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<document id="64781789B7F07D0B7C164D5B2EF4C1F5" ID-DOI="10.11646/zootaxa.3700.3.9" ID-GBIF-Dataset="378fa437-543f-42a9-ada1-1778983a5eba" ID-ISSN="1175-5326" ID-Zenodo-Dep="219974" ID-ZooBank="8ABAD147-4B64-4F0C-80D1-0D977B93D7D3" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460664016035" checkinUser="plazi" docAuthor="Bekchiev, Rostislav &amp; Hlaváč, Peter" docDate="2013" docId="03B887FE06508E1BBC910B43322760C2" docLanguage="en" docName="zt03700p498.pdf" docOrigin="Zootaxa 3700 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pselaphodes unicornis Bekchiev &amp; Hlaváč, 2013, sp.n." docType="treatment" docUuid="89D368F5-3096-4015-85D7-3B593246468F" docUuidSource="ZooBank" docVersion="8" lastPageNumber="498" masterDocId="FF81FF8606528E18BC060D6031206345" masterDocTitle="Description of a new unusual Pselaphodes Westwood, 1870 (Coleoptera: Staphylinidae: Pselaphinae) from Nepal" masterLastPageNumber="498" masterPageNumber="495" pageNumber="497" updateTime="1698347700654" updateUser="plazi">
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<mods:title id="6F8D79563B47BCA06963A2DE7A319E53">Description of a new unusual Pselaphodes Westwood, 1870 (Coleoptera: Staphylinidae: Pselaphinae) from Nepal</mods:title>
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<mods:name id="C608EF0689BE9EA80625F41247F8B053" type="personal">
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<mods:namePart id="27E19B91826BDE586DA9A74AD38C6876">Bekchiev, Rostislav</mods:namePart>
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<mods:namePart id="8FC7B580721D7DF1147958AB68476355">Hlaváč, Peter</mods:namePart>
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<mods:typeOfResource id="44C87E7D3FEAC92B0C22C5B3F2F43A49">text</mods:typeOfResource>
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<mods:title id="06086B095D19DD1227C4D5D4CEA4618A">Zootaxa</mods:title>
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<mods:date id="9835665B1AA3DD941B4EC8BE0BE842E4">2013</mods:date>
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<mods:number id="083C4BD8DC2C22B5768EEC37D5D4976F">3700</mods:number>
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<mods:identifier id="06D63EFA8F95BEC683EF86CCD89A6469" type="DOI">10.11646/zootaxa.3700.3.9</mods:identifier>
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<treatment id="03B887FE06508E1BBC910B43322760C2" ID-DOI="http://doi.org/10.5281/zenodo.6154541" ID-GBIF-Taxon="119574894" ID-Zenodo-Dep="6154541" LSID="urn:lsid:zoobank.org:act:89D368F5-3096-4015-85D7-3B593246468F" httpUri="http://treatment.plazi.org/id/03B887FE06508E1BBC910B43322760C2" lastPageId="3" lastPageNumber="498" pageId="2" pageNumber="497">
<subSubSection id="C30B656306508E1ABC910B43302C6519" pageId="2" pageNumber="497" type="nomenclature">
<paragraph id="8BAE36E806508E1ABC910B4330CA6578" blockId="2.[151,490,1571,1628]" box="[151,490,1571,1597]" pageId="2" pageNumber="497">
<heading id="D0E6818406508E1ABC910B4330CA6578" bold="true" box="[151,490,1571,1597]" fontSize="11" level="1" pageId="2" pageNumber="497" reason="1">
<emphasis id="B965EAFA06508E1ABC910B4330CA6578" bold="true" box="[151,490,1571,1597]" pageId="2" pageNumber="497">
<taxonomicName id="4C114D6B06508E1ABC910B4330886578" ID-CoL="6WD4Q" box="[151,424,1571,1597]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="497" phylum="Arthropoda" rank="species" species="unicornis" status="sp. nov.">
