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<mods:title id="1EB067EA24816E43023BB5F39CCF1A4D">Systematics of the green lacewing tribe Ankylopterygini Navás, 1910 (Neuroptera: Chrysopidae: Chrysopinae) from China</mods:title>
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<mods:namePart id="4279BA721710F5A23E2DB7EC8A82C5CB">Wu, Jingyu</mods:namePart>
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<mods:affiliation id="3E49C0C9B351B0A6EB4C41EC3E1118F4">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659</mods:affiliation>
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<mods:affiliation id="B01818EF1EA050B72E89F80948BBCC52">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659</mods:affiliation>
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<subSubSection id="C30B650DFFE6FFE49EE7CB869CFF48AD" box="[151,985,1411,1438]" pageId="65" pageNumber="66" type="nomenclature">
<paragraph id="8BAE3686FFE6FFE49EE7CB869CFF48AD" blockId="65.[151,985,1411,1474]" box="[151,985,1411,1438]" pageId="65" pageNumber="66">
<emphasis id="B965EA94FFE6FFE49EE7CB869CFF48AD" bold="true" box="[151,985,1411,1438]" pageId="65" pageNumber="66">
[25]
<taxonomicName id="4C114D05FFE6FFE49EBFCB819C6D48AD" authority="(Yang &amp; Yang, 1989)" authorityName="Wu &amp; Liu" authorityYear="2024" baseAuthorityName="Yang &amp; Yang" baseAuthorityYear="1989" box="[207,843,1411,1438]" class="Insecta" family="Chrysopidae" genus="Tumeochrysa" kingdom="Animalia" order="Neuroptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="abunda" status="comb. nov." subGenus="Nineta">
<emphasis id="B965EA94FFE6FFE49EBFCB819E5548AD" bold="true" box="[207,371,1411,1438]" italics="true" pageId="65" pageNumber="66">Tumeochrysa</emphasis>
(
<emphasis id="B965EA94FFE6FFE49FF4CB869EF348AD" bold="true" box="[388,469,1412,1438]" italics="true" pageId="65" pageNumber="66">Nineta</emphasis>
)
<emphasis id="B965EA94FFE6FFE49F97CB819D6448AD" bold="true" box="[487,578,1411,1438]" italics="true" pageId="65" pageNumber="66">abunda</emphasis>
(
<bibRefCitation id="EF804B77FFE6FFE49C21CB869C6548AD" author="Yang, C. K. &amp; Yang, X. K." box="[593,835,1412,1438]" pageId="65" pageNumber="66" pagination="13 - 30" refId="ref91761" refString="Yang, C. K. &amp; Yang, X. K. (1989) Fourteen new species of green lacewings from Shaanxi Province (Neuroptera: Chrysopidae). Entomotaxonomia, 11, 13 - 30." type="journal article" year="1989">Yang &amp; Yang, 1989</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A25657EFFFE6FFE49D23CB869CFF48AD" box="[851,985,1412,1438]" pageId="65" pageNumber="66" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
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<subSubSection id="C30B650DFFE6FFE49EE7CBAA9E0148F1" box="[151,295,1448,1474]" pageId="65" pageNumber="66" type="description">
<paragraph id="8BAE3686FFE6FFE49EE7CBAA9E0148F1" blockId="65.[151,985,1411,1474]" box="[151,295,1448,1474]" pageId="65" pageNumber="66">
(
