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<document id="C1D6E8FB0A07287222C748D533FC0417" ID-CLB-Dataset="8896" ID-DOI="10.11646/zootaxa.4868.1.4" ID-GBIF-Dataset="cfd5c06e-06f0-4f1f-ae4b-40b7b46f44a3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4417281" ID-ZooBank="DC4B8EC5-CBDA-49D2-B370-381C96AF3A10" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609805123592" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2020" docId="03A887A1A9721970AFE9FF5F440567FD" docLanguage="en" docName="zootaxa.4868.1.4.pdf" docOrigin="Zootaxa 4868 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus subovatus Perkins 2020, new species" docType="treatment" docVersion="8" lastPageNumber="81" masterDocId="FF91FFD9A9611964AF7EFFC8470B6435" masterDocTitle="Taxonomy of Peruvian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="89" masterPageNumber="61" pageNumber="80" updateTime="1698852338306" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2C5AE0875462FD493E3453526671FD01">Taxonomy of Peruvian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
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<mods:namePart id="B6962AAB7CC9E2BC112A0A393F7BA8BE">Perkins, Philip D.</mods:namePart>
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<mods:date id="514ADA08E9F6D100C8CF000863E7E6E0">2020</mods:date>
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<mods:number id="808C9EC12F9C7CBBC62F0A4659670F20">2020-10-23</mods:number>
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<treatment id="03A887A1A9721970AFE9FF5F440567FD" ID-DOI="http://doi.org/10.5281/zenodo.4417474" ID-GBIF-Taxon="176581831" ID-Zenodo-Dep="4417474" LSID="urn:lsid:plazi:treatment:03A887A1A9721970AFE9FF5F440567FD" httpUri="http://treatment.plazi.org/id/03A887A1A9721970AFE9FF5F440567FD" lastPageId="20" lastPageNumber="81" pageId="19" pageNumber="80">
<subSubSection id="C31B653CA9721977AFE9FF5F45306484" box="[151,571,151,178]" pageId="19" pageNumber="80" type="nomenclature">
<paragraph id="8BBE36B7A9721977AFE9FF5F45306484" blockId="19.[151,571,151,213]" box="[151,571,151,178]" pageId="19" pageNumber="80">
<heading id="D0F681DBA9721977AFE9FF5F45306484" bold="true" box="[151,571,151,178]" fontSize="11" level="1" pageId="19" pageNumber="80" reason="1">
<emphasis id="B975EAA5A9721977AFE9FF5F45306484" bold="true" box="[151,571,151,178]" pageId="19" pageNumber="80">
<taxonomicName id="4C014D34A9721977AFE9FF5F46966487" ID-CoL="8Y4RL" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[151,413,151,178]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="80" phylum="Arthropoda" rank="species" species="subovatus" status="sp. nov.">
<emphasis id="B975EAA5A9721977AFE9FF5F46966487" bold="true" box="[151,413,151,178]" italics="true" pageId="19" pageNumber="80">Hydrochus subovatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24657DEA9721977AED5FF5F45306484" box="[427,571,151,177]" pageId="19" pageNumber="80" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA9721977AFE9FF73460664E0" box="[151,269,187,213]" pageId="19" pageNumber="80" type="description">
<paragraph id="8BBE36B7A9721977AFE9FF73460664E0" blockId="19.[151,571,151,213]" box="[151,269,187,213]" pageId="19" pageNumber="80">
<figureCitation id="133A2A32A9721977AFE9FF7347E864E0" box="[151,227,187,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1971,1995]" captionTargetBox="[157,1430,309,1948]" captionTargetId="figure-37@2.[157,1430,309,1948]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.4417283" httpUri="https://zenodo.org/record/4417283/files/figure.png" pageId="19" pageNumber="80">Figs. 1</figureCitation>
,
<figureCitation id="133A2A32A9721977AF8FFF73460664E0" box="[241,269,187,213]" captionStart="FIGURE 13" captionStartId="19.[151,250,1788,1812]" captionTargetBox="[175,1422,241,1753]" captionTargetId="figure-23@19.[151,1435,222,1772]" captionTargetPageId="19" captionText="FIGURE 13. Hydrochus subovatus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417313" httpUri="https://zenodo.org/record/4417313/files/figure.png" pageId="19" pageNumber="80">13</figureCitation>
</paragraph>
</subSubSection>
<caption id="DF7E663FA9721977AFE9F93444DC6321" ID-DOI="http://doi.org/10.5281/zenodo.4417313" ID-Zenodo-Dep="4417313" box="[151,983,1788,1812]" httpUri="https://zenodo.org/record/4417313/files/figure.png" pageId="19" pageNumber="80" startId="19.[151,250,1788,1812]" targetBox="[175,1422,241,1753]" targetPageId="19">
<paragraph id="8BBE36B7A9721977AFE9F93444DC6321" blockId="19.[151,983,1788,1812]" box="[151,983,1788,1812]" pageId="19" pageNumber="80">
<emphasis id="B975EAA5A9721977AFE9F934462B6321" bold="true" box="[151,288,1788,1812]" pageId="19" pageNumber="80">FIGURE 13.</emphasis>
<taxonomicName id="4C014D34A9721977AE58F934450B6321" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[294,512,1788,1812]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="80" phylum="Arthropoda" rank="species" species="subovatus" status="sp. nov.">
<emphasis id="B975EAA5A9721977AE58F934450B6321" box="[294,512,1788,1812]" italics="true" pageId="19" pageNumber="80">Hydrochus subovatus</emphasis>
</taxonomicName>
<emphasis id="B975EAA5A9721977AD78F93445346321" bold="true" box="[518,575,1788,1812]" pageId="19" pageNumber="80">
<taxonomicNameLabel id="A24657DEA9721977AD78F93445346321" box="[518,575,1788,1812]" pageId="19" pageNumber="80" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, habitus and male genitalia of holotype.
