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<document id="DFBA06B5B0A25266357C61178247AC98" ID-CLB-Dataset="102843" ID-DOI="10.11646/zootaxa.5168.2.5" ID-GBIF-Dataset="16df2448-1e2c-4410-ac7b-af3623ff401f" ID-ISSN="1175-5326" ID-Zenodo-Dep="6877267" ID-ZooBank="36831FDB-E9BE-4255-82B9-BFFCDCD40098" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658420700313" checkinUser="plazi" docAuthor="Zhao, Ming-Zhi &amp; Zorn, Carsten" docDate="2022" docId="03B887EBDF62FFAA17AC05320BE3F83B" docLanguage="en" docName="zootaxa.5168.2.5.pdf" docOrigin="Zootaxa 5168 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anomala longilobata Zhao &amp; Zorn 2022, new species" docType="treatment" docUuid="5F2B62DB-827C-4E3F-962F-A4ABB5183036" docUuidSource="ZooBank" docVersion="6" lastPageNumber="194" masterDocId="FF81FF93DF72FFB9173B045C0836FF81" masterDocTitle="Contribution to the genus Anomala Samouelle, 1819 (Coleoptera: Scarabaeidae Rutelinae) of China and adjacent regions. Part II: six new species from Taiwan and Hainan" masterLastPageNumber="195" masterPageNumber="175" pageNumber="191" updateTime="1699361553354" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="465C9EE563612D3138B8DACCF343367C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B071B1F9DA95E553D52A289E6EAD67E6">
<mods:title id="47F4C40B1393C2F71BEFB861F7AF3E76">Contribution to the genus Anomala Samouelle, 1819 (Coleoptera: Scarabaeidae Rutelinae) of China and adjacent regions. Part II: six new species from Taiwan and Hainan</mods:title>
</mods:titleInfo>
<mods:name id="C8564EF3202D7EDF6FC4B2F0E1FFB084" type="personal">
<mods:role id="3DD45D1D1925609C24B17A12432EE467">
<mods:roleTerm id="F5020AA82247EC6CCF0CA05D90D3F81B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D15C897389B65FEDE2AEE7413C5120F9">Zhao, Ming-Zhi</mods:namePart>
<mods:nameIdentifier id="97CC7DE4EC463808BD825CBAC1235A9E" type="ORCID">0000-0002-3158-6177</mods:nameIdentifier>
<mods:affiliation id="9873AB9AF0EE1DC2BACBA76E86EDF347">College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, P. R. China. zhaomzhai @ 126. com; https: // orcid. org / 0000 - 0002 - 3158 - 6177</mods:affiliation>
</mods:name>
<mods:name id="893BF2499DA8E2E576C7ED172CB30322" type="personal">
<mods:role id="50E38C718054AA93FC9184944EF14372">
<mods:roleTerm id="F89B5ABCE663F54D325ED2029E8AC977">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F861C820266F0AFF7A82409C2AB7E077">Zorn, Carsten</mods:namePart>
<mods:affiliation id="98B17C258F01290429D64B69146B9D6F">Rostocker Strasse 1 a, Gnoien, 17179, Germany.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="3238A1E0E623ED4E265CEAE07DE2CE2B">text</mods:typeOfResource>
<mods:relatedItem id="AA9C03828586B31CBB5A8B076D4B94DB" type="host">
<mods:titleInfo id="AB959B970757F8DF7231E2C685B95871">
<mods:title id="B7D1864A2812724689A2A265EDD6B13B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6F7A6E4BF9DD1C8E776AF46F7807A0A9">
<mods:date id="0C42AA2FC64E868BBA42397723EEB221">2022</mods:date>
<mods:detail id="5B554066039E810C24B53C785D849A4C" type="pubDate">
<mods:number id="41A2F125929D9785B36B5AC5F237894C">2022-07-21</mods:number>
</mods:detail>
<mods:detail id="253A36CE130C0430F3C8BFCE90BEABBC" type="volume">
<mods:number id="6CD7E6C97C0B4E03FA02E229CC9E4571">5168</mods:number>
</mods:detail>
<mods:detail id="A4BEBDD04ED66234E814E49D0A84684F" type="issue">
<mods:number id="BE85AC18AA02C62511265B296EEE88BE">2</mods:number>
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<mods:start id="A7C51E7892CCFEEB7C115BD6965DE00F">175</mods:start>
<mods:end id="1A407848928118F63D1B367ABD6C0578">195</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D3FB1719E5C88C3A4361A951872A8FD2">journal article</mods:classification>
<mods:identifier id="2FC18B561C02DCC619FBE957E08DBC80" type="CLB-Dataset">102843</mods:identifier>
<mods:identifier id="88D47FA6C27510509ECBB2D36735ABA7" type="DOI">10.11646/zootaxa.5168.2.5</mods:identifier>
<mods:identifier id="2E22D6D2316235C99FFEB2A665AFC006" type="GBIF-Dataset">16df2448-1e2c-4410-ac7b-af3623ff401f</mods:identifier>
<mods:identifier id="3D65394CF53341B4A4233057B69D8DB1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DBF981F7B4610964ACA6882DA6655A90" type="Zenodo-Dep">6877267</mods:identifier>
<mods:identifier id="796D131532D082911CDAF1CC6F680295" type="ZooBank">36831FDB-E9BE-4255-82B9-BFFCDCD40098</mods:identifier>
</mods:mods>
<treatment id="03B887EBDF62FFAA17AC05320BE3F83B" ID-DOI="http://doi.org/10.5281/zenodo.6877223" ID-GBIF-Taxon="197841324" ID-Zenodo-Dep="6877223" LSID="urn:lsid:zoobank.org:act:5F2B62DB-827C-4E3F-962F-A4ABB5183036" httpUri="http://treatment.plazi.org/id/03B887EBDF62FFAA17AC05320BE3F83B" lastPageId="19" lastPageNumber="194" pageId="16" pageNumber="191">
<subSubSection id="C30B6576DF62FFA917AC05320B99FE08" box="[151,943,366,393]" pageId="16" pageNumber="191" type="nomenclature">
<paragraph id="8BAE36FDDF62FFA917AC05320B99FE08" blockId="16.[151,943,366,429]" box="[151,943,366,393]" pageId="16" pageNumber="191">
<heading id="D0E68191DF62FFA917AC05320B99FE08" bold="true" box="[151,943,366,393]" fontSize="11" level="1" pageId="16" pageNumber="191" reason="1">
<emphasis id="B965EAEFDF62FFA917AC05320B99FE08" bold="true" box="[151,943,366,393]" pageId="16" pageNumber="191">
<taxonomicName id="4C114D7EDF62FFA917AC05320A71FE08" authority="Zhao &amp; Zorn" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[151,583,366,393]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="191" phylum="Arthropoda" rank="species" species="longilobata" status="sp. nov.">
<emphasis id="B965EAEFDF62FFA917AC053209A0FE08" bold="true" box="[151,406,366,393]" italics="true" pageId="16" pageNumber="191">Anomala longilobata</emphasis>
Zhao &amp; Zorn
</taxonomicName>
,
<taxonomicNameLabel id="A2565794DF62FFA9156F05330AD2FE09" box="[596,740,366,393]" pageId="16" pageNumber="191" rank="species">new species</taxonomicNameLabel>
[ḴNJ异fflŵoi]
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30B6576DF62FFA917AC05CF09C4FE2C" box="[151,498,403,429]" pageId="16" pageNumber="191" type="description">
<paragraph id="8BAE36FDDF62FFA917AC05CF09C4FE2C" blockId="16.[151,943,366,429]" box="[151,498,403,429]" pageId="16" pageNumber="191">
(
