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<mods:title id="C88C9749D988805E2BA5AECB9EE2AAD4">Systematics of the green lacewing tribe Ankylopterygini Navás, 1910 (Neuroptera: Chrysopidae: Chrysopinae) from China</mods:title>
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<mods:affiliation id="C69DFFFB0B3A210C3439EAEE7556BBD6">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659</mods:affiliation>
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<paragraph id="8BAE3686FFDEFFDC9EE7CD899D4B4E97" blockId="121.[151,621,906,969]" box="[151,621,906,933]" pageId="121" pageNumber="122">
<emphasis id="B965EA94FFDEFFDC9EE7CD899D4B4E97" bold="true" box="[151,621,906,933]" pageId="121" pageNumber="122">
[57]
<taxonomicName id="4C114D05FFDEFFDC9EBFCD899D4B4E97" ID-CoL="6X48R" authority="Brooks, 1986" authorityName="Brooks" authorityYear="1986" box="[207,621,906,933]" class="Insecta" family="Chrysopidae" genus="Retipenna" kingdom="Animalia" order="Neuroptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="variegata">
<emphasis id="B965EA94FFDEFFDC9EBFCD899EE54E96" bold="true" box="[207,451,907,933]" italics="true" pageId="121" pageNumber="122">Retipenna variegata</emphasis>
<bibRefCitation id="EF804B77FFDEFFDC9FBACD899D4B4E97" author="Brooks, S. J." box="[458,621,906,933]" pageId="121" pageNumber="122" pagination="35 - 48" refId="ref88537" refString="Brooks, S. J. (1986) A new genus of Ankylopterygini (Chrysopidae). Neuroptera International, 4, 35 - 48." type="journal article" year="1986">Brooks, 1986</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
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<subSubSection id="C30B650DFFDEFFDC9EE7CDAD9E014EFA" box="[151,295,943,969]" pageId="121" pageNumber="122" type="description">
<paragraph id="8BAE3686FFDEFFDC9EE7CDAD9E014EFA" blockId="121.[151,621,906,969]" box="[151,295,943,969]" pageId="121" pageNumber="122">
(
<figureCitation id="132A2A03FFDEFFDC9EEFCDAD9FD04EFA" box="[159,246,943,969]" captionStart="FIGURE 67" captionStartId="122.[151,250,1940,1964]" captionTargetBox="[174,1387,200,1895]" captionTargetId="figure-33@122.[164,1423,181,1917]" captionTargetPageId="122" captionText="FIGURE 67. Retipenna variegata Brooks, 1986. A. Dorsal habitus; B. Head, frontal view; C. Head, lateral view; D. Thorax, dorsal view; E. Male genitalia, lateral view; F. Gonarcal complex, dorsal view; G. Gonarcal complex, lateral view; H. Arcessus; I. Female genitalia, lateral view; J. Subgenitale; K. Spermatheca. Scale bar: 5 mm (A), 1 mm (BD)." figureDoi="http://doi.org/10.5281/zenodo.14240680" httpUri="https://zenodo.org/record/14240680/files/figure.png" pageId="121" pageNumber="122">Figs 67</figureCitation>
,
<figureCitation id="132A2A03FFDEFFDC9F72CDAD9E394EFA" box="[258,287,943,969]" captionStart="FIGURE 87" captionStartId="146.[151,250,1890,1914]" captionTargetBox="[197,1391,1096,1867]" captionTargetId="figure-46@146.[197,1391,1096,1867]" captionTargetPageId="146" captionText="FIGURE 87. Distribution map of species of Retipenna from China." figureDoi="http://doi.org/10.5281/zenodo.14240720" httpUri="https://zenodo.org/record/14240720/files/figure.png" pageId="121" pageNumber="122">87</figureCitation>
)
</paragraph>
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<paragraph id="8BAE3686FFDEFFDC9EE7CDF69B724938" blockId="121.[151,1108,1011,1036]" box="[151,1108,1011,1036]" pageId="121" pageNumber="122">
<treatmentCitationGroup id="AB0111A8FFDEFFDC9EE7CDF69B724938" box="[151,1108,1011,1036]" pageId="121" pageNumber="122">
<treatmentCitation id="0AB01097FFDEFFDC9EE7CDF69D384938" author="Brooks, S. J." box="[151,542,1011,1035]" page="67" pageId="121" pageNumber="122" year="1986">
<taxonomicName id="4C114D05FFDEFFDC9EE7CDF69D384938" ID-CoL="6X48R" authority="Brooks, 1986: 67" authorityName="Brooks" authorityPageNumber="67" authorityYear="1986" box="[151,542,1011,1035]" class="Insecta" family="Chrysopidae" genus="Retipenna" kingdom="Animalia" order="Neuroptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="variegata">
<emphasis id="B965EA94FFDEFFDC9EE7CDF69E434938" box="[151,357,1012,1035]" italics="true" pageId="121" pageNumber="122">Retipenna variegata</emphasis>
<bibRefCitation id="EF804B77FFDEFFDC9F1CCDF19D384938" author="Brooks, S. J." box="[364,542,1011,1035]" pageId="121" pageNumber="122" pagination="35 - 48" refId="ref88537" refString="Brooks, S. J. (1986) A new genus of Ankylopterygini (Chrysopidae). Neuroptera International, 4, 35 - 48." type="journal article" year="1986">Brooks, 1986: 67</bibRefCitation>
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. Type locality:
<collectingCountry id="F3067616FFDEFFDC9CC9CDF19DCB4938" box="[697,749,1011,1035]" name="India" pageId="121" pageNumber="122">India</collectingCountry>
(Khasi Hills).