</paragraph>
</caption>
<subSubSection id="C31B653CA9721977AFE9F88B429363B4" pageId="19" pageNumber="80" type="materials_examined">
<paragraph id="8BBE36B7A9721977AFE9F88B429363B4" blockId="19.[151,1436,1858,2029]" pageId="19" pageNumber="80">
<emphasis id="B975EAA5A9721977AFE9F88B46426369" bold="true" box="[151,329,1858,1885]" pageId="19" pageNumber="80">Type Material.</emphasis>
<materialsCitation id="3B693CEAA9721977AE2CF88B448463B4" ID-GBIF-Occurrence="3015416333" collectingDate="1983-09-11" collectionCode="NMNH" collectorName="J. J. Anderson" country="Peru" county="Tambopata" latitude="-13.0" location="Drinking Quebrada" longLatPrecision="1291" longitude="-69.55" municipality="Terrafirme" pageId="19" pageNumber="80" specimenCount="1" stateProvince="Madre de Dios" typeStatus="holotype">
<emphasis id="B975EAA5A9721977AE2CF88B45176369" bold="true" box="[338,540,1858,1885]" pageId="19" pageNumber="80">
<typeStatus id="54BA8815A9721977AE2CF88B46B56368" box="[338,446,1859,1885]" pageId="19" pageNumber="80" type="holotype">Holotype</typeStatus>
(male):
</emphasis>
<collectingCountry id="F3167627A9721977AD51F88B45766368" box="[559,637,1859,1885]" name="Peru" pageId="19" pageNumber="80">PERU</collectingCountry>
:
<collectingRegion id="49C5F855A9721977ADF4F88A443D6369" box="[650,822,1858,1884]" country="Peru" name="Madre de Dios" pageId="19" pageNumber="80">Madre de Dios</collectingRegion>
,
<collectingCounty id="62DF4E3BA9721977AC3AF88A44C96369" box="[836,962,1858,1884]" pageId="19" pageNumber="80">Tambopata</collectingCounty>
,
<collectingMunicipality id="6BDAACCDA9721977ACB0F88A434C6369" box="[974,1095,1858,1884]" pageId="19" pageNumber="80">Terrafirme</collectingMunicipality>
,
<location id="8EDE606CA9721977AB2BF88A42396369" LSID="urn:lsid:plazi:treatment:03A887A1A9721970AFE9FF5F440567FD:8EDE606CA9721977AB2BF88A42396369" box="[1109,1330,1858,1884]" country="Peru" county="Tambopata" latitude="-13.0" longLatPrecision="1291" longitude="-69.55" municipality="Terrafirme" name="Drinking Quebrada" pageId="19" pageNumber="80" stateProvince="Madre de Dios">Drinking Quebrada</location>
, (
<geoCoordinate id="EE355070A9721977AA36F88B42936368" box="[1352,1432,1859,1885]" degrees="13" direction="south" minutes="0" orientation="latitude" pageId="19" pageNumber="80" precision="925" value="-13.0">13°0S</geoCoordinate>
,
<geoCoordinate id="EE355070A9721977AFE9F8AE47F563B4" box="[151,254,1894,1921]" degrees="69" direction="west" minutes="33" orientation="longitude" pageId="19" pageNumber="80" precision="925" value="-69.55">69°33W</geoCoordinate>
) //
<date id="FFBF1077A9721977AE5AF8AE469063B5" box="[292,411,1894,1920]" pageId="19" pageNumber="80" value="1983-09-11">
<collectingDate id="EFFBE99FA9721977AE5AF8AE469063B5" box="[292,411,1894,1920]" pageId="19" pageNumber="80" value="1983-09-11">11.ix.1983</collectingDate>
</date>
, leg.