<figureCitation id="132A2A78DF62FFA917A405CF08CBFE2C" box="[159,253,403,429]" captionStart="FIGURE 10" captionStartId="17.[151,250,1856,1880]" captionTargetBox="[162,1430,290,1814]" captionTargetId="figure-56@17.[151,1435,273,1833]" captionTargetPageId="17" captionText="FIGURE 10. Habitus of Anomala longilobata Zhao &amp; Zorn, new species. A–C, holotype; D–F, female paratype. A, D, dorsal view; B, E, lateral view; C, F, ventral view. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877291" httpUri="https://zenodo.org/record/6877291/files/figure.png" pageId="16" pageNumber="191">Figs. 10</figureCitation>
,
<figureCitation id="132A2A78DF62FFA9163205CF09BAFE2C" box="[265,396,403,429]" captionStart="FIGURE 11" captionStartId="18.[151,250,1839,1863]" captionTargetBox="[164,1424,193,1807]" captionTargetId="figure-21@18.[151,1435,181,1816]" captionTargetPageId="18" captionText="FIGURE 11. Habitus of Anomala spp., in dorsal view.A–E, I, paratypes of A. longilobata Zhao &amp; Zorn, new species; F–H, J–L, A. imperialis Arrow, 1899 (China: Guangxi). A–H, male; I–L, female. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877295" httpUri="https://zenodo.org/record/6877295/files/figure.png" pageId="16" pageNumber="191">11A–E, 11I</figureCitation>
,
<figureCitation id="132A2A78DF62FFA916AC05CF09DCFE2C" box="[407,490,403,429]" captionStart="FIGURE 12" captionStartId="19.[151,250,730,754]" captionTargetBox="[162,1428,184,697]" captionTargetId="figure-18@19.[151,1435,181,705]" captionTargetPageId="19" captionText="FIGURE 12. Male genitalia of Anomala spp. A–C, holotype of A. longilobata Zhao &amp; Zorn, new species; D–F, A. imperialis Arrow, 1899 (China: Guangxi). A, D, dorsal view; B, E, right lateral view; C, F, ventral view. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6877297" httpUri="https://zenodo.org/record/6877297/files/figure.png" pageId="16" pageNumber="191">12A–C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30B6576DF62FFA917AC05870A6AFC94" pageId="16" pageNumber="191" type="materials_examined">
<paragraph id="8BAE36FDDF62FFA917AC05870A6AFC94" blockId="16.[151,1437,474,2013]" pageId="16" pageNumber="191">
<emphasis id="B965EAEFDF62FFA917AC05870975FE75" bold="true" box="[151,323,474,501]" pageId="16" pageNumber="191">Type material.</emphasis>
<materialsCitation id="3B793CA0DF62FFA9167305870AA5FD98" ID-GBIF-Occurrence="3864839311" collectionCode="SCAU" collectorName="Shi-Liang Mo" county="Island" elevation="1412" latitude="18.710709" location="Mts. Jianfengling" longLatPrecision="1" longitude="108.87581" municipality="Ledong" pageId="16" pageNumber="191" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="holotype">
<emphasis id="B965EAEFDF62FFA9167305870988FE74" bold="true" box="[328,446,475,501]" pageId="16" pageNumber="191">
<typeStatus id="54AA885FDF62FFA916730587098FFE74" box="[328,441,475,501]" pageId="16" pageNumber="191" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D17FD74DF62FFA916F8058709EEFE74" box="[451,472,475,501]" pageId="16" pageNumber="191" type="male">♂</specimenCount>
(
<collectionCode id="ED00AE38DF62FFA916DE05870A03FE74" box="[485,565,475,501]" pageId="16" pageNumber="191">SCAU</collectionCode>
),
<collectingRegion id="49D5F81FDF62FFA9157C05870AAEFE74" box="[583,664,475,501]" country="China" name="Hainan" pageId="16" pageNumber="191">Hainan</collectingRegion>
<collectingCounty id="62CF4E71DF62FFA915A605870AD2FE74" box="[669,740,475,501]" pageId="16" pageNumber="191">Island</collectingCounty>
,
<collectingMunicipality id="6BCAAC87DF62FFA915D505870B70FE74" box="[750,838,475,501]" pageId="16" pageNumber="191">Ledong</collectingMunicipality>
,
<location id="8ECE6026DF62FFA9147405870C27FE74" LSID="urn:lsid:plazi:treatment:03B887EBDF62FFAA17AC05320BE3F83B:8ECE6026DF62FFA9147405870C27FE74" box="[847,1041,475,501]" county="Island" latitude="18.710709" longLatPrecision="1" longitude="108.87581" municipality="Ledong" name="Mts. Jianfengling" pageId="16" pageNumber="191" stateProvince="Hainan">Mts. Jianfengling</location>
[=
<emphasis id="B965EAEFDF62FFA9130E05870CBFFE73" bold="true" box="[1077,1161,475,498]" pageId="16" pageNumber="191">尖峰岭</emphasis>
], alt.
<quantity id="4CE99B18DF62FFA913FD05870D2DFE74" box="[1222,1307,475,501]" metricMagnitude="3" metricUnit="m" metricValue="1.412" pageId="16" pageNumber="191" unit="m" value="1412.0">
<elevation id="003CD1CEDF62FFA913FD05870D2DFE74" box="[1222,1307,475,501]" metricMagnitude="3" metricUnit="m" metricValue="1.412" pageId="16" pageNumber="191" unit="m" value="1412.0">1412 m</elevation>
</quantity>
,
<geoCoordinate id="EE25503ADF62FFA9121E05870898FD98" degrees="18.710708" direction="north" orientation="latitude" pageId="16" pageNumber="191" precision="1" value="18.710709">18.710708 N</geoCoordinate>
,
<geoCoordinate id="EE25503ADF62FFA9178305A20960FD98" box="[184,342,510,537]" degrees="108.875812" direction="east" orientation="longitude" pageId="16" pageNumber="191" precision="1" value="108.87581">108.875812 E</geoCoordinate>
, 2019.IV,
<collectorName id="26E4532BDF62FFA916FC05A30A55FD98" box="[455,611,511,537]" pageId="16" pageNumber="191">Shi-Liang Mo</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B793CA0DF62FFA915A305A30D32FD98" ID-GBIF-Occurrence="3864839333" box="[664,1284,511,537]" collectionCode="ZMPC" collectorName="Shi-Liang Mo" county="Island" elevation="1412" latitude="18.710709" location="Mts. Jianfengling" longLatPrecision="1" longitude="108.87581" municipality="Ledong" pageId="16" pageNumber="191" specimenCount="9" specimenCount-female="2" specimenCount-male="7" stateProvince="Hainan" typeStatus="paratype">
<emphasis id="B965EAEFDF62FFA915A305A30B2FFD98" bold="true" box="[664,793,511,537]" pageId="16" pageNumber="191">
<typeStatus id="54AA885FDF62FFA915A305A30B22FD98" box="[664,788,511,537]" pageId="16" pageNumber="191" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D17FD74DF62FFA9142405A30B6FFD98" box="[799,857,511,537]" pageId="16" pageNumber="191" type="male">7♂♂</specimenCount>
,
<specimenCount id="9D17FD74DF62FFA9145F05A30BAAFD98" box="[868,924,511,537]" pageId="16" pageNumber="191" type="female">2♀♀</specimenCount>
(
<collectionCode id="ED00AE38DF62FFA9149205A30BCDFD98" box="[937,1019,511,537]" pageId="16" pageNumber="191">ZMPC</collectionCode>
), same data as holotype
</materialsCitation>
;