<typeStatus id="54AA8824FFDEFFDC9DF4CDF19CC44938" box="[900,994,1011,1035]" pageId="121" pageNumber="122" type="holotype">Holotype</typeStatus>
in NHML.
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</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFDEFFDC9EE7CA3B9B6749A8" pageId="121" pageNumber="122" type="diagnosis">
<paragraph id="8BAE3686FFDEFFDC9EE7CA3B9B6749A8" blockId="121.[151,1437,1081,1971]" pageId="121" pageNumber="122">
<emphasis id="B965EA94FFDEFFDC9EE7CA3B9E344960" bold="true" box="[151,274,1081,1107]" pageId="121" pageNumber="122">Diagnosis.</emphasis>
This species is characterized by the head with spots on the gena, the thorax without markings, the dark brown veins in basal areas and gradate series of fore- and hindwings, the median arch of male gonarcus with shallow U-shaped invagination, the long narrow arcessus rounded at base and pointed at tip.
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFDEFFDC9EB7CAA79C7F4A80" pageId="121" pageNumber="122" type="description">
<paragraph id="8BAE3686FFDEFFDC9EB7CAA79BC049F3" blockId="121.[151,1437,1081,1971]" box="[199,1254,1189,1216]" pageId="121" pageNumber="122">
<emphasis id="B965EA94FFDEFFDC9EB7CAA79E7F498C" bold="true" box="[199,345,1189,1215]" pageId="121" pageNumber="122">Description.</emphasis>
Male. Body length
<quantity id="4CE99B63FFDEFFDC9C48CAA79DA149F3" box="[568,647,1189,1216]" metricMagnitude="1" metricUnit="m" metricValue="1.18" pageId="121" pageNumber="122" unit="m" value="11.8">11.8 m</quantity>
, forewing length
<quantity id="4CE99B63FFDEFFDC9D3CCAA79C9449F3" box="[844,946,1189,1216]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="121" pageNumber="122" unit="mm" value="14.7">14.7 mm</quantity>
, hindwing length
<quantity id="4CE99B63FFDEFFDC9A0BCAA79BC449F3" box="[1147,1250,1189,1216]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="121" pageNumber="122" unit="mm" value="13.3">13.3 mm</quantity>
.
</paragraph>
<paragraph id="8BAE3686FFDEFFDC9EB7CACB9FCF4818" blockId="121.[151,1437,1081,1971]" pageId="121" pageNumber="122">Head yellow, vertex raised. Compound eyes prominent and black. Antennae yellow, scape and pedicel wider than flagellum. Frons and clypeus not marked. Gena marked black spots. Maxillary and labial palpi brownish yellow.</paragraph>
<paragraph id="8BAE3686FFDEFFDC9EB7CB349A68487C" blockId="121.[151,1437,1081,1971]" box="[199,1358,1333,1360]" pageId="121" pageNumber="122">Pronotum anterior margin shorter than posterior margin, mesonotum and metanotum without markings.</paragraph>
<paragraph id="8BAE3686FFDEFFDC9EB7CB589C804840" blockId="121.[151,1437,1081,1971]" box="[199,934,1369,1396]" pageId="121" pageNumber="122">Legs with dense setae. Pretarsal claws brown, with basal dilation.</paragraph>
<paragraph id="8BAE3686FFDEFFDC9EB7CB7F9E8E4B14" blockId="121.[151,1437,1081,1971]" pageId="121" pageNumber="122">
Forewing slightly narrow, dark brown veins present in three areas: veins in basal area, inner gradate series and apical radial crossveins, outer gradates and forking marginal veins. Pterostigma pale brown. Costal area with 2025 crossveins; 1215 radial crossveins present.