<collectorName id="26F45361A9721977AEA9F8AF457E63B5" box="[471,629,1894,1921]" pageId="19" pageNumber="80">J. J. Anderson</collectorName>
, ex leaf packs” (
<collectionCode id="ED10AE72A9721977AC52F8AF448C63B4" box="[812,903,1895,1921]" country="USA" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="80" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B693CEAA9721977ACE7F8AF429E63B4" ID-GBIF-Occurrence="3015416331" box="[921,1429,1894,1921]" collectingDate="1983-09-11" collectionCode="NMNH" collectorName="J. J. Anderson" country="Peru" county="Tambopata" latitude="-13.0" location="Drinking Quebrada" longLatPrecision="1291" longitude="-69.55" municipality="Terrafirme" pageId="19" pageNumber="80" specimenCount="1" stateProvince="Madre de Dios" typeStatus="paratype">
<emphasis id="B975EAA5A9721977ACE7F8AF430563B4" bold="true" box="[921,1038,1895,1921]" pageId="19" pageNumber="80">
<typeStatus id="54BA8815A9721977ACE7F8AF430263B4" box="[921,1033,1895,1921]" pageId="19" pageNumber="80" type="paratype">Paratype</typeStatus>
:
</emphasis>
Same data as holotype (1
<collectionCode id="ED10AE72A9721977AA4DF8AF428663B4" box="[1331,1421,1895,1921]" country="USA" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="80" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA9721977AFB9F84247E763D8" pageId="19" pageNumber="80" type="diagnosis">
<paragraph id="8BBE36B7A9721977AFB9F84247E763D8" blockId="19.[151,1436,1858,2029]" pageId="19" pageNumber="80">
<emphasis id="B975EAA5A9721977AFB9F84246D86391" bold="true" box="[199,467,1930,1956]" pageId="19" pageNumber="80">Differential Diagnosis.</emphasis>
Among Peruvian species, recognized by the combination of small size (ca.
<quantity id="4CF99B52A9721977AA56F843429B6390" box="[1320,1424,1931,1957]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="19" pageNumber="80" unit="mm" value="2.3">2.30 mm</quantity>
), the sub-oval body form, the elytra with odd-numbered interstriae raised, and the male genitalia (
<figureCitation id="133A2A32A9721977ABAFF866422E63FC" box="[1233,1317,1966,1993]" captionStart="FIGURE 13" captionStartId="19.[151,250,1788,1812]" captionTargetBox="[175,1422,241,1753]" captionTargetId="figure-23@19.[151,1435,222,1772]" captionTargetPageId="19" captionText="FIGURE 13. Hydrochus subovatus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417313" httpUri="https://zenodo.org/record/4417313/files/figure.png" pageId="19" pageNumber="80">Fig. 13</figureCitation>
; described below).
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA9751970AFB9FF5E45476791" pageId="20" pageNumber="81" type="description">
<paragraph id="8BBE36B7A9751970AFB9FF5E458D6551" blockId="20.[151,1437,150,969]" pageId="20" pageNumber="81">
<emphasis id="B975EAA5A9751970AFB9FF5E465A6485" bold="true" box="[199,337,150,176]" pageId="20" pageNumber="81">Description</emphasis>
. Habitus as illustrated (
<figureCitation id="133A2A32A9751970AD27FF5E45A76485" box="[601,684,150,176]" captionStart="FIGURE 13" captionStartId="19.[151,250,1788,1812]" captionTargetBox="[175,1422,241,1753]" captionTargetId="figure-23@19.[151,1435,222,1772]" captionTargetPageId="19" captionText="FIGURE 13. Hydrochus subovatus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417313" httpUri="https://zenodo.org/record/4417313/files/figure.png" pageId="20" pageNumber="81">Fig. 13</figureCitation>
). Size:
<typeStatus id="54BA8815A9751970AD80FF5E446B6485" box="[766,864,150,176]" pageId="20" pageNumber="81" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 2.30/0.93; head width 0.58; pronotum l/w 0.53/0.60; PA 0.55; PB 0.47; elytra 1.43/0.93. Body size moderate (ca.