<materialsCitation id="3B793CA0DF62FFA9123505A308F5FDE0" ID-GBIF-Occurrence="3864839314" collectingDate="2019-04-14" collectingDateMax="2019-04-16" collectingDateMin="2019-04-14" collectionCode="ZMPC" collectorName="Hai-Tian Song" county="Island" elevation="1412" latitude="18.717" location="Jianfengling" longLatPrecision="7" longitude="108.8716" municipality="Ledong" pageId="16" pageNumber="191" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="9D17FD74DF62FFA9123505A30D07FD98" box="[1294,1329,511,537]" pageId="16" pageNumber="191" type="male">1♂</specimenCount>
(
<collectionCode id="ED00AE38DF62FFA9120505A30DA6FD98" box="[1342,1424,511,537]" pageId="16" pageNumber="191">ZMPC</collectionCode>
),
<collectingRegion id="49D5F81FDF62FFA917AC067F08DEFDBC" box="[151,232,547,573]" country="China" name="Hainan" pageId="16" pageNumber="191">Hainan</collectingRegion>
<collectingCounty id="62CF4E71DF62FFA917D5067F0903FDBC" box="[238,309,547,573]" pageId="16" pageNumber="191">Island</collectingCounty>
,
<collectingMunicipality id="6BCAAC87DF62FFA91604067F09A1FDBC" box="[319,407,547,573]" pageId="16" pageNumber="191">Ledong</collectingMunicipality>
,
<location id="8ECE6026DF62FFA9169A067F0A1DFDBC" LSID="urn:lsid:plazi:treatment:03B887EBDF62FFAA17AC05320BE3F83B:8ECE6026DF62FFA9169A067F0A1DFDBC" box="[417,555,547,573]" county="Island" latitude="18.717" longLatPrecision="7" longitude="108.8716" municipality="Ledong" name="Jianfengling" pageId="16" pageNumber="191" stateProvince="Hainan">Jianfengling</location>
main peak,
<date id="FFAF103DDF62FFA9158A067F0B43FDBC" box="[689,885,547,573]" pageId="16" pageNumber="191" value="2019-04-14" valueMax="2019-04-16" valueMin="2019-04-14">
<collectingDate id="EFEBE9D5DF62FFA9158A067F0B43FDBC" box="[689,885,547,573]" pageId="16" pageNumber="191" value="2019-04-14" valueMax="2019-04-16" valueMin="2019-04-14">2019-Apr.-14–16</collectingDate>
</date>
,
<quantity id="4CE99B18DF62FFA91444067F0BE3FDBC" box="[895,981,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.412" pageId="16" pageNumber="191" unit="m" value="1412.0">
<elevation id="003CD1CEDF62FFA91444067F0BE3FDBC" box="[895,981,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.412" pageId="16" pageNumber="191" unit="m" value="1412.0">1412 m</elevation>
</quantity>
,
<geoCoordinate id="EE25503ADF62FFA914DB067F0C6AFDBC" box="[992,1116,547,573]" degrees="18.7170" direction="north" orientation="latitude" pageId="16" pageNumber="191" precision="5" value="18.717">18.7170°N</geoCoordinate>
,
<geoCoordinate id="EE25503ADF62FFA9135C067F0CD9FDBC" box="[1127,1263,547,573]" degrees="108.8716" direction="east" orientation="longitude" pageId="16" pageNumber="191" precision="5" value="108.8716">108.8716°E</geoCoordinate>
,
<collectorName id="26E4532BDF62FFA913C2067F0DABFDBC" box="[1273,1437,547,573]" pageId="16" pageNumber="191">Hai-Tian Song</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B793CA0DF62FFA917EA061B0BF2FDE0" ID-GBIF-Occurrence="3864839310" box="[209,964,583,609]" collectionCode="CCPC" collectorName="Ying-Hui Li" elevation="1412" location="Jianfengling" pageId="16" pageNumber="191" specimenCount="2" specimenCount-male="2" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="9D17FD74DF62FFA917EA061B093FFDE0" box="[209,265,583,609]" pageId="16" pageNumber="191" type="male">2♂♂</specimenCount>
(
<collectionCode id="ED00AE38DF62FFA91622061B0950FDE0" box="[281,358,583,609]" pageId="16" pageNumber="191">CCPC</collectionCode>
),
<location id="8ECE6026DF62FFA91640061B0A3EFDE0" LSID="urn:lsid:plazi:treatment:03B887EBDF62FFAA17AC05320BE3F83B:8ECE6026DF62FFA91640061B0A3EFDE0" box="[379,520,583,609]" name="Jianfengling" pageId="16" pageNumber="191" stateProvince="Hainan">Jianfengling</location>
,
<quantity id="4CE99B18DF62FFA9152E061B0A58FDE0" box="[533,622,583,609]" metricMagnitude="3" metricUnit="m" metricValue="1.412" pageId="16" pageNumber="191" unit="m" value="1412.0">
<elevation id="003CD1CEDF62FFA9152E061B0A58FDE0" box="[533,622,583,609]" metricMagnitude="3" metricUnit="m" metricValue="1.412" pageId="16" pageNumber="191" unit="m" value="1412.0">1412 m</elevation>
</quantity>
, 2020.IV–V,
<collectorName id="26E4532BDF62FFA9143F061B0BA6FDE0" box="[772,912,583,609]" pageId="16" pageNumber="191">Ying-Hui Li</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B793CA0DF62FFA914E9061B0C90FD04" ID-GBIF-Occurrence="3864839320" collectingDate="2018-04-22" collectionCode="IZAS" collectorName="Bin Liu" elevation="1412" location="Jianfengling" municipality="Ledong" pageId="16" pageNumber="191" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="9D17FD74DF62FFA914E9061B0BC3FDE0" box="[978,1013,583,609]" pageId="16" pageNumber="191" type="male">1♂</specimenCount>
(
<collectionCode id="ED00AE38DF62FFA9133E061B0C7EFDE0" box="[1029,1096,583,609]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="16" pageNumber="191">IZAS</collectionCode>
),
<collectingRegion id="49D5F81FDF62FFA91365061B0CC2FDE0" box="[1118,1268,583,609]" country="China" name="Hainan" pageId="16" pageNumber="191">Hainan Prov.</collectingRegion>
,
<collectingMunicipality id="6BCAAC87DF62FFA9123A061B0D6FFDE0" box="[1281,1369,583,609]" pageId="16" pageNumber="191">Ledong</collectingMunicipality>
,
<location id="8ECE6026DF62FFA9125D061B08C2FD04" LSID="urn:lsid:plazi:treatment:03B887EBDF62FFAA17AC05320BE3F83B:8ECE6026DF62FFA9125D061B08C2FD04" municipality="Ledong" name="Jianfengling" pageId="16" pageNumber="191" stateProvince="Hainan">Jianfengling</location>
main peak,
<date id="FFAF103DDF62FFA9164406370A38FD04" box="[383,526,619,645]" pageId="16" pageNumber="191" value="2018-04-22">
<collectingDate id="EFEBE9D5DF62FFA9164406370A38FD04" box="[383,526,619,645]" pageId="16" pageNumber="191" value="2018-04-22">2018. IV. 22</collectingDate>
</date>
,
<quantity id="4CE99B18DF62FFA9152206370A47FD04" box="[537,625,619,645]" metricMagnitude="3" metricUnit="m" metricValue="1.412" pageId="16" pageNumber="191" unit="m" value="1412.0">
<elevation id="003CD1CEDF62FFA9152206370A47FD04" box="[537,625,619,645]" metricMagnitude="3" metricUnit="m" metricValue="1.412" pageId="16" pageNumber="191" unit="m" value="1412.0">1412 m</elevation>
</quantity>
, 18°43’0.85’’(N), 108°52’17.74’’(E),
<collectorName id="26E4532BDF62FFA9132506370C43FD04" box="[1054,1141,619,645]" pageId="16" pageNumber="191">Bin Liu</collectorName>
leg.