<emphasis id="B965EAFA06508E1ABC910B4330886578" bold="true" box="[151,424,1571,1597]" italics="true" pageId="2" pageNumber="497">Pselaphodes unicornis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A256578106508E1ABDA90B4330CA6578" box="[431,490,1571,1597]" pageId="2" pageNumber="497" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36E806508E1ABC910B25302C6519" blockId="2.[151,490,1571,1628]" box="[151,268,1605,1628]" pageId="2" pageNumber="497">
(
<figureCitation id="132A2A6D06508E1ABC990B2531CA6519" box="[159,234,1605,1628]" captionStart="FIGURE 2" captionStartId="1.[151,244,1643,1665]" captionTargetBox="[151,1428,200,1609]" captionTargetId="figure@1.[151,1436,194,1622]" captionTargetPageId="1" captionText="FIGURE 2. Diagnostic features of Pselaphodes unicornis in male: a) habitus, dorsal view scale 0.2 mm; b) head, lateral view scale 0.6 mm; c) metaventral processes, lateral view scale 0.4 mm; d) protrochanter and profemur scale 1.0 mm; e) mesotrochanter and mesofemur scale 1.0 mm." httpUri="https://zenodo.org/record/219976/files/figure.png" pageId="2" pageNumber="497">Figs. 2</figureCitation>
,
<figureCitation id="132A2A6D06508E1ABCF30B2530226519" box="[245,258,1605,1628]" captionStart="FIGURE 3" captionStartId="2.[151,244,1471,1493]" captionTargetBox="[163,1401,446,1427]" captionTargetId="figure@2.[151,1436,437,1451]" captionTargetPageId="2" captionText="FIGURE 3. Aedeagus of Pselaphodes unicornis: a) dorsal view; b) lateral view, scale 0.2 mm." httpUri="https://zenodo.org/record/219977/files/figure.png" pageId="2" pageNumber="497">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30B656306508E1ABC910BEB3354644F" pageId="2" pageNumber="497" type="materials_examined">
<paragraph id="8BAE36E806508E1ABC910BEB3354644F" blockId="2.[151,1436,1675,2010]" pageId="2" pageNumber="497">
<emphasis id="B965EAFA06508E1ABC910BEB305D65E7" bold="true" box="[151,381,1675,1698]" pageId="2" pageNumber="497">Material examined:</emphasis>
(
<specimenCount id="9D17FD6106508E1ABD940BEB30CD65E7" box="[402,493,1675,1698]" pageId="2" pageNumber="497" type="male">4 males</specimenCount>
,
<specimenCount id="9D17FD6106508E1ABDF80BEB334365E7" box="[510,611,1675,1698]" pageId="2" pageNumber="497" type="female">1 female</specimenCount>
):
<typeStatus id="54AA884A06508E1ABE790BEB322E65E7" box="[639,782,1675,1698]" pageId="2" pageNumber="497" type="holotype">HOLOTYPE</typeStatus>
, male, labelled as follows: (p)
<collectingCountry id="F306767806508E1AB87C0BEB359C65E7" box="[1146,1212,1675,1698]" name="Nepal" pageId="2" pageNumber="497">Nepal</collectingCountry>
, Parbat, Ghorepani, Rhodendron forest,
<quantity id="4CE99B0D06508E1ABD6D0BCD309D6581" box="[363,445,1709,1732]" metricMagnitude="3" metricUnit="m" metricValue="2.86" pageId="2" pageNumber="497" unit="m" value="2860.0">2860 m</quantity>
,
<date id="FFAF102806508E1ABDCE0BCD33626581" box="[456,578,1709,1732]" pageId="2" pageNumber="497" value="2006-10-13">13.10.2006</date>
, leg. D. Bechev / red label (p)
<typeStatus id="54AA884A06508E1ABF800BCD35336581" box="[902,1043,1709,1732]" pageId="2" pageNumber="497" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C114D6B06508E1AB81F0BCE35F76580" box="[1049,1239,1710,1733]" class="Insecta" family="Staphylinidae" genus="Lasinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="497" phylum="Arthropoda" rank="species" species="unicornis" status="sp. nov.">
<emphasis id="B965EAFA06508E1AB81F0BCE35F76580" box="[1049,1239,1710,1733]" italics="true" pageId="2" pageNumber="497">Lasinus unicornis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A256578106508E1AB8DB0BCD34316581" box="[1245,1297,1709,1732]" pageId="2" pageNumber="497" rank="species">sp.n.</taxonomicNameLabel>
, Bekchiev &amp; Hlaváč det. 2013. (NMNHS).
