341,889,915]" name="United States of America" pageId="26" pageNumber="69">USA</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDB32197B0E6B60FF06FC0E8696A2D7" blockId="26.[113,756,828,1397]" box="[151,727,919,945]" pageId="26" pageNumber="69">
<emphasis id="B910EE0B7B0E6B60FF06FC0E8505A2D7" bold="true" box="[151,324,919,945]" pageId="26" pageNumber="69">Geological age</emphasis>
: Cannonball Formation, Paleocene.
</paragraph>
<paragraph id="8BDB32197B0E6B60FF06FC2C85B4A2A9" blockId="26.[113,756,828,1397]" box="[151,501,949,976]" pageId="26" pageNumber="69">
<emphasis id="B910EE0B7B0E6B60FF06FC2C853CA2A9" bold="true" box="[151,381,949,975]" pageId="26" pageNumber="69">Examined material</emphasis>
:
<typeStatus id="54DF8CBB7B0E6B60FE1AFC2C85B0A2A9" box="[395,497,949,975]" pageId="26" pageNumber="69" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E61927B0E6B60FF06FC4A803EA3B8" pageId="26" pageNumber="69" type="description">
<paragraph id="8BDB32197B0E6B60FF06FC4A803EA3B8" blockId="26.[113,756,828,1397]" lastBlockId="26.[832,1475,135,975]" pageId="26" pageNumber="69">
<emphasis id="B910EE0B7B0E6B60FF06FC4A86EAA28B" bold="true" box="[151,683,979,1005]" pageId="26" pageNumber="69">
Description by
<bibRefCitation id="EFF54FE87B0E6B60FEC5FC4A86EAA28B" author="Holland F. D. Jr. &amp; Cvancara A. M." box="[340,683,979,1005]" pageId="26" pageNumber="69" pagination="495 - 5" refId="ref24095" refString="Holland F. D. Jr. &amp; Cvancara A. M., 1958 - Crabs from the Cannonbal Formation (Paleocene) of North Dakota. Journal of Paleontology, 32 (3): 495 - 5." type="journal article" year="1958">
<collectingCountry id="F37372897B0E6B60FEC5FC4A85F4A28B" box="[340,437,979,1005]" name="Netherlands" pageId="26" pageNumber="69">Holland</collectingCountry>
&amp; Cvancara (1958)
</bibRefCitation>
</emphasis>
: “
<emphasis id="B910EE0B7B0E6B60FD5BFC4A8035A3B8" italics="true" pageId="26" pageNumber="69">
The
<typeStatus id="54DF8CBB7B0E6B60FFE0FC688493A56D" box="[113,210,1009,1035]" pageId="26" pageNumber="69" type="holotype">holotype</typeStatus>
consists of the broken distal portion (twothirds?) of the left manus, the stub of the pollex or immovable finger, and the proximal part of the dactylus of a rather large propodus; the ma us, though laterally crushed, appears to have been oblong in cross-section with the width about three-fourths the height; the outer surface of the manus is covered with irregular spaced, asymmetrical tubercles which are inclined forward; between the tubercles are smaller, more symmetrical, rounded granules; inner surface similar but with the tubercles less pronounced; the tubercles apparently disappear toward the upper surface and on the pollex; on upper distal corners of the manus near the articulation of the dactylus is situated a pair of spine bases; the base of a single, small, forward-directed spine can still be seen on the outer surface near the junction with the pollex; the dactylus is so broken that it is impossible to determine its length; on the inferior surface a rounded projection appears near the break, 6.5 mm anterior to the junction with the manus; this projection is presumed to be the first prehensile tooth; at this place the dactylus is subovate but it expands rapidly in a proximal direction and becomes flatter on the superior surface; posteriorly directed projections occur on the upper proximal corners of the dactylus; the projections apparently articulated with poorly preserved sockets in the upper distal corners of the manus; a spine base of an anteriorly directed spine is borne on the distal edge of each projection (the inner spine was observed by the writer but broke during preparation); the dactylus is covered with irregularly spaced shallow pits; although little remains of the pollex, it appears almost rectangular in cross-section, pitted like the dactylus, and inclined downward from the manus.
</emphasis>
”
</paragraph>
</subSubSection>
<subSubSection id="C37E61927B0E6B60FCF4FD7B8079A2A9" pageId="26" pageNumber="69" type="discussion">
<paragraph id="8BDB32197B0E6B60FCF4FD7B8079A2A9" blockId="26.[832,1475,135,975]" pageId="26" pageNumber="69">
<emphasis id="B910EE0B7B0E6B60FCF4FD7B87A2A39A" bold="true" box="[869,995,738,764]" pageId="26" pageNumber="69">Discussion</emphasis>
. Based upon the above-reported original description, this manus has strong propodus and dactylus that do not fit clearly the typical characters of a raninid manus. Indeed according to Feldmann (pers. comm., 2015) the manus clearly resembles that of a lobster or a xanthid. In conclusion, this species cannot be assigned to
<taxonomicName id="4C64499A7B0E6B60FCD1FC0E87D0A2D6" box="[832,913,919,944]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B910EE0B7B0E6B60FCD1FC0E87D0A2D6" box="[832,913,919,944]" italics="true" pageId="26" pageNumber="69">Ranina</emphasis>
</taxonomicName>
and this manus might well belong to a variety of other decapod groups.
</paragraph>
</subSubSection>
</treatment>
</document>
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