</materialsCitation>
IOZ(E) 1967013;
<materialsCitation id="3B793CA0DF62FFA9124206370AC1FD28" ID-GBIF-Occurrence="3864839304" collectingDate="2018-05-06" collectionCode="IZAS" collectorName="Bin Liu" elevation="1412" location="Jianfengling" municipality="Ledong" pageId="16" pageNumber="191" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="9D17FD74DF62FFA9124206370DAAFD04" box="[1401,1436,619,645]" pageId="16" pageNumber="191" type="male">1♂</specimenCount>
(
<collectionCode id="ED00AE38DF62FFA917A406D308D4FD28" box="[159,226,655,681]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="16" pageNumber="191">IZAS</collectionCode>
), ditto, but “
<date id="FFAF103DDF62FFA9165006D309E9FD28" box="[363,479,655,681]" pageId="16" pageNumber="191" value="2018-05-06">
<collectingDate id="EFEBE9D5DF62FFA9165006D309E9FD28" box="[363,479,655,681]" pageId="16" pageNumber="191" value="2018-05-06">2018. V. 6</collectingDate>
</date>
” and “IOZ(E) 1967014”
</materialsCitation>
;
<materialsCitation id="3B793CA0DF62FFA9143906D30DA1FD28" ID-GBIF-Occurrence="3864839308" box="[770,1431,654,681]" collectingDate="2018-05-11" collectionCode="IZAS" collectorName="Bin Liu" elevation="1412" location="Jianfengling" municipality="Ledong" pageId="16" pageNumber="191" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="9D17FD74DF62FFA9143906D30B13FD28" box="[770,805,655,681]" pageId="16" pageNumber="191" type="male">1♂</specimenCount>
(
<collectionCode id="ED00AE38DF62FFA9140806D30B40FD28" box="[819,886,655,681]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="16" pageNumber="191">IZAS</collectionCode>
), ditto, but “
<date id="FFAF103DDF62FFA914C406D30CB6FD28" box="[1023,1152,655,681]" pageId="16" pageNumber="191" value="2018-05-11">
<collectingDate id="EFEBE9D5DF62FFA914C406D30CB6FD28" box="[1023,1152,655,681]" pageId="16" pageNumber="191" value="2018-05-11">2018. V. 11</collectingDate>
</date>
” and “IOZ(E) 1967015”
</materialsCitation>
;
<materialsCitation id="3B793CA0DF62FFA917AC06EF0B70FD4C" ID-GBIF-Occurrence="3864839326" box="[151,838,691,717]" collectingDate="2018-04-28" collectionCode="IZAS" collectorName="Bin Liu" elevation="1412" location="Jianfengling" municipality="Ledong" pageId="16" pageNumber="191" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="9D17FD74DF62FFA917AC06EF088CFD4C" box="[151,186,691,717]" pageId="16" pageNumber="191" type="male">1♂</specimenCount>
(
<collectionCode id="ED00AE38DF62FFA917F106EF093BFD4C" box="[202,269,691,717]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="16" pageNumber="191">IZAS</collectionCode>
), ditto, but “
<date id="FFAF103DDF62FFA916A006EF0A1CFD4C" box="[411,554,691,717]" pageId="16" pageNumber="191" value="2018-04-28">
<collectingDate id="EFEBE9D5DF62FFA916A006EF0A1CFD4C" box="[411,554,691,717]" pageId="16" pageNumber="191" value="2018-04-28">2018. IV. 28</collectingDate>
</date>
” and “IOZ(E) 1967016”
</materialsCitation>
;
<materialsCitation id="3B793CA0DF62FFA9146906EF094FFD70" ID-GBIF-Occurrence="3864839305" collectingDate="2018-04-20" collectionCode="IZAS" collectorName="Bin Liu" elevation="1412" location="Jianfengling" municipality="Ledong" pageId="16" pageNumber="191" specimenCount="2" specimenCount-male="2" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="9D17FD74DF62FFA9146906EF0BBCFD4C" box="[850,906,691,717]" pageId="16" pageNumber="191" type="male">2♂♂</specimenCount>
(
<collectionCode id="ED00AE38DF62FFA914A106EF0BEBFD4C" box="[922,989,691,717]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="16" pageNumber="191">IZAS</collectionCode>
), ditto, but “
<date id="FFAF103DDF62FFA9135006EF0CCCFD4C" box="[1131,1274,691,717]" pageId="16" pageNumber="191" value="2018-04-20">
<collectingDate id="EFEBE9D5DF62FFA9135006EF0CCCFD4C" box="[1131,1274,691,717]" pageId="16" pageNumber="191" value="2018-04-20">2018. IV. 20</collectingDate>
</date>
” and “IOZ(E) 1967017–1967018”
</materialsCitation>
;
<materialsCitation id="3B793CA0DF62FFA916BF068B0CB4FD70" ID-GBIF-Occurrence="3864839302" box="[388,1154,727,753]" collectingDate="2009-04-19" collectionCode="CCPC" collectorName="Yu-Feng Hsu" location="Wuzhishan" pageId="16" pageNumber="191" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="9D17FD74DF62FFA916BF068B0991FD70" box="[388,423,727,753]" pageId="16" pageNumber="191" type="male">1♂</specimenCount>
(
<collectionCode id="ED00AE38DF62FFA9168D068B0A35FD70" box="[438,515,727,753]" pageId="16" pageNumber="191">CCPC</collectionCode>
),
<collectingRegion id="49D5F81FDF62FFA9152C068B0A5DFD70" box="[535,619,727,753]" country="China" name="Hainan" pageId="16" pageNumber="191">Hainan</collectingRegion>
,
<location id="8ECE6026DF62FFA9154E068B0AC3FD70" LSID="urn:lsid:plazi:treatment:03B887EBDF62FFAA17AC05320BE3F83B:8ECE6026DF62FFA9154E068B0AC3FD70" box="[629,757,727,753]" name="Wuzhishan" pageId="16" pageNumber="191" stateProvince="Hainan">Wuzhishan</location>
,
<date id="FFAF103DDF62FFA9143B068B0BB3FD70" box="[768,901,727,753]" pageId="16" pageNumber="191" value="2009-04-19">
<collectingDate id="EFEBE9D5DF62FFA9143B068B0BB3FD70" box="[768,901,727,753]" pageId="16" pageNumber="191" value="2009-04-19">2009-IV-19</collectingDate>
</date>
~24,
<collectorName id="26E4532BDF62FFA91481068B0C78FD70" box="[954,1102,727,753]" pageId="16" pageNumber="191">Yu-Feng Hsu</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B793CA0DF62FFA913B5068B0A6AFC94" ID-GBIF-Occurrence="3864839328" collectingDate="2011-04-25" collectionCode="CCPC" collectorName="Yi-Ting Chung" elevation="700" location="Wuzhishan" pageId="16" pageNumber="191" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="9D17FD74DF62FFA913B5068B0C86FD70" box="[1166,1200,727,753]" pageId="16" pageNumber="191" type="male">1♂</specimenCount>
(
<collectionCode id="ED00AE38DF62FFA91384068B0D3AFD70" box="[1215,1292,727,753]" pageId="16" pageNumber="191">CCPC</collectionCode>
),
<location id="8ECE6026DF62FFA91224068B0DAAFD70" LSID="urn:lsid:plazi:treatment:03B887EBDF62FFAA17AC05320BE3F83B:8ECE6026DF62FFA91224068B0DAAFD70" box="[1311,1436,727,753]" name="Wuzhishan" pageId="16" pageNumber="191" stateProvince="Hainan">Wuzhishan</location>
~
<quantity id="4CE99B18DF62FFA9179E06A708DFFC94" box="[165,233,763,789]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="16" pageNumber="191" unit="m" value="700.0">
<elevation id="003CD1CEDF62FFA9179E06A708DFFC94" box="[165,233,763,789]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="16" pageNumber="191" unit="m" value="700.0">700m</elevation>
</quantity>
,
<date id="FFAF103DDF62FFA917CF06A6094EFC94" box="[244,376,762,789]" pageId="16" pageNumber="191" value="2011-04-25">
<collectingDate id="EFEBE9D5DF62FFA917CF06A6094EFC94" box="[244,376,762,789]" pageId="16" pageNumber="191" value="2011-04-25">2011-IV-25</collectingDate>
</date>
,
<collectorName id="26E4532BDF62FFA916B906A70A1DFC94" box="[386,555,763,789]" pageId="16" pageNumber="191">Yi-Ting Chung</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30B6576DF62FFAA17FC07420D1AF9F7" lastPageId="19" lastPageNumber="194" pageId="16" pageNumber="191" type="description">
<paragraph id="8BAE36FDDF62FFA917FC07420D01FCB8" blockId="16.[151,1437,474,2013]" box="[199,1335,798,825]" pageId="16" pageNumber="191">
<emphasis id="B965EAEFDF62FFA917FC07420A38FCB8" bold="true" box="[199,526,798,825]" pageId="16" pageNumber="191">
Description.