<typeStatus id="54AA884A06508E1ABDEC0BB033A265A2" box="[490,642,1744,1767]" pageId="2" pageNumber="497" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D17FD6106508E1ABE8B0BB033C465A2" box="[653,740,1744,1767]" pageId="2" pageNumber="497" type="male">3 males</specimenCount>
,
<specimenCount id="9D17FD6106508E1ABEF70BB0327165A2" box="[753,849,1744,1767]" pageId="2" pageNumber="497" type="female">1 female</specimenCount>
:
<collectingCountry id="F306767806508E1ABF670BB0328365A2" box="[865,931,1744,1767]" name="Nepal" pageId="2" pageNumber="497">Nepal</collectingCountry>
, Langtang Valley. Lama longe,
<quantity id="4CE99B0D06508E1AB9060BB034B765A2" box="[1280,1431,1744,1767]" metricMagnitude="3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="2" pageNumber="497" unit="m" value="2650.0" valueMax="2800.0" valueMin="2500.0">25002800 m</quantity>
,
<date id="FFAF102806508E1ABC910B933032644F" box="[151,274,1779,1802]" pageId="2" pageNumber="497" value="1984-10-25">25.10.1984</date>
, leg. P. Beron (NMNHS, PCPH).
</paragraph>
</subSubSection>
<subSubSection id="C30B656306508E1BBCC10A7530206134" lastPageId="3" lastPageNumber="498" pageId="2" pageNumber="497" type="description">
<paragraph id="8BAE36E806508E1ABCC10A7530AE640A" blockId="2.[151,1436,1675,2010]" pageId="2" pageNumber="497">
<emphasis id="B965EAFA06508E1ABCC10A7530746469" bold="true" box="[199,340,1813,1836]" pageId="2" pageNumber="497">Description:</emphasis>
Body length (BL) 2.703.00 mm (
<figureCitation id="132A2A6D06508E1ABEE40A7532076469" box="[738,807,1813,1836]" captionStart="FIGURE 2" captionStartId="1.[151,244,1643,1665]" captionTargetBox="[151,1428,200,1609]" captionTargetId="figure@1.[151,1436,194,1622]" captionTargetPageId="1" captionText="FIGURE 2. Diagnostic features of Pselaphodes unicornis in male: a) habitus, dorsal view scale 0.2 mm; b) head, lateral view scale 0.6 mm; c) metaventral processes, lateral view scale 0.4 mm; d) protrochanter and profemur scale 1.0 mm; e) mesotrochanter and mesofemur scale 1.0 mm." httpUri="https://zenodo.org/record/219976/files/figure.png" pageId="2" pageNumber="497">Fig. 2</figureCitation>
a), unicoloured, reddish-brown, abdomen slightly darker, maxillary palpi yellow.