<emphasis id="B965EA94FFDEFFDC9CFFCBC79D9748EC" box="[655,689,1477,1503]" italics="true" pageId="121" pageNumber="122">m1</emphasis>
smaller than
<emphasis id="B965EA94FFDEFFDC9D3CCBC79C4848EC" box="[844,878,1477,1503]" italics="true" pageId="121" pageNumber="122">m2</emphasis>
;
<emphasis id="B965EA94FFDEFFDC9D0ECBC49CBC48EC" box="[894,922,1478,1503]" italics="true" pageId="121" pageNumber="122">im</emphasis>
ovoid; 1011 psm-psc crossveins present.
<emphasis id="B965EA94FFDEFFDC9BF2CBC79ABB48EC" box="[1410,1437,1477,1503]" italics="true" pageId="121" pageNumber="122">c1</emphasis>
smaller than
<emphasis id="B965EA94FFDEFFDC9F5ACBEB9E624B30" box="[298,324,1513,1539]" italics="true" pageId="121" pageNumber="122">c2</emphasis>
,
<emphasis id="B965EA94FFDEFFDC9F23CBEB9E5C4B30" box="[339,378,1513,1539]" italics="true" pageId="121" pageNumber="122">dcc</emphasis>
closed. A1 forked, A2 and A3 simple. Inner gradate series with 67 crossveins, outer gradate series with 8 crossveins.
</paragraph>
<paragraph id="8BAE3686FFDEFFDC9EB7C8339E474BA0" blockId="121.[151,1437,1081,1971]" pageId="121" pageNumber="122">Hindwing narrow, apical radial crossveins and gradate series brown. Costal area with 1517 crossveins; 1314 radial crossveins and 8 psm-psc crossveins present. Inner gradate series with 67 crossveins, outer gradate series with 7 crossveins.</paragraph>
<paragraph id="8BAE3686FFDEFFDC9EB7C89F9A714A10" blockId="121.[151,1437,1081,1971]" pageId="121" pageNumber="122">Abdomen with dense setae. Tergum VII and sternum VII subquadrate in lateral view; tergum VIII slightly shorter than tergum VII; sternum VIII fused with sternum IX and with small tubercle near tip; tergum IX fused with ectoproct. Callus cerci ovoid. Gonarcus arcuate, median arch with shallow U-shaped invagination, lateral arms narrow at base and wide at tip; entoprocessus widest at tip; arcessus long, rounded at base and pointed at tip.</paragraph>
<paragraph id="8BAE3686FFDEFFDC9EB7C92F9B5B4A7B" blockId="121.[151,1437,1081,1971]" box="[199,1149,1837,1864]" pageId="121" pageNumber="122">Female. Body length 10.0 mm, forewing length 16.0 mm, hindwing length 14.0 mm.</paragraph>
<paragraph id="8BAE3686FFDEFFDC9EB7C9539C7F4A80" blockId="121.[151,1437,1081,1971]" pageId="121" pageNumber="122">Sternum VII longer than tergum VII; tergum VII subquadrate in lateral view; tergum VIII shorter than tergum VII; tergum IX fused with ectoproct. Callus cerci ovoid. Subgenitale bilobed apically; spermatheca thick, vela extremely short, ventral impression deep, duct long and coiled.</paragraph>
</subSubSection>
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<paragraph id="8BAE3686FFDDFFDF9EE7C9969BB54AC6" blockId="122.[151,1437,1940,2037]" pageId="122" pageNumber="123">
<emphasis id="B965EA94FFDDFFDF9EE7C9969E074A9E" bold="true" box="[151,289,1940,1965]" pageId="122" pageNumber="123">FIGURE 67.</emphasis>
<taxonomicName id="4C114D05FFDDFFDF9F58C9979DAA4A9E" authority="Brooks, 1986" authorityName="Brooks" authorityYear="1986" box="[296,652,1941,1965]" class="Insecta" family="Chrysopidae" genus="Retipenna" kingdom="Animalia" order="Neuroptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="species" species="variegata">
<emphasis id="B965EA94FFDDFFDF9F58C9979ED14A9F" box="[296,503,1941,1964]" italics="true" pageId="122" pageNumber="123">Retipenna variegata</emphasis>
<bibRefCitation id="EF804B77FFDDFFDF9F8FC9979DAA4A9E" author="Brooks, S. J." box="[511,652,1941,1965]" pageId="122" pageNumber="123" pagination="35 - 48" refId="ref88537" refString="Brooks, S. J. (1986) A new genus of Ankylopterygini (Chrysopidae). Neuroptera International, 4, 35 - 48." type="journal article" year="1986">Brooks, 1986</bibRefCitation>
</taxonomicName>
. A. Dorsal habitus; B. Head, frontal view; C. Head, lateral view; D. Thorax, dorsal view; E. Male genitalia, lateral view; F. Gonarcal complex, dorsal view; G. Gonarcal complex, lateral view; H. Arcessus; I. Female genitalia, lateral view; J. Subgenitale; K. Spermatheca. Scale bar: 5 mm (A), 1 mm (BD).