<typeStatus id="54AA885FDF62FFA9165B074309FBFCB8" box="[352,461,799,825]" pageId="16" pageNumber="191" type="holotype">Holotype</typeStatus>
male
</emphasis>
(
<figureCitation id="132A2A78DF62FFA9152707430A99FCB8" box="[540,687,799,825]" captionStart="FIGURE 10" captionStartId="17.[151,250,1856,1880]" captionTargetBox="[162,1430,290,1814]" captionTargetId="figure-56@17.[151,1435,273,1833]" captionTargetPageId="17" captionText="FIGURE 10. Habitus of Anomala longilobata Zhao &amp; Zorn, new species. A–C, holotype; D–F, female paratype. A, D, dorsal view; B, E, lateral view; C, F, ventral view. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877291" httpUri="https://zenodo.org/record/6877291/files/figure.png" pageId="16" pageNumber="191">Figs. 10A–C</figureCitation>
,
<figureCitation id="132A2A78DF62FFA9158107430B3BFCB8" box="[698,781,799,825]" captionStart="FIGURE 12" captionStartId="19.[151,250,730,754]" captionTargetBox="[162,1428,184,697]" captionTargetId="figure-18@19.[151,1435,181,705]" captionTargetPageId="19" captionText="FIGURE 12. Male genitalia of Anomala spp. A–C, holotype of A. longilobata Zhao &amp; Zorn, new species; D–F, A. imperialis Arrow, 1899 (China: Guangxi). A, D, dorsal view; B, E, right lateral view; C, F, ventral view. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6877297" httpUri="https://zenodo.org/record/6877297/files/figure.png" pageId="16" pageNumber="191">12A–C</figureCitation>
). Body length:
<quantity id="4CE99B18DF62FFA9148007430C17FCB8" box="[955,1057,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="16" pageNumber="191" unit="mm" value="13.7">13.7 mm</quantity>
, greatest width: 8.0 mm.
</paragraph>
<paragraph id="8BAE36FDDF62FFA917FC071E0D5DFC6C" blockId="16.[151,1437,474,2013]" pageId="16" pageNumber="191">
<emphasis id="B965EAEFDF62FFA917FC071E09F6FCDC" bold="true" box="[199,448,834,861]" pageId="16" pageNumber="191">General appearance.</emphasis>
Body shape elongated ovoid, strongly convex. Generally dark green, with reddish sheen and weak metallic luster; marginal part of clypeus, mouth parts, lateral part of pronotum (except for a small spot at middle), ventral thoracic surface including coxae and femora (except for metasternal ventrite and anterior half of metepisternum), lateral part of abdominal ventrites and pygidium (except for basal and apical portions) yellow; antennomeres 1–4 brown, other antennomeres dark brown. Protibial teeth and claws of all legs reddish brown.
</paragraph>
<paragraph id="8BAE36FDDF62FFA917FC07AB09C7FBFC" blockId="16.[151,1437,474,2013]" pageId="16" pageNumber="191">
<emphasis id="B965EAEFDF62FFA917FC07AB0938FB90" bold="true" box="[199,270,1015,1041]" pageId="16" pageNumber="191">Head.</emphasis>
Clypeus subtrapezoidal, anterior corner broadly rounded; anterior margin feebly arched and moderately reflexed; surface with very dense and coarse punctures. Frontoclypeal suture distinct. Frons with very dense and coarse punctures, dense in vertex. Antennal club distinctly longer than antennomeres 2–6 combined. Inner margin of eye with several short setae.
</paragraph>
<paragraph id="8BAE36FDDF62FFA917FC00DB0BCBFA8C" blockId="16.[151,1437,474,2013]" pageId="16" pageNumber="191">
<emphasis id="B965EAEFDF62FFA917FC00DB0971FB20" bold="true" box="[199,327,1159,1185]" pageId="16" pageNumber="191">Pronotum.</emphasis>
Sides gently arched and convergent anteriad. Anterior angle acute and protruding, posterior angle blunt. Basal marginal line interrupted before scutellum, other marginal lines complete, lateral marginal line indistinct. With dense small punctures, punctures being coalescent at marginal portions; with a longitudinal medial furrow not reaching the base. Lateral margin with several moderately long setae.
</paragraph>
<paragraph id="8BAE36FDDF62FFA917FC014A09FDFAD4" blockId="16.[151,1437,474,2013]" pageId="16" pageNumber="191">
<emphasis id="B965EAEFDF62FFA917FC014A0970FAB1" bold="true" box="[199,326,1302,1328]" pageId="16" pageNumber="191">Scutellum.</emphasis>
Almost triangular, lateral margin feebly arched. Margins without setae, disc with dense, somewhat coalescent small punctures.
</paragraph>
<paragraph id="8BAE36FDDF62FFA917FC01030D31F918" blockId="16.[151,1437,474,2013]" pageId="16" pageNumber="191">
<emphasis id="B965EAEFDF62FFA917FC0103092CFAF8" bold="true" box="[199,282,1375,1401]" pageId="16" pageNumber="191">Elytra.</emphasis>
Intervals strongly convex; strial punctures very dense, annulated and small, usually strongly coalescent and larger at basal part of each stria and in lateral portions of each elytron; the whole surface with scattered minute punctures. Interstices I to IV each with a secondary stria; interstice I (the subsutural interstice) broadest, divided by a wide secondary stria into two costae, the secondary stria is formed by dense and coalescent large punctures; secondary stria of interstice II irregularly doubled, obsolete at basal third of elytra; secondary stria of interstice III emerging from internal portions of humeral umbone, irregularly doubled and wide at base, gradually narrower posteriad and obsolete before basal half of elytra; secondary stria on interstice IV shortly doubled at base, obsolete behind apical protuberance. Primary costae without additional striae. Humeral umbone and apical protuberance moderately bulging. Lateral carina distinct. Epipleura with a row of sparse short setae. Marginal membrane complete.
</paragraph>
<paragraph id="8BAE36FDDF62FFA917FC02FE08EBF960" blockId="16.[151,1437,474,2013]" pageId="16" pageNumber="191">
<emphasis id="B965EAEFDF62FFA917FC02FE095DF93D" bold="true" box="[199,363,1698,1724]" pageId="16" pageNumber="191">Propygidium.</emphasis>
Surface with dense transverse striolation. With a transverse row of sparse short setae proximally.
</paragraph>
<paragraph id="8BAE36FDDF62FFA917FC02B60920F8CC" blockId="16.[151,1437,474,2013]" pageId="16" pageNumber="191">
<emphasis id="B965EAEFDF62FFA917FC02B60974F885" bold="true" box="[199,322,1770,1796]" pageId="16" pageNumber="191">Pygidium.</emphasis>
Distinctly bulging, posterior margin arched. Surface with dense transverse striolation. With a short row of long setae proximally, and a transverse row of long setae at posterior margin near apex, disc with scattered short setae.
</paragraph>
<paragraph id="8BAE36FDDF62FFA917FC030B0AE6F838" blockId="16.[151,1437,474,2013]" pageId="16" pageNumber="191">
<emphasis id="B965EAEFDF62FFA917FC030B09C7F8F1" bold="true" box="[199,497,1878,1905]" pageId="16" pageNumber="191">Ventral thoracic surface.</emphasis>
Ventral pro- and mesothoracic surface with dense coalescent punctures and sparse short setae. Ventral metathoracic surface with dense large punctures and dense long setae, the punctures being smaller and sparser in the glabrous medial portion.
</paragraph>
<paragraph id="8BAE36FDDF62FFA817FC039E096EFF78" blockId="16.[151,1437,474,2013]" lastBlockId="17.[151,1437,151,249]" lastPageId="17" lastPageNumber="192" pageId="16" pageNumber="191">
<emphasis id="B965EAEFDF62FFA917FC039E09F5F85D" bold="true" box="[199,451,1986,2012]" pageId="16" pageNumber="191">Abdominal ventrites.</emphasis>
Ventrites 1–2 and anterior half of 3 weakly carinate laterally. Ventrites 2–5 with moderately dense transverse large punctures, usually coalescent near each side; ventrite 6 transversely striolated. Ventrites 2–5 each with a complete and transverse row of sparse, short or moderately long setae, the row of ventrite 6 at posterior margin.
</paragraph>
<caption id="DF6E6675DF63FFA817AC031C0B3BF8FD" ID-DOI="http://doi.org/10.5281/zenodo.6877291" ID-Zenodo-Dep="6877291" httpUri="https://zenodo.org/record/6877291/files/figure.png" pageId="17" pageNumber="192" startId="17.[151,250,1856,1880]" targetBox="[162,1430,290,1814]" targetPageId="17">
<paragraph id="8BAE36FDDF63FFA817AC031C0B3BF8FD" blockId="17.[151,1436,1856,1916]" pageId="17" pageNumber="192">
<emphasis id="B965EAEFDF63FFA817AC031C0916F8D9" bold="true" box="[151,288,1856,1880]" pageId="17" pageNumber="192">FIGURE 10.</emphasis>
Habitus of
<taxonomicName id="4C114D7EDF63FFA816A2031C0B37F8D9" authority="Zhao &amp; Zorn" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[409,769,1856,1880]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="longilobata" status="sp. nov.">
<emphasis id="B965EAEFDF63FFA816A2031C0A46F8D9" box="[409,624,1856,1880]" italics="true" pageId="17" pageNumber="192">Anomala longilobata</emphasis>
Zhao &amp; Zorn
</taxonomicName>
,
<emphasis id="B965EAEFDF63FFA81430031C0BBFF8D9" bold="true" box="[779,905,1856,1880]" pageId="17" pageNumber="192">
<taxonomicNameLabel id="A2565794DF63FFA81430031C0BBFF8D9" box="[779,905,1856,1880]" pageId="17" pageNumber="192" rank="species">new species</taxonomicNameLabel>
</emphasis>
. A–C, holotype; D–F, female paratype. A, D, dorsal view; B, E, lateral view; C, F, ventral view. Scale bar = 5 mm.