<quantity id="4CF99B52A9751970AFE9FF16460B64CC" box="[151,256,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="20" pageNumber="81" unit="mm" value="2.3">2.30 mm</quantity>
). Dorsum dark brown, with slight iridescence, elytra without black spots. Legs brown, with tibiofemoral joints and basal ½ of femora darker. Punctation on elytra ca. 23x that of pronotum. Elytra interstriae ca. 0.51x strial puncture diameter. Interstria 9
<superScript id="7C749BFFA9751970AD54FEEE453C6501" attach="left" box="[554,567,294,308]" fontSize="6" pageId="20" pageNumber="81">th</superScript>
more convex than others, overhanging 10
<superScript id="7C749BFFA9751970AB6DFEEE432B6501" attach="left" box="[1043,1056,294,308]" fontSize="6" pageId="20" pageNumber="81">th</superScript>
interstria. Usual area of callus on 5
<superScript id="7C749BFFA9751970AFDBFE8247B9656D" attach="left" box="[165,178,330,344]" fontSize="6" pageId="20" pageNumber="81">th</superScript>
very weakly raised, moderately elongate.
</paragraph>
<paragraph id="8BBE36B7A9751970AFB9FEA743BC65E5" blockId="20.[151,1437,150,969]" pageId="20" pageNumber="81">Pronotum slightly wider than long, widest slightly behind anterior margin, narrowed at base, sides arcuate or very weakly sinuate, denticulate, each process with moderately long arcuate seta; anterior margin bisinuate; depressions, especially anterior, deep, much more densely punctate than reliefs between depressions.</paragraph>
<paragraph id="8BBE36B7A9751970AFB9FE1246516609" blockId="20.[151,1437,150,969]" pageId="20" pageNumber="81">Elytra with odd numbered interstriae raised, spaces between strial punctures ca. 0.51x their diameter; apices rather sharply conjointly rounded in dorsal view, with sublateral punctate groove, in lateral view outer margin with slight angulation.</paragraph>
<paragraph id="8BBE36B7A9751970AFB9FD8E47D966B0" blockId="20.[151,1437,150,969]" pageId="20" pageNumber="81">Ventral characters: Mentum moderately densely punctate, with median fovea. Submentum with two deep foveae.</paragraph>
<paragraph id="8BBE36B7A9751970AFB9FD46430A670D" blockId="20.[151,1437,150,969]" pageId="20" pageNumber="81">
Male genitalia (
<figureCitation id="133A2A32A9751970AE0BFD4646CF669D" box="[373,452,654,680]" captionStart="FIGURE 13" captionStartId="19.[151,250,1788,1812]" captionTargetBox="[175,1422,241,1753]" captionTargetId="figure-23@19.[151,1435,222,1772]" captionTargetPageId="19" captionText="FIGURE 13. Hydrochus subovatus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417313" httpUri="https://zenodo.org/record/4417313/files/figure.png" pageId="20" pageNumber="81">Fig. 13</figureCitation>
) general characters: genitalia very small, very slender, length ca. 6x width; paramere distal ends sharply rounded in dorsal/ventral and lateral views, slightly longer than aedeagus, lacking plsa, tips beyond aedeagus with medial margins parallel or nearly so, distal ½ of parameres with distinct pmsm; aedeagus narrowly lanceolate, agw slightly distal to midlength, alw just distal to adbl; basal piece slightly longer than parameres, width slightly greater in dorsal/ventral than lateral views, orifice weakly sclerotized.
</paragraph>
<paragraph id="8BBE36B7A9751970AFB9FC8A42186769" blockId="20.[151,1437,150,969]" box="[199,1299,834,861]" pageId="20" pageNumber="81">Dorsal surface: adbl moderately large, almost as wide as agw; pdmm approximating shape of alm.</paragraph>
<paragraph id="8BBE36B7A9751970AFB9FCAE45476791" blockId="20.[151,1437,150,969]" pageId="20" pageNumber="81">Ventral surface: pvmm basally overlapping aedeagus, margins very narrowly separated one from the other, otherwise approximating shape of alm.</paragraph>
</subSubSection>
<subSubSection id="C31B653CA9751970AFB9FC66440567FD" box="[199,782,942,969]" pageId="20" pageNumber="81" type="etymology">
<paragraph id="8BBE36B7A9751970AFB9FC66440567FD" blockId="20.[151,1437,150,969]" box="[199,782,942,969]" pageId="20" pageNumber="81">
<emphasis id="B975EAA5A9751970AFB9FC66464567FD" bold="true" box="[199,334,942,968]" pageId="20" pageNumber="81">Etymology.</emphasis>
Named in reference to the oval habitus.
</paragraph>
</subSubSection>
</treatment>
</document>