</paragraph>
</caption>
<subSubSection id="C30B650DFFDCFFDE9EB7CE949DEA4DE6" pageId="123" pageNumber="124" type="materials_examined">
<paragraph id="8BAE3686FFDCFFDE9EB7CE949DEA4DE6" blockId="123.[151,1437,150,321]" pageId="123" pageNumber="124">
<emphasis id="B965EA94FFDCFFDE9EB7CE949E104D83" bold="true" box="[199,310,150,176]" pageId="123" pageNumber="124">Material.</emphasis>
<materialsCitation id="3B793CDBFFDCFFDE9F31CE959CFC4D82" box="[321,986,151,177]" collectionCode="CAU" collectorName="Yingjiang &amp; Hulukou" country="China" location="Yunnan" pageId="123" pageNumber="124" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="9D17FD0FFFDCFFDE9F31CE959E434D82" box="[321,357,151,177]" count="1" pageId="123" pageNumber="124" type="male">1♂</specimenCount>
,
<collectingCountry id="F3067616FFDCFFDE9F06CE959E9A4D82" box="[374,444,151,177]" name="China" pageId="123" pageNumber="124">China</collectingCountry>
,
<collectingRegion id="49D5F864FFDCFFDE9FB9CE959D044D82" box="[457,546,151,177]" country="China" name="Yunnan" pageId="123" pageNumber="124">Yunnan</collectingRegion>
,
<collectorName id="26E45350FFDCFFDE9C5FCE959D864D82" box="[559,672,151,177]" pageId="123" pageNumber="124">Yingjiang</collectorName>
,
<collectorName id="26E45350FFDCFFDE9CDFCE959C324D82" box="[687,788,151,177]" pageId="123" pageNumber="124">Hulukou</collectorName>
, 2018.IV (
<collectionCode id="ED00AE43FFDCFFDE9DE1CE959CF44D82" box="[913,978,151,177]" country="China" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="123" pageNumber="124">CAU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B793CDBFFDCFFDE9D99CE959DEA4DE6" collectingDate="2019-05-30" collectionCode="CAU" collectorName="Rongrong Shen &amp; Yingnan He &amp; Distribution. &amp; Khasi Hills &amp; Remarks. &amp; This" country="China" elevation="778" location="Medog" pageId="123" pageNumber="124" specimenCount="1" specimenCount-female="1" stateProvince="Xizang">
<specimenCount id="9D17FD0FFFDCFFDE9D99CE959B2A4D82" box="[1001,1036,151,177]" count="1" pageId="123" pageNumber="124" type="female">1♀</specimenCount>
,
<collectingCountry id="F3067616FFDCFFDE9A6ECE959B424D82" box="[1054,1124,151,177]" name="China" pageId="123" pageNumber="124">China</collectingCountry>
,
<collectingRegion id="49D5F864FFDCFFDE9A02CE959BE04D82" box="[1138,1222,151,177]" country="China" name="Xizang" pageId="123" pageNumber="124">Xizang</collectingRegion>
,
<location id="8ECE605DFFDCFFDE9AA5CE959A014D82" LSID="urn:lsid:plazi:treatment:03B88790FFDEFFDE9EE7CD899DAC4C72:8ECE605DFFDCFFDE9AA5CE959A014D82" box="[1237,1319,151,177]" country="China" name="Medog" pageId="123" pageNumber="124" stateProvince="Xizang">Mêdog</location>
,
<quantity id="4CE99B63FFDCFFDE9B46CE959AB14D82" box="[1334,1431,151,177]" metricMagnitude="2" metricUnit="m" metricValue="7.778999999999999" pageId="123" pageNumber="124" unit="m" value="777.9">
<elevation id="003CD1B5FFDCFFDE9B46CE959AB14D82" box="[1334,1431,151,177]" metricMagnitude="2" metricUnit="m" metricValue="7.778999999999999" pageId="123" pageNumber="124" unit="m" value="777.9">777.9 m</elevation>
</quantity>
,
<date id="FFAF1046FFDCFFDE9EE7CEB99E2A4DE6" box="[151,268,187,213]" pageId="123" pageNumber="124" value="2019-05-30">
<collectingDate id="EFEBE9AEFFDCFFDE9EE7CEB99E2A4DE6" box="[151,268,187,213]" pageId="123" pageNumber="124" value="2019-05-30">2019.V.30</collectingDate>
</date>
,