</paragraph>
</caption>
<caption id="DF6E6675DF60FFAB17AC03730C2AF8EA" ID-DOI="http://doi.org/10.5281/zenodo.6877295" ID-Zenodo-Dep="6877295" httpUri="https://zenodo.org/record/6877295/files/figure.png" pageId="18" pageNumber="193" startId="18.[151,250,1839,1863]" targetBox="[164,1424,193,1807]" targetPageId="18">
<paragraph id="8BAE36FDDF60FFAB17AC03730C2AF8EA" blockId="18.[151,1436,1839,1900]" pageId="18" pageNumber="193">
<emphasis id="B965EAEFDF60FFAB17AC0373092BF8C6" bold="true" box="[151,285,1839,1863]" pageId="18" pageNumber="193">FIGURE 11.</emphasis>
Habitus of
<taxonomicName id="4C114D7EDF60FFAB16AA036C0A28F8C6" box="[401,542,1840,1864]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="193" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B965EAEFDF60FFAB16AA036C09DAF8C9" box="[401,492,1840,1864]" italics="true" pageId="18" pageNumber="193">Anomala</emphasis>
spp.
</taxonomicName>
, in dorsal view. A–E, I, paratypes of
<taxonomicName id="4C114D7EDF60FFAB14B0036C0C91F8C6" authority="Zhao &amp; Zorn" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[907,1191,1840,1864]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="193" phylum="Arthropoda" rank="species" species="longilobata" status="sp. nov.">
<emphasis id="B965EAEFDF60FFAB14B0036C0C2DF8C9" box="[907,1051,1840,1864]" italics="true" pageId="18" pageNumber="193">A. longilobata</emphasis>
Zhao &amp; Zorn
</taxonomicName>
,
<emphasis id="B965EAEFDF60FFAB138B036C0D1DF8C6" bold="true" box="[1200,1323,1839,1864]" pageId="18" pageNumber="193">
<taxonomicNameLabel id="A2565794DF60FFAB138B036C0D1DF8C6" box="[1200,1323,1839,1864]" pageId="18" pageNumber="193" rank="species">new species</taxonomicNameLabel>
</emphasis>
; F–H, J–L,
<taxonomicName id="4C114D7EDF60FFAB17AC030809A9F8ED" authority="Arrow, 1899" authorityName="Arrow" authorityYear="1899" box="[151,415,1876,1900]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="193" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="B965EAEFDF60FFAB17AC0308092CF8ED" box="[151,282,1876,1900]" italics="true" pageId="18" pageNumber="193">A. imperialis</emphasis>
Arrow, 1899
</taxonomicName>
(China: Guangxi). A–H, male; I–L, female. Scale bar = 5 mm.
</paragraph>
</caption>
<caption id="DF6E6675DF61FFAA17AC06860D3FFC94" ID-DOI="http://doi.org/10.5281/zenodo.6877297" ID-Zenodo-Dep="6877297" httpUri="https://zenodo.org/record/6877297/files/figure.png" pageId="19" pageNumber="194" startId="19.[151,250,730,754]" targetBox="[162,1428,184,697]" targetPageId="19">
<paragraph id="8BAE36FDDF61FFAA17AC06860D3FFC94" blockId="19.[151,1436,730,790]" pageId="19" pageNumber="194">
<emphasis id="B965EAEFDF61FFAA17AC06860916FD73" bold="true" box="[151,288,730,754]" pageId="19" pageNumber="194">FIGURE 12.</emphasis>
Male genitalia of
<taxonomicName id="4C114D7EDF61FFAA16E606860AB4FD73" box="[477,642,730,754]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="194" phylum="Arthropoda" rank="species" species="undefined-a">
<emphasis id="B965EAEFDF61FFAA16E606860A0EFD73" box="[477,568,730,754]" italics="true" pageId="19" pageNumber="194">Anomala</emphasis>
spp. A
</taxonomicName>
–C, holotype of
<taxonomicName id="4C114D7EDF61FFAA141306860C7CFD70" authority="Zhao &amp; Zorn" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[808,1098,730,754]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="194" phylum="Arthropoda" rank="species" species="longilobata" status="sp. nov.">
<emphasis id="B965EAEFDF61FFAA141306860B8CFD73" box="[808,954,730,754]" italics="true" pageId="19" pageNumber="194">A. longilobata</emphasis>
Zhao &amp; Zorn
</taxonomicName>
,
<emphasis id="B965EAEFDF61FFAA136E06860CE5FD73" bold="true" box="[1109,1235,730,754]" pageId="19" pageNumber="194">
<taxonomicNameLabel id="A2565794DF61FFAA136E06860CE5FD73" box="[1109,1235,730,754]" pageId="19" pageNumber="194" rank="species">new species</taxonomicNameLabel>
</emphasis>
; D–F,
<taxonomicName id="4C114D7EDF61FFAA122206860921FC97" authority="Arrow, 1899" authorityName="Arrow" authorityYear="1899" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="194" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="B965EAEFDF61FFAA122206860DAAFD73" box="[1305,1436,730,754]" italics="true" pageId="19" pageNumber="194">A. imperialis</emphasis>
Arrow, 1899
</taxonomicName>
(China: Guangxi). A, D, dorsal view; B, E, right lateral view; C, F, ventral view. Scale bar = 2 mm.
</paragraph>
</caption>
<paragraph id="8BAE36FDDF61FFAA17FC07180D32FBFF" blockId="19.[151,1437,836,1979]" pageId="19" pageNumber="194">
<emphasis id="B965EAEFDF61FFAA17FC07180930FCDF" bold="true" box="[199,262,836,862]" pageId="19" pageNumber="194">Legs.</emphasis>
Protibia bidentate, both teeth distinct, blunt at apices; apical tooth extending to level of halfway of protarsomere 2. Inner spur situated at the level of the proximal tooth. Inner protarsal claw and outer mesotarsal claw split apically; lower branch of inner protarsal claw concave internobasally, longer and distinctly wider than the upper branch; lower branch of outer mesotarsal claw slightly wider than upper branch, almost equal in length. Each tarsomere 5 with an internomedial denticle, those of meso- and metatarsomere 5 small. Outer metatarsal claw slightly longer than inner one. Mesofemoral surface with transverse rows of long setae, several irregular rows between two regular rows at anterior margin and situated behind middle, the latter spiniform, a row at posterior margin regular with shorter setae. Metafemoral surface with three transverse rows of sparse short setae, the first row at anterior margin, a second row situated behind middle with robuster setae, and a third row at posterior margin.
</paragraph>
<paragraph id="8BAE36FDDF61FFAA17FC00D40A0DFB23" blockId="19.[151,1437,836,1979]" box="[199,571,1160,1186]" pageId="19" pageNumber="194">
<emphasis id="B965EAEFDF61FFAA17FC00D4094FFB23" bold="true" box="[199,377,1160,1186]" pageId="19" pageNumber="194">Male genitalia.</emphasis>
See
<figureCitation id="132A2A78DF61FFAA169400D40A00FB23" box="[431,566,1160,1186]" captionStart="FIGURE 12" captionStartId="19.[151,250,730,754]" captionTargetBox="[162,1428,184,697]" captionTargetId="figure-18@19.[151,1435,181,705]" captionTargetPageId="19" captionText="FIGURE 12. Male genitalia of Anomala spp. A–C, holotype of A. longilobata Zhao &amp; Zorn, new species; D–F, A. imperialis Arrow, 1899 (China: Guangxi). A, D, dorsal view; B, E, right lateral view; C, F, ventral view. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6877297" httpUri="https://zenodo.org/record/6877297/files/figure.png" pageId="19" pageNumber="194">Fig. 12A–C</figureCitation>
.