<figureCitation id="132A2A03FFE6FFE49EEFCBAA9FD048F1" box="[159,246,1448,1474]" captionStart="FIGURE 33" captionStartId="66.[151,250,1488,1512]" captionTargetBox="[169,1418,193,1449]" captionTargetId="figure-33@66.[151,1436,181,1464]" captionTargetPageId="66" captionText="FIGURE 33. Tumoechrysa (Nineta) abunda (Yang &amp; Yang, 1989) comb. nov. A. Dorsal habitus; B. Head, frontal view; C. Head, lateral view; D. Thorax, dorsal view. Scale bar: 5 mm (A), 1 mm (BD)." figureDoi="http://doi.org/10.5281/zenodo.14240596" httpUri="https://zenodo.org/record/14240596/files/figure.png" pageId="65" pageNumber="66">Figs 33</figureCitation>
,
<figureCitation id="132A2A03FFE6FFE49F72CBAA9E3948F1" box="[258,287,1448,1474]" captionStart="FIGURE 83" captionStartId="144.[151,250,1890,1914]" captionTargetBox="[197,1391,1096,1867]" captionTargetId="figure-46@144.[197,1391,1096,1867]" captionTargetPageId="144" captionText="FIGURE 83. Distribution map of species of Nineta from China." figureDoi="http://doi.org/10.5281/zenodo.14240712" httpUri="https://zenodo.org/record/14240712/files/figure.png" pageId="65" pageNumber="66">83</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFE6FFE49EE7CBEF9BBA4B37" box="[151,1180,1516,1541]" pageId="65" pageNumber="66" type="reference_group">
<paragraph id="8BAE3686FFE6FFE49EE7CBEF9BBA4B37" blockId="65.[151,1180,1516,1541]" box="[151,1180,1516,1541]" pageId="65" pageNumber="66">
<treatmentCitationGroup id="AB0111A8FFE6FFE49EE7CBEF9BBA4B37" box="[151,1180,1516,1541]" pageId="65" pageNumber="66">
<treatmentCitation id="0AB01097FFE6FFE49EE7CBEF9D054B36" author="Yang, C. K. &amp; Yang, X. K." box="[151,547,1516,1541]" page="24" pageId="65" pageNumber="66" year="1989">
<taxonomicName id="4C114D05FFE6FFE49EE7CBEF9D054B36" ID-CoL="6RZSK" authority="Yang &amp; Yang, 1989: 24" authorityName="Yang &amp; Yang" authorityPageNumber="24" authorityYear="1989" box="[151,547,1516,1541]" class="Insecta" family="Chrysopidae" genus="Nineta" kingdom="Animalia" order="Neuroptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="abunda">
<emphasis id="B965EA94FFE6FFE49EE7CBEF9E0B4B37" box="[151,301,1516,1540]" italics="true" pageId="65" pageNumber="66">Nineta abunda</emphasis>
<materialsCitation id="3B793CDBFFE6FFE49F42CBEF9D054B36" box="[306,547,1517,1541]" country="China" location="Qingquan" pageId="65" pageNumber="66" specimenCount="1" stateProvince="Shaanxi" typeStatus="holotype">
<bibRefCitation id="EF804B77FFE6FFE49F42CBEF9D054B36" author="Yang, C. K. &amp; Yang, X. K." box="[306,547,1517,1541]" pageId="65" pageNumber="66" pagination="13 - 30" refId="ref91761" refString="Yang, C. K. &amp; Yang, X. K. (1989) Fourteen new species of green lacewings from Shaanxi Province (Neuroptera: Chrysopidae). Entomotaxonomia, 11, 13 - 30." type="journal article" year="1989">Yang &amp; Yang, 1989: 24</bibRefCitation>
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. Type locality:
<collectingCountry id="F3067616FFE6FFE49CCFCBEE9DDA4B37" box="[703,764,1516,1540]" name="China" pageId="65" pageNumber="66">China</collectingCountry>
(
<collectingRegion id="49D5F864FFE6FFE49D79CBEE9C784B37" box="[777,862,1516,1540]" country="China" name="Shaanxi" pageId="65" pageNumber="66">Shaanxi</collectingRegion>
,
<location id="8ECE605DFFE6FFE49D18CBEE9CF74B37" LSID="urn:lsid:plazi:treatment:03B88790FFE6FFE79EE7CB869C694AC4:8ECE605DFFE6FFE49D18CBEE9CF74B37" box="[872,977,1516,1540]" country="China" name="Qingquan" pageId="65" pageNumber="66" stateProvince="Shaanxi">Qingquan</location>
).