</paragraph>
<paragraph id="8BAE36E806508E1ABCC10A3B33F6649F" blockId="2.[151,1436,1675,2010]" pageId="2" pageNumber="497">
Head elongate with very prominent eyes, longer than wide (HL: 0.60062 mm; HW:
<quantity id="4CE99B0D06508E1AB8620A3B34246437" box="[1124,1284,1883,1906]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="2" pageNumber="497" unit="mm" value="0.49" valueMax="0.5" valueMin="0.48">0.480.50 mm</quantity>
), vertexal and frontal foveae well-defined, median sulcus deep only in frontal part, vanishing on vertex, clypeus with strongly downward curved horn (
<figureCitation id="132A2A6D06508E1ABD980AC030FF64F2" box="[414,479,1952,1975]" captionStart="FIGURE 2" captionStartId="1.[151,244,1643,1665]" captionTargetBox="[151,1428,200,1609]" captionTargetId="figure@1.[151,1436,194,1622]" captionTargetPageId="1" captionText="FIGURE 2. Diagnostic features of Pselaphodes unicornis in male: a) habitus, dorsal view scale 0.2 mm; b) head, lateral view scale 0.6 mm; c) metaventral processes, lateral view scale 0.4 mm; d) protrochanter and profemur scale 1.0 mm; e) mesotrochanter and mesofemur scale 1.0 mm." httpUri="https://zenodo.org/record/219976/files/figure.png" pageId="2" pageNumber="497">Fig. 2</figureCitation>
b), maxillary palpi with palpomere III indistinctly projected laterally. Antennae relatively short,
<quantity id="4CE99B0D06508E1ABCDF0AA33055649F" box="[217,373,1987,2010]" metricMagnitude="-3" metricUnit="m" metricValue="1.495" metricValueMax="1.51" metricValueMin="1.48" pageId="2" pageNumber="497" unit="mm" value="1.495" valueMax="1.51" valueMin="1.48">1.481.51 mm</quantity>
long, antennal club as in
<figureCitation id="132A2A6D06508E1ABE830AA333E5649F" box="[645,709,1987,2010]" captionStart="FIGURE 2" captionStartId="1.[151,244,1643,1665]" captionTargetBox="[151,1428,200,1609]" captionTargetId="figure@1.[151,1436,194,1622]" captionTargetPageId="1" captionText="FIGURE 2. Diagnostic features of Pselaphodes unicornis in male: a) habitus, dorsal view scale 0.2 mm; b) head, lateral view scale 0.6 mm; c) metaventral processes, lateral view scale 0.4 mm; d) protrochanter and profemur scale 1.0 mm; e) mesotrochanter and mesofemur scale 1.0 mm." httpUri="https://zenodo.org/record/219976/files/figure.png" pageId="2" pageNumber="497">Fig. 2</figureCitation>
a.
</paragraph>
<paragraph id="8BAE36E806518E1BBCC10DF835B66252" blockId="3.[151,1437,152,695]" pageId="3" pageNumber="498">
Pronotum slightly longer than wide (PL:
<quantity id="4CE99B0D06518E1BBE8B0DF8320E63EA" box="[653,814,152,175]" metricMagnitude="-4" metricUnit="m" metricValue="6.05" metricValueMax="6.2" metricValueMin="5.8999999999999995" pageId="3" pageNumber="498" unit="mm" value="0.605" valueMax="0.62" valueMin="0.59">0.590.62 mm</quantity>
; PW:
<quantity id="4CE99B0D06518E1BBF740DF8353463EA" box="[882,1044,152,175]" metricMagnitude="-4" metricUnit="m" metricValue="4.95" metricValueMax="5.1" metricValueMin="4.8" pageId="3" pageNumber="498" unit="mm" value="0.495" valueMax="0.51" valueMin="0.48">0.480.51 mm</quantity>
), disc smooth, lateral parts wrinkly, lateral and median foveae glabrous, well-defined, median sulcus almost reaching anterior margin of pronotum. Metaventrite with two long (
<quantity id="4CE99B0D06518E1BBDD40DBD331963B1" box="[466,569,221,244]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="498" unit="mm" value="0.14">0.14 mm</quantity>
), slightly curved, apically bifid basal horny processes and two small triangular processes behind them (
<figureCitation id="132A2A6D06518E1BBD910C6030F86252" box="[407,472,256,279]" captionStart="FIGURE 2" captionStartId="1.[151,244,1643,1665]" captionTargetBox="[151,1428,200,1609]" captionTargetId="figure@1.[151,1436,194,1622]" captionTargetPageId="1" captionText="FIGURE 2. Diagnostic features of Pselaphodes unicornis in male: a) habitus, dorsal view scale 0.2 mm; b) head, lateral view scale 0.6 mm; c) metaventral processes, lateral view scale 0.4 mm; d) protrochanter and profemur scale 1.0 mm; e) mesotrochanter and mesofemur scale 1.0 mm." httpUri="https://zenodo.org/record/219976/files/figure.png" pageId="3" pageNumber="498">Fig. 2</figureCitation>
c). Elytra wider than long, EL
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, EW
<quantity id="4CE99B0D06518E1BBFF20C6035B26252" box="[1012,1170,256,279]" metricMagnitude="-3" metricUnit="m" metricValue="1.07" metricValueMax="1.08" metricValueMin="1.06" pageId="3" pageNumber="498" unit="mm" value="1.07" valueMax="1.08" valueMin="1.06">1.061.08 mm</quantity>
.