</paragraph>
<paragraph id="8BAE36FDDF61FFAA17FC00F00D1AF9F7" blockId="19.[151,1437,836,1979]" pageId="19" pageNumber="194">
<emphasis id="B965EAEFDF61FFAA17FC00F009A1FB47" bold="true" box="[199,407,1196,1222]" pageId="19" pageNumber="194">
<typeStatus id="54AA885FDF61FFAA17FC00F00977FB47" box="[199,321,1196,1222]" pageId="19" pageNumber="194" type="paratype">Paratypes</typeStatus>
. Males
</emphasis>
(
<figureCitation id="132A2A78DF61FFAA169C00F00A1BFB47" box="[423,557,1196,1222]" captionStart="FIGURE 11" captionStartId="18.[151,250,1839,1863]" captionTargetBox="[164,1424,193,1807]" captionTargetId="figure-21@18.[151,1435,181,1816]" captionTargetPageId="18" captionText="FIGURE 11. Habitus of Anomala spp., in dorsal view.A–E, I, paratypes of A. longilobata Zhao &amp; Zorn, new species; F–H, J–L, A. imperialis Arrow, 1899 (China: Guangxi). A–H, male; I–L, female. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877295" httpUri="https://zenodo.org/record/6877295/files/figure.png" pageId="19" pageNumber="194">Fig. 11A–E</figureCitation>
). Body length:
<quantity id="4CE99B18DF61FFAA15DB00F00BB1FB46" box="[736,903,1196,1223]" metricMagnitude="-2" metricUnit="m" metricValue="1.425" metricValueMax="1.44" metricValueMin="1.41" pageId="19" pageNumber="194" unit="mm" value="14.25" valueMax="14.4" valueMin="14.1">14.1–14.4 mm</quantity>
, greatest width:
<quantity id="4CE99B18DF61FFAA137F00F00CE6FB46" box="[1092,1232,1196,1223]" metricMagnitude="-3" metricUnit="m" metricValue="8.05" metricValueMax="8.3" metricValueMin="7.8" pageId="19" pageNumber="194" unit="mm" value="8.05" valueMax="8.3" valueMin="7.8">7.8–8.3 mm</quantity>
. External features and shape of parameres consistent but color variable. The general color varies from green, reddish green, purplish black, coal black (
<figureCitation id="132A2A78DF61FFAA165B00A809DFFA8F" box="[352,489,1268,1294]" captionStart="FIGURE 11" captionStartId="18.[151,250,1839,1863]" captionTargetBox="[164,1424,193,1807]" captionTargetId="figure-21@18.[151,1435,181,1816]" captionTargetPageId="18" captionText="FIGURE 11. Habitus of Anomala spp., in dorsal view.A–E, I, paratypes of A. longilobata Zhao &amp; Zorn, new species; F–H, J–L, A. imperialis Arrow, 1899 (China: Guangxi). A–H, male; I–L, female. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877295" httpUri="https://zenodo.org/record/6877295/files/figure.png" pageId="19" pageNumber="194">Fig. 11A–D</figureCitation>
), or even yellowish brown or reddish brown with two separated subtriangular patches in discal portion of pronotum (
<figureCitation id="132A2A78DF61FFAA16CD01440A6CFAB3" box="[502,602,1304,1330]" captionStart="FIGURE 11" captionStartId="18.[151,250,1839,1863]" captionTargetBox="[164,1424,193,1807]" captionTargetId="figure-21@18.[151,1435,181,1816]" captionTargetPageId="18" captionText="FIGURE 11. Habitus of Anomala spp., in dorsal view.A–E, I, paratypes of A. longilobata Zhao &amp; Zorn, new species; F–H, J–L, A. imperialis Arrow, 1899 (China: Guangxi). A–H, male; I–L, female. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877295" httpUri="https://zenodo.org/record/6877295/files/figure.png" pageId="19" pageNumber="194">Fig. 11E</figureCitation>
), one yellowish specimen lacks of any pronotal patch; ventral surface including pygidium and all femora sometimes entirely yellowish brown.
<emphasis id="B965EAEFDF61FFAA14FE01600C11FAD7" bold="true" box="[965,1063,1340,1366]" pageId="19" pageNumber="194">Females</emphasis>
(
<figureCitation id="132A2A78DF61FFAA130201600C8BFAD7" box="[1081,1213,1340,1366]" captionStart="FIGURE 10" captionStartId="17.[151,250,1856,1880]" captionTargetBox="[162,1430,290,1814]" captionTargetId="figure-56@17.[151,1435,273,1833]" captionTargetPageId="17" captionText="FIGURE 10. Habitus of Anomala longilobata Zhao &amp; Zorn, new species. A–C, holotype; D–F, female paratype. A, D, dorsal view; B, E, lateral view; C, F, ventral view. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877291" httpUri="https://zenodo.org/record/6877291/files/figure.png" pageId="19" pageNumber="194">Fig. 10D–F</figureCitation>
,
<figureCitation id="132A2A78DF61FFAA13F001600CC6FAD7" box="[1227,1264,1340,1366]" captionStart="FIGURE 11" captionStartId="18.[151,250,1839,1863]" captionTargetBox="[164,1424,193,1807]" captionTargetId="figure-21@18.[151,1435,181,1816]" captionTargetPageId="18" captionText="FIGURE 11. Habitus of Anomala spp., in dorsal view.A–E, I, paratypes of A. longilobata Zhao &amp; Zorn, new species; F–H, J–L, A. imperialis Arrow, 1899 (China: Guangxi). A–H, male; I–L, female. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877295" httpUri="https://zenodo.org/record/6877295/files/figure.png" pageId="19" pageNumber="194">11I</figureCitation>
). Body length:
<quantity id="4CE99B18DF61FFAA17AC013C090DFAFA" box="[151,315,1376,1403]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" metricValueMax="1.54" metricValueMin="1.44" pageId="19" pageNumber="194" unit="mm" value="14.9" valueMax="15.4" valueMin="14.4">14.4–15.4 mm</quantity>
, greatest width:
<quantity id="4CE99B18DF61FFAA16CB013C0A4FFAFA" box="[496,633,1376,1403]" metricMagnitude="-3" metricUnit="m" metricValue="8.899999999999999" metricValueMax="9.1" metricValueMin="8.7" pageId="19" pageNumber="194" unit="mm" value="8.899999999999999" valueMax="9.1" valueMin="8.7">8.7–9.1 mm</quantity>
. Generally similar to male but shape more broadened behind. Color generally yellowish brown with greenish sheen and weak metallic luster; each side of pronotum with a greenish patch located in a fovea. Antennal club slightly longer than antennomeres 1–6 combined. Basal marginal line distinct but less furrowed. Lateral carina slightly thicker. Pygidium almost triangular in shape and less bulging in lateral view. Setae on ventral surface distinctly denser and longer. Protibia slightly thinner than in male; apical protibial tooth round at apex, extending to level of base of protarsomere 3, proximal tooth blunt; inner spur situated at halfway of protibia; internomedial protuberance of all tarsomeres 5 small; upper and lower branches of inner protarsal claw and outer mesotarsal claw almost equal in length but the lower branches are slightly thicker; metatibia wider.