<typeStatus id="54AA8824FFE6FFE49D91CBEE9B194B37" box="[993,1087,1516,1540]" pageId="65" pageNumber="66" type="holotype">Holotype</typeStatus>
in CAU.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFE6FFE49EE7C82E9E7D4B59" pageId="65" pageNumber="66" type="diagnosis">
<paragraph id="8BAE3686FFE6FFE49EE7C82E9E7D4B59" blockId="65.[151,1437,1580,2039]" pageId="65" pageNumber="66">
<emphasis id="B965EA94FFE6FFE49EE7C82E9E344B75" bold="true" box="[151,274,1580,1606]" pageId="65" pageNumber="66">Diagnosis.</emphasis>
This species is characterized by the body without markings, and the length of female subgenitale slightly longer than wide.
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFE6FFE79EB7C8769DA94B80" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66" type="description">
<paragraph id="8BAE3686FFE6FFE49EB7C8769A304BBC" blockId="65.[151,1437,1580,2039]" box="[199,1302,1652,1679]" pageId="65" pageNumber="66">
<emphasis id="B965EA94FFE6FFE49EB7C8769E7F4BBD" bold="true" box="[199,345,1652,1678]" pageId="65" pageNumber="66">Description.</emphasis>
Female. Body length
<quantity id="4CE99B63FFE6FFE49C21C8769D914BBC" box="[593,695,1652,1679]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="65" pageNumber="66" unit="mm" value="13.5">13.5 mm</quantity>
, forewing length
<quantity id="4CE99B63FFE6FFE49D0BC8769CC74BBC" box="[891,993,1652,1679]" metricMagnitude="-2" metricUnit="m" metricValue="1.83" pageId="65" pageNumber="66" unit="mm" value="18.3">18.3 mm</quantity>
, hindwing length
<quantity id="4CE99B63FFE6FFE49ADBC8769A344BBC" box="[1195,1298,1652,1679]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="65" pageNumber="66" unit="mm" value="16.8">16.8 mm</quantity>
.
</paragraph>
<paragraph id="8BAE3686FFE6FFE49EB7C89A9EB14BC9" blockId="65.[151,1437,1580,2039]" pageId="65" pageNumber="66">Head without spots, vertex raised. Compound eye prominent and black. Antennae yellow, scape and pedicel wider than flagellum. Frons, clypeus and gena without spots. Maxillary palpi with brown stripe on outer side, labial palpi brownish yellow.</paragraph>
<paragraph id="8BAE3686FFE6FFE49EB7C9079E214A71" blockId="65.[151,1437,1580,2039]" pageId="65" pageNumber="66">Pronotum anterior margin shorter than posterior margin, without markings. Mesonotum and metanotum without markings.</paragraph>
<paragraph id="8BAE3686FFE6FFE49EB7C94F9B6C4A55" blockId="65.[151,1437,1580,2039]" box="[199,1098,1868,1895]" pageId="65" pageNumber="66">Legs with dense setae, without spots. Pretarsal claws brown, with basal dilation.</paragraph>
<paragraph id="8BAE3686FFE6FFE49EB7C9729D764AC5" blockId="65.[151,1437,1580,2039]" pageId="65" pageNumber="66">
Forewing broad, without shadings. Pterostigma indistinct. Costal area with 30 crossveins; 19 radial crossveins present.
<emphasis id="B965EA94FFE6FFE49E87C9969E3F4A9D" box="[247,281,1940,1966]" italics="true" pageId="65" pageNumber="66">m1</emphasis>
smaller than
<emphasis id="B965EA94FFE6FFE49FC2C9969EF24A9D" box="[434,468,1940,1966]" italics="true" pageId="65" pageNumber="66">m2</emphasis>
;
<emphasis id="B965EA94FFE6FFE49F94C9979D264A9D" box="[484,512,1941,1966]" italics="true" pageId="65" pageNumber="66">im</emphasis>
ovoid; 11 psm-psc crossveins present.