</paragraph>
<paragraph id="8BAE36E806518E1BBCC10C4235B36219" blockId="3.[151,1437,152,695]" pageId="3" pageNumber="498">
Legs long and slender, protrochanters with small ventral spine, profemora with long and robust ventral spine at middle (
<figureCitation id="132A2A6D06518E1BBCE90C25300F6219" box="[239,303,325,348]" captionStart="FIGURE 2" captionStartId="1.[151,244,1643,1665]" captionTargetBox="[151,1428,200,1609]" captionTargetId="figure@1.[151,1436,194,1622]" captionTargetPageId="1" captionText="FIGURE 2. Diagnostic features of Pselaphodes unicornis in male: a) habitus, dorsal view scale 0.2 mm; b) head, lateral view scale 0.6 mm; c) metaventral processes, lateral view scale 0.4 mm; d) protrochanter and profemur scale 1.0 mm; e) mesotrochanter and mesofemur scale 1.0 mm." httpUri="https://zenodo.org/record/219976/files/figure.png" pageId="3" pageNumber="498">Fig. 2</figureCitation>
d), mesotrochanter with two spines at apex, mesofemora without spines (
<figureCitation id="132A2A6D06518E1BB83E0C2535596219" box="[1080,1145,325,348]" captionStart="FIGURE 2" captionStartId="1.[151,244,1643,1665]" captionTargetBox="[151,1428,200,1609]" captionTargetId="figure@1.[151,1436,194,1622]" captionTargetPageId="1" captionText="FIGURE 2. Diagnostic features of Pselaphodes unicornis in male: a) habitus, dorsal view scale 0.2 mm; b) head, lateral view scale 0.6 mm; c) metaventral processes, lateral view scale 0.4 mm; d) protrochanter and profemur scale 1.0 mm; e) mesotrochanter and mesofemur scale 1.0 mm." httpUri="https://zenodo.org/record/219976/files/figure.png" pageId="3" pageNumber="498">Fig. 2</figureCitation>
e).
</paragraph>
<paragraph id="8BAE36E806518E1BBCC10C0835BF62E4" blockId="3.[151,1437,152,695]" pageId="3" pageNumber="498">
Abdomen broad at base and narrowed apically, AL
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, AW 1.00
<quantity id="4CE99B0D06518E1BB82A0C0835B1623A" box="[1068,1169,360,383]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="498" unit="mm" value="1.1">1.10 mm</quantity>
. First visible abdominal tergite (IV) glabrous, basal carinae absent. Aedeagus:
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long, as in
<figureCitation id="132A2A6D06518E1BBFE90CEA351662E4" box="[1007,1078,394,417]" captionStart="FIGURE 3" captionStartId="2.[151,244,1471,1493]" captionTargetBox="[163,1401,446,1427]" captionTargetId="figure@2.[151,1436,437,1451]" captionTargetPageId="2" captionText="FIGURE 3. Aedeagus of Pselaphodes unicornis: a) dorsal view; b) lateral view, scale 0.2 mm." httpUri="https://zenodo.org/record/219977/files/figure.png" pageId="3" pageNumber="498">Figs. 3</figureCitation>
a and, 3b.