</paragraph>
</subSubSection>
<subSubSection id="C30B6576DF61FFAA17FC02DC0ADDF8CF" pageId="19" pageNumber="194" type="diagnosis">
<paragraph id="8BAE36FDDF61FFAA17FC02DC0ADDF8CF" blockId="19.[151,1437,836,1979]" pageId="19" pageNumber="194">
<emphasis id="B965EAEFDF61FFAA17FC02DC09F9F91B" bold="true" box="[199,463,1664,1690]" pageId="19" pageNumber="194">Differential diagnosis.</emphasis>
<taxonomicName id="4C114D7EDF61FFAA16EC02DC0B50F91A" authority="Zhao &amp; Zorn" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[471,870,1664,1691]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="194" phylum="Arthropoda" rank="species" species="longilobata" status="sp. nov.">
<emphasis id="B965EAEFDF61FFAA16EC02DC0AF3F91B" box="[471,709,1664,1690]" italics="true" pageId="19" pageNumber="194">Anomala longilobata</emphasis>
Zhao &amp; Zorn
</taxonomicName>
,
<emphasis id="B965EAEFDF61FFAA144802DC0BCBF91B" bold="true" box="[883,1021,1664,1690]" pageId="19" pageNumber="194">
<taxonomicNameLabel id="A2565794DF61FFAA144802DC0BCBF91B" box="[883,1021,1664,1690]" pageId="19" pageNumber="194" rank="species">new species</taxonomicNameLabel>
</emphasis>
is closely allied to
<taxonomicName id="4C114D7EDF61FFAA13E702DC093EF93F" authority="Arrow, 1899" authorityName="Arrow" authorityYear="1899" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="194" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="B965EAEFDF61FFAA13E702DC0D58F91B" box="[1244,1390,1664,1690]" italics="true" pageId="19" pageNumber="194">A. imperialis</emphasis>
Arrow, 1899
</taxonomicName>
(
<figureCitation id="132A2A78DF61FFAA162202F809CBF93F" box="[281,509,1700,1726]" captionStart="FIGURE 11" captionStartId="18.[151,250,1839,1863]" captionTargetBox="[164,1424,193,1807]" captionTargetId="figure-21@18.[151,1435,181,1816]" captionTargetPageId="18" captionText="FIGURE 11. Habitus of Anomala spp., in dorsal view.A–E, I, paratypes of A. longilobata Zhao &amp; Zorn, new species; F–H, J–L, A. imperialis Arrow, 1899 (China: Guangxi). A–H, male; I–L, female. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877295" httpUri="https://zenodo.org/record/6877295/files/figure.png" pageId="19" pageNumber="194">Figs. 11F–H, 11J–L</figureCitation>
,
<figureCitation id="132A2A78DF61FFAA153202F80A6CF93F" box="[521,602,1700,1726]" captionStart="FIGURE 12" captionStartId="19.[151,250,730,754]" captionTargetBox="[162,1428,184,697]" captionTargetId="figure-18@19.[151,1435,181,705]" captionTargetPageId="19" captionText="FIGURE 12. Male genitalia of Anomala spp. A–C, holotype of A. longilobata Zhao &amp; Zorn, new species; D–F, A. imperialis Arrow, 1899 (China: Guangxi). A, D, dorsal view; B, E, right lateral view; C, F, ventral view. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6877297" httpUri="https://zenodo.org/record/6877297/files/figure.png" pageId="19" pageNumber="194">12D–F</figureCitation>
;
<typeStatus id="54AA885FDF61FFAA155C02F90A88F93E" box="[615,702,1701,1727]" pageId="19" pageNumber="194" type="syntype">syntype</typeStatus>
examined) from
<collectingCountry id="F306766DDF61FFAA14B802F80BFFF93F" box="[899,969,1700,1726]" name="China" pageId="19" pageNumber="194">China</collectingCountry>
,
<collectingCountry id="F306766DDF61FFAA14EE02F90C38F93E" box="[981,1038,1701,1727]" name="Laos" pageId="19" pageNumber="194">Laos</collectingCountry>
, and
<collectingCountry id="F306766DDF61FFAA137002F80C98F93F" box="[1099,1198,1700,1726]" name="Vietnam" pageId="19" pageNumber="194">Vietnam</collectingCountry>
. The elytral intervals of
<taxonomicName id="4C114D7EDF61FFAA178E029409AEF963" authority="Zhao" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[181,408,1736,1762]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="194" phylum="Arthropoda" rank="species" species="longilobata" status="sp. nov.">
<emphasis id="B965EAEFDF61FFAA178E02940963F963" box="[181,341,1736,1762]" italics="true" pageId="19" pageNumber="194">A. longilobata</emphasis>
Zhao
</taxonomicName>
,
<emphasis id="B965EAEFDF61FFAA169802940A1AF963" bold="true" box="[419,556,1736,1762]" pageId="19" pageNumber="194">
<taxonomicNameLabel id="A2565794DF61FFAA169802940A1AF963" box="[419,556,1736,1762]" pageId="19" pageNumber="194" rank="species">new species</taxonomicNameLabel>
</emphasis>
are usually slightly more convex. The lower branches of parameres are longer, with the right one evenly pointed at apex (enlarged into a plate at apex in
<taxonomicName id="4C114D7EDF61FFAA14F702B00C6BF887" authorityName="Arrow" authorityYear="1899" box="[972,1117,1772,1798]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="194" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="B965EAEFDF61FFAA14F702B00C6BF887" box="[972,1117,1772,1798]" italics="true" pageId="19" pageNumber="194">A. imperialis</emphasis>
</taxonomicName>
). The ventral plate is simply rounded at apex but distinctly emarginated in
<emphasis id="B965EAEFDF61FFAA159F034C0B08F8AB" box="[676,830,1808,1834]" italics="true" pageId="19" pageNumber="194">
<taxonomicName id="4C114D7EDF61FFAA159F034C0B0CF8AB" authorityName="Arrow" authorityYear="1899" box="[676,826,1808,1834]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="194" phylum="Arthropoda" rank="species" species="imperialis">A. imperialis</taxonomicName>
.
</emphasis>
The generally greenish or purplish (pronotum green) female is only known for
<taxonomicName id="4C114D7EDF61FFAA168303680A7FF8CF" authorityName="Arrow" authorityYear="1899" box="[440,585,1844,1870]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="194" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="B965EAEFDF61FFAA168303680A7FF8CF" box="[440,585,1844,1870]" italics="true" pageId="19" pageNumber="194">A. imperialis</emphasis>
</taxonomicName>
(
<figureCitation id="132A2A78DF61FFAA156303680AE9F8CF" box="[600,735,1844,1870]" captionStart="FIGURE 11" captionStartId="18.[151,250,1839,1863]" captionTargetBox="[164,1424,193,1807]" captionTargetId="figure-21@18.[151,1435,181,1816]" captionTargetPageId="18" captionText="FIGURE 11. Habitus of Anomala spp., in dorsal view.A–E, I, paratypes of A. longilobata Zhao &amp; Zorn, new species; F–H, J–L, A. imperialis Arrow, 1899 (China: Guangxi). A–H, male; I–L, female. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877295" httpUri="https://zenodo.org/record/6877295/files/figure.png" pageId="19" pageNumber="194">Fig. 11K–L</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B6576DF61FFAA17FC03040B03F816" pageId="19" pageNumber="194" type="etymology">
<paragraph id="8BAE36FDDF61FFAA17FC03040B03F816" blockId="19.[151,1437,836,1979]" pageId="19" pageNumber="194">
<emphasis id="B965EAEFDF61FFAA17FC03040978F8F3" bold="true" box="[199,334,1880,1906]" pageId="19" pageNumber="194">Etymology.</emphasis>
The specific epithet derives from the Latin prefix “
<emphasis id="B965EAEFDF61FFAA149803040BD0F8F3" box="[931,998,1880,1906]" italics="true" pageId="19" pageNumber="194">longi-</emphasis>
” and the adjective “
<emphasis id="B965EAEFDF61FFAA13E803040DBFF8F3" box="[1235,1417,1880,1906]" italics="true" pageId="19" pageNumber="194">lobatus, -a, -um</emphasis>
”, alluding to the slender and lobed lower branch of paramere.
</paragraph>
</subSubSection>
<subSubSection id="C30B6576DF61FFAA17FC03FC0BE3F83B" box="[199,981,1952,1979]" pageId="19" pageNumber="194" type="distribution">
<paragraph id="8BAE36FDDF61FFAA17FC03FC0BE3F83B" blockId="19.[151,1437,836,1979]" box="[199,981,1952,1979]" pageId="19" pageNumber="194">
<emphasis id="B965EAEFDF61FFAA17FC03FC0957F83B" bold="true" box="[199,353,1952,1978]" pageId="19" pageNumber="194">Distribution.</emphasis>
<collectingRegion id="49D5F81FDF61FFAA165C03FC098EF83B" box="[359,440,1952,1978]" country="China" name="Hainan" pageId="19" pageNumber="194">Hainan</collectingRegion>
Island (Mts. Jianfengling and Mts. Wuzhishan).
</paragraph>
</subSubSection>
</treatment>
</document>
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