<emphasis id="B965EA94FFE6FFE49DC9C9969CF24A9D" box="[953,980,1940,1966]" italics="true" pageId="65" pageNumber="66">c1</emphasis>
smaller than
<emphasis id="B965EA94FFE6FFE49A1DC9969BAE4A9D" box="[1133,1160,1940,1966]" italics="true" pageId="65" pageNumber="66">c2</emphasis>
,
<emphasis id="B965EA94FFE6FFE49AE6C9969B9B4A9D" box="[1174,1213,1940,1966]" italics="true" pageId="65" pageNumber="66">dcc</emphasis>
opened. A1 forked, A2 and A3 simple. Inner gradate series with 13 crossveins, basal crossvein of inner gradate series meeting Psm; outer gradate series with 12 crossveins.
</paragraph>
<caption id="DF6E660EFFE5FFE79EE7CBD29C904B3E" ID-DOI="http://doi.org/10.5281/zenodo.14240596" ID-Zenodo-Dep="14240596" httpUri="https://zenodo.org/record/14240596/files/figure.png" pageId="66" pageNumber="67" startId="66.[151,250,1488,1512]" targetBox="[169,1418,193,1449]" targetPageId="66" targetType="figure">
<paragraph id="8BAE3686FFE5FFE79EE7CBD29C904B3E" blockId="66.[151,1437,1488,1549]" pageId="66" pageNumber="67">
<emphasis id="B965EA94FFE5FFE79EE7CBD29E0448DB" bold="true" box="[151,290,1488,1512]" pageId="66" pageNumber="67">FIGURE 33.</emphasis>
<taxonomicName id="4C114D05FFE5FFE79F5ACBD29C6F48DA" authority="(Yang &amp; Yang, 1989)" authorityName="Wu &amp; Liu" authorityYear="2024" baseAuthorityName="Yang &amp; Yang" baseAuthorityYear="1989" box="[298,841,1488,1513]" class="Insecta" family="Chrysopidae" genus="Tumoechrysa" kingdom="Animalia" order="Neuroptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="abunda" status="comb. nov." subGenus="Nineta">
<emphasis id="B965EA94FFE5FFE79F5ACBD29E9748DB" box="[298,433,1488,1512]" italics="true" pageId="66" pageNumber="67">Tumoechrysa</emphasis>
(
<emphasis id="B965EA94FFE5FFE79FB2CBD39D2048DB" box="[450,518,1489,1512]" italics="true" pageId="66" pageNumber="67">Nineta</emphasis>
)
<emphasis id="B965EA94FFE5FFE79C66CBD29D4448DB" box="[534,610,1488,1512]" italics="true" pageId="66" pageNumber="67">abunda</emphasis>
(
<bibRefCitation id="EF804B77FFE5FFE79C03CBD39C6448DA" author="Yang, C. K. &amp; Yang, X. K." box="[627,834,1489,1513]" pageId="66" pageNumber="67" pagination="13 - 30" refId="ref91761" refString="Yang, C. K. &amp; Yang, X. K. (1989) Fourteen new species of green lacewings from Shaanxi Province (Neuroptera: Chrysopidae). Entomotaxonomia, 11, 13 - 30." type="journal article" year="1989">Yang &amp; Yang, 1989</bibRefCitation>
)
</taxonomicName>
<emphasis id="B965EA94FFE5FFE79D22CBD39CEE48DA" bold="true" box="[850,968,1489,1513]" pageId="66" pageNumber="67">
<taxonomicNameLabel id="A25657EFFFE5FFE79D22CBD39CEE48DA" box="[850,968,1489,1513]" pageId="66" pageNumber="67" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