</paragraph>
<paragraph id="8BAE36E806518E1BBCC10CCD326662A2" blockId="3.[151,1437,152,695]" pageId="3" pageNumber="498">
<emphasis id="B965EAFA06518E1BBCC10CCD30906281" bold="true" box="[199,432,429,452]" pageId="3" pageNumber="498">Sexual dimorphism:</emphasis>
Female readily differs from male by simple unmodified antennal club, the clypeus lacking horn and metaventrite without prominent metaventral processes.
</paragraph>
<paragraph id="8BAE36E806518E1BBCC10C9230206134" blockId="3.[151,1437,152,695]" pageId="3" pageNumber="498">
<emphasis id="B965EAFA06518E1BBCC10C9230E4614C" bold="true" box="[199,452,498,521]" pageId="3" pageNumber="498">Differential diagnosis:</emphasis>
<taxonomicName id="4C114D6B06518E1BBDCA0C92336D614C" box="[460,589,498,521]" class="Insecta" family="Staphylinidae" genus="Lasinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="B965EAFA06518E1BBDCA0C92336D614C" box="[460,589,498,521]" italics="true" pageId="3" pageNumber="498">P. unicornis</emphasis>
</taxonomicName>
is readily separated from all other species of the genus by the maxillary palpi with segment III indistinctly projected laterally and by the presence of a down-curved horn on the clypeus. It is closely related to
<taxonomicName id="4C114D6B06518E1BBD0E0F5830B9610A" box="[264,409,568,591]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="monoceros">
<emphasis id="B965EAFA06518E1BBD0E0F5830B9610A" box="[264,409,568,591]" italics="true" pageId="3" pageNumber="498">P. monoceros</emphasis>
</taxonomicName>
but differs from it mainly by the shape of the metaventral process, the clypeal horn and the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C30B656306518E1BBCC10F1D308961D1" box="[199,425,637,660]" pageId="3" pageNumber="498" type="distribution">
<paragraph id="8BAE36E806518E1BBCC10F1D308961D1" blockId="3.[151,1437,152,695]" box="[199,425,637,660]" pageId="3" pageNumber="498">
<emphasis id="B965EAFA06518E1BBCC10F1D307B61D1" bold="true" box="[199,347,637,660]" pageId="3" pageNumber="498">Distribution:</emphasis>
<collectingCountry id="F306767806518E1BBD640F1D308561D1" box="[354,421,637,660]" name="Nepal" pageId="3" pageNumber="498">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B656306518E1BBCC10FC0322760C2" pageId="3" pageNumber="498" type="etymology">
<paragraph id="8BAE36E806518E1BBCC10FC0328961F2" blockId="3.[151,1437,152,695]" box="[199,937,672,695]" pageId="3" pageNumber="498">
<emphasis id="B965EAFA06518E1BBCC10FC0306A61F2" bold="true" box="[199,330,672,695]" pageId="3" pageNumber="498">Etymology:</emphasis>
The name is referring to the horned clypeus of the male.
</paragraph>
<paragraph id="8BAE36E806518E1BBC910E683053605A" blockId="3.[151,371,776,799]" box="[151,371,776,799]" pageId="3" pageNumber="498">
<emphasis id="B965EAFA06518E1BBC910E683053605A" bold="true" box="[151,371,776,799]" pageId="3" pageNumber="498">Аcknowledgements</emphasis>
</paragraph>
<paragraph id="8BAE36E806518E1BBC910E2D322760C2" blockId="3.[151,1436,845,903]" pageId="3" pageNumber="498">
We want to thank our colleagues and reviewers George Coulon (
<collectingCountry id="F306767806518E1BBF630E2D329A6021" box="[869,954,845,868]" name="Canada" pageId="3" pageNumber="498">Canada</collectingCountry>
), and Zi-Wei Yin (
<collectingCountry id="F306767806518E1BB8900E2D35F96021" box="[1174,1241,845,868]" name="China" pageId="3" pageNumber="498">China</collectingCountry>
) for the critically reading of the manuscript and providing useful comments.
</paragraph>
</subSubSection>
</treatment>
</document>