<collectorName id="26E45350FFDCFFDE9F68CEB99EE14DE6" box="[280,455,187,213]" pageId="123" pageNumber="124">Rongrong Shen</collectorName>
&amp;
<collectorName id="26E45350FFDCFFDE9F9ACEB99D564DE6" box="[490,624,187,213]" pageId="123" pageNumber="124">Yingnan He</collectorName>
(
<collectionCode id="ED00AE43FFDCFFDE9C0FCEB99D9A4DE6" box="[639,700,187,213]" country="China" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="123" pageNumber="124">CAU</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFDCFFDE9EB7CEDC9C794DCA" box="[199,863,222,249]" pageId="123" pageNumber="124" type="distribution">
<paragraph id="8BAE3686FFDCFFDE9EB7CEDC9C794DCA" blockId="123.[151,1437,150,321]" box="[199,863,222,249]" pageId="123" pageNumber="124">
<emphasis id="B965EA94FFDCFFDE9EB7CEDC9E474DCB" bold="true" box="[199,353,222,248]" pageId="123" pageNumber="124">
<collectorName id="26E45350FFDCFFDE9EB7CEDC9E474DCB" box="[199,353,222,248]" pageId="123" pageNumber="124">Distribution.</collectorName>
</emphasis>
<collectingCountry id="F3067616FFDCFFDE9F17CEDD9E8C4DCA" box="[359,426,223,249]" name="China" pageId="123" pageNumber="124">China</collectingCountry>
(
<collectingRegion id="49D5F864FFDCFFDE9FC9CEDD9D284DCA" box="[441,526,223,249]" country="China" name="Xizang" pageId="123" pageNumber="124">Xizang</collectingRegion>
,
<collectingRegion id="49D5F864FFDCFFDE9C68CEDD9D554DCA" box="[536,627,223,249]" country="China" name="Yunnan" pageId="123" pageNumber="124">Yunnan</collectingRegion>
);
<collectingCountry id="F3067616FFDCFFDE9CF6CEDD9DE64DCA" box="[646,704,223,249]" name="India" pageId="123" pageNumber="124">India</collectingCountry>
(
<collectorName id="26E45350FFDCFFDE9CA0CEDD9C754DCA" box="[720,851,223,249]" pageId="123" pageNumber="124">Khasi Hills</collectorName>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFDCFFDE9EB7CF019DAC4C72" pageId="123" pageNumber="124" type="discussion">
<paragraph id="8BAE3686FFDCFFDE9EB7CF019DAC4C72" blockId="123.[151,1437,150,321]" pageId="123" pageNumber="124">
<emphasis id="B965EA94FFDCFFDE9EB7CF019E1D4C2E" bold="true" box="[199,315,259,285]" pageId="123" pageNumber="124">
<collectorName id="26E45350FFDCFFDE9EB7CF019E1D4C2E" box="[199,315,259,285]" pageId="123" pageNumber="124">Remarks.</collectorName>
</emphasis>
<bibRefCitation id="EF804B77FFDCFFDE9F35CF019ECC4C2E" author="Brooks, S. J." box="[325,490,259,285]" pageId="123" pageNumber="124" pagination="35 - 48" refId="ref88537" refString="Brooks, S. J. (1986) A new genus of Ankylopterygini (Chrysopidae). Neuroptera International, 4, 35 - 48." type="journal article" year="1986">Brooks (1986)</bibRefCitation>
established
<taxonomicName id="4C114D05FFDCFFDE9C0ACF019C464C2F" authorityName="Brooks" authorityYear="1986" box="[634,864,259,284]" class="Insecta" family="Chrysopidae" genus="Retipenna" kingdom="Animalia" order="Neuroptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="variegata">
<emphasis id="B965EA94FFDCFFDE9C0ACF019C464C2F" box="[634,864,259,284]" italics="true" pageId="123" pageNumber="124">Retipenna variegata</emphasis>
</taxonomicName>
and provided genital information.
<collectorName id="26E45350FFDCFFDE9A84CF019A004C2E" box="[1268,1318,259,285]" pageId="123" pageNumber="124">This</collectorName>
species is firstly recorded from
<collectingCountry id="F3067616FFDCFFDE9FF7CF259EEC4C72" box="[391,458,295,321]" name="China" pageId="123" pageNumber="124">China</collectingCountry>
and redescribed.
</paragraph>
</subSubSection>
</treatment>
</document>