A. Dorsal habitus; B. Head, frontal view; C. Head, lateral view; D. Thorax, dorsal view. Scale bar: 5 mm (A), 1 mm (BD).
</paragraph>
</caption>
<paragraph id="8BAE3686FFE5FFE79EB7C8539DA94B80" blockId="66.[151,1437,1617,2039]" pageId="66" pageNumber="67">Hindwing slightly narrow. Pterostigma indistinct. Costal area with 26 crossveins; 18 radial crossveins and 10 psm-psc crossveins present. Inner gradate series with 11 crossveins, basal crossvein of inner gradate series meeting Psm; outer gradate series with 10 crossveins.</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFE5FFE79EB7C8BF9FCA4BCF" pageId="66" pageNumber="67" type="materials_examined">
<paragraph id="8BAE3686FFE5FFE79EB7C8BF9FCA4BCF" blockId="66.[151,1437,1617,2039]" pageId="66" pageNumber="67">
<emphasis id="B965EA94FFE5FFE79EB7C8BF9E104BE4" bold="true" box="[199,310,1725,1751]" pageId="66" pageNumber="67">Material.</emphasis>
<materialsCitation id="3B793CDBFFE5FFE79F35C8BF9FCA4BCF" collectingDate="1971-06-28" collectionCode="CAU" country="China" county="Ganquan County" location="Qingquan" municipality="Yan'an" pageId="66" pageNumber="67" specimenCount="1" specimenCount-female="1" stateProvince="Shaanxi" typeStatus="holotype">
<typeStatus id="54AA8824FFE5FFE79F35C8BF9E8B4BE4" box="[325,429,1725,1751]" pageId="66" pageNumber="67" type="holotype">Holotype</typeStatus>
<specimenCount id="9D17FD0FFFE5FFE79FCBC8BC9EF64BEB" box="[443,464,1726,1752]" count="1" pageId="66" pageNumber="67" type="female"></specimenCount>
,
<collectingCountry id="F3067616FFE5FFE79F95C8BF9D0D4BE4" box="[485,555,1725,1751]" name="China" pageId="66" pageNumber="67">China</collectingCountry>
,
<collectingRegion id="49D5F864FFE5FFE79C4EC8BF9DBD4BE4" box="[574,667,1725,1751]" country="China" name="Shaanxi" pageId="66" pageNumber="67">Shaanxi</collectingRegion>
,
<collectingMunicipality id="6BCAACFCFFE5FFE79CDCC8BF9DD84BE4" box="[684,766,1725,1751]" pageId="66" pageNumber="67">Yanan</collectingMunicipality>
,
<collectingCounty id="62CF4E0AFFE5FFE79D61C8BF9CFE4BE4" box="[785,984,1725,1751]" pageId="66" pageNumber="67">Ganquan County</collectingCounty>
,
<location id="8ECE605DFFE5FFE79D9AC8BF9B7D4BE4" LSID="urn:lsid:plazi:treatment:03B88790FFE6FFE79EE7CB869C694AC4:8ECE605DFFE5FFE79D9AC8BF9B7D4BE4" box="[1002,1115,1725,1751]" country="China" county="Ganquan County" municipality="Yan'an" name="Qingquan" pageId="66" pageNumber="67" stateProvince="Shaanxi">Qingquan</location>
,
<date id="FFAF1046FFE5FFE79A1EC8BF9BD64BE4" box="[1134,1264,1725,1751]" pageId="66" pageNumber="67" value="1971-06-28">
<collectingDate id="EFEBE9AEFFE5FFE79A1EC8BF9BD64BE4" box="[1134,1264,1725,1751]" pageId="66" pageNumber="67" value="1971-06-28">1971.VI.28</collectingDate>
</date>
,
<location id="8ECE605DFFE5FFE79B73C8BF9ABA4BEB" LSID="urn:lsid:plazi:treatment:03B88790FFE6FFE79EE7CB869C694AC4:8ECE605DFFE5FFE79B73C8BF9ABA4BEB" box="[1283,1436,1725,1752]" country="China" county="Ganquan County" municipality="Yan'an" name="Chikun Yang" pageId="66" pageNumber="67" stateProvince="Shaanxi">Chikun Yang</location>
(
<collectionCode id="ED00AE43FFE5FFE79EEFC8E09FFA4BCF" box="[159,220,1762,1788]" country="China" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="66" pageNumber="67">CAU</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFE5FFE79EB7C9079D004A2C" box="[199,550,1797,1823]" pageId="66" pageNumber="67" type="distribution">
<paragraph id="8BAE3686FFE5FFE79EB7C9079D004A2C" blockId="66.[151,1437,1617,2039]" box="[199,550,1797,1823]" pageId="66" pageNumber="67">
<emphasis id="B965EA94FFE5FFE79EB7C9079E474A2C" bold="true" box="[199,353,1797,1823]" pageId="66" pageNumber="67">Distribution.</emphasis>
<collectingCountry id="F3067616FFE5FFE79F17C9079E8C4A2C" box="[359,426,1797,1823]" name="China" pageId="66" pageNumber="67">China</collectingCountry>
(
<collectingRegion id="49D5F864FFE5FFE79FC9C9079D3F4A2C" box="[441,537,1797,1823]" country="China" name="Shaanxi" pageId="66" pageNumber="67">Shaanxi</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFE5FFE79EB7C92B9C694AC4" pageId="66" pageNumber="67" type="discussion">
<paragraph id="8BAE3686FFE5FFE79EB7C92B9C694AC4" blockId="66.[151,1437,1617,2039]" pageId="66" pageNumber="67">
<emphasis id="B965EA94FFE5FFE79EB7C92B9E1D4A70" bold="true" box="[199,315,1833,1859]" pageId="66" pageNumber="67">Remarks.</emphasis>
This species was established based on a female specimen, and its genitalia may have been lost. The basal crossvein of inner gradate series of forewing and hindwing of
<typeStatus id="54AA8824FFE5FFE79DE4C94C9CE24A5B" box="[916,964,1870,1896]" pageId="66" pageNumber="67">type</typeStatus>
specimen attached to Psm directly, which only occurs occasionally in
<taxonomicName id="4C114D05FFE5FFE79FA4C9709D394AB8" authorityName="Navas" authorityYear="1912" box="[468,543,1906,1931]" class="Insecta" family="Chrysopidae" genus="Nineta" kingdom="Animalia" order="Neuroptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFE5FFE79FA4C9709D394AB8" box="[468,543,1906,1931]" italics="true" pageId="66" pageNumber="67">Nineta</emphasis>
</taxonomicName>
. Other morphological characters of this species are consistent with
<taxonomicName id="4C114D05FFE5FFE79B50C9709A4D4AB8" authorityName="Navas" authorityYear="1912" box="[1312,1387,1906,1931]" class="Insecta" family="Chrysopidae" genus="Nineta" kingdom="Animalia" order="Neuroptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFE5FFE79B50C9709A4D4AB8" box="[1312,1387,1906,1931]" italics="true" pageId="66" pageNumber="67">Nineta</emphasis>
</taxonomicName>
: the wide head, the broad antennal scape, the thorax without markings, the wings without shadings and the body without markings. We consider this species as a valid species of
<taxonomicName id="4C114D05FFE5FFE79D54C9B89C494AE0" authorityName="Navas" authorityYear="1912" box="[804,879,1978,2003]" class="Insecta" family="Chrysopidae" genus="Nineta" kingdom="Animalia" order="Neuroptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFE5FFE79D54C9B89C494AE0" box="[804,879,1978,2003]" italics="true" pageId="66" pageNumber="67">Nineta</emphasis>
</taxonomicName>
. According to original description, the length of female subgenitale of this species is slightly longer than wide.
</paragraph>
</subSubSection>
</treatment>
</document>