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<document id="EEDBBD8152AC63AEF44D62D689A742F7" ID-CLB-Dataset="8896" ID-DOI="10.11646/zootaxa.4868.1.4" ID-GBIF-Dataset="cfd5c06e-06f0-4f1f-ae4b-40b7b46f44a3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4417281" ID-ZooBank="DC4B8EC5-CBDA-49D2-B370-381C96AF3A10" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609805123592" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2020" docId="03A887A1A969196EAFE9FAF244616528" docLanguage="en" docName="zootaxa.4868.1.4.pdf" docOrigin="Zootaxa 4868 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus bucapitus Perkins 2020, new species" docType="treatment" docVersion="8" lastPageNumber="71" masterDocId="FF91FFD9A9611964AF7EFFC8470B6435" masterDocTitle="Taxonomy of Peruvian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="89" masterPageNumber="61" pageNumber="69" updateTime="1698852338306" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="310C98E5D4E42C4C4ABF312D9B79C099">Taxonomy of Peruvian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
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<mods:namePart id="9F85153E68D202E73EB2D7B39F637D35">Perkins, Philip D.</mods:namePart>
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<mods:date id="CC7034B50A1F5BAD74577907FD17B1B3">2020</mods:date>
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<mods:number id="408EDB2FCE79AFA503307D39EEDF16F6">2020-10-23</mods:number>
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<treatment id="03A887A1A969196EAFE9FAF244616528" ID-DOI="http://doi.org/10.5281/zenodo.4417464" ID-GBIF-Taxon="176581838" ID-Zenodo-Dep="4417464" LSID="urn:lsid:plazi:treatment:03A887A1A969196EAFE9FAF244616528" httpUri="http://treatment.plazi.org/id/03A887A1A969196EAFE9FAF244616528" lastPageId="10" lastPageNumber="71" pageId="8" pageNumber="69">
<subSubSection id="C31B653CA969196CAFE9FAF245336161" box="[151,568,1338,1365]" pageId="8" pageNumber="69" type="nomenclature">
<paragraph id="8BBE36B7A969196CAFE9FAF245336161" blockId="8.[151,568,1338,1400]" box="[151,568,1338,1365]" pageId="8" pageNumber="69">
<heading id="D0F681DBA969196CAFE9FAF245336161" bold="true" box="[151,568,1338,1365]" fontSize="11" level="1" pageId="8" pageNumber="69" reason="1">
<emphasis id="B975EAA5A969196CAFE9FAF245336161" bold="true" box="[151,568,1338,1365]" pageId="8" pageNumber="69">
<taxonomicName id="4C014D34A969196CAFE9FAF246926160" ID-CoL="8Y4PH" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[151,409,1338,1365]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="species" species="bucapitus" status="sp. nov.">
<emphasis id="B975EAA5A969196CAFE9FAF246926160" bold="true" box="[151,409,1338,1365]" italics="true" pageId="8" pageNumber="69">Hydrochus bucapitus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24657DEA969196CAED6FAF245336161" box="[424,568,1338,1364]" pageId="8" pageNumber="69" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C31B653CA969196CAFE9FA9647F4614D" box="[151,255,1374,1400]" pageId="8" pageNumber="69" type="description">
<paragraph id="8BBE36B7A969196CAFE9FA9647F4614D" blockId="8.[151,568,1338,1400]" box="[151,255,1374,1400]" pageId="8" pageNumber="69">
<figureCitation id="133A2A32A969196CAFE9FA9647E8614D" box="[151,227,1374,1400]" captionStart="FIGURE 1" captionStartId="2.[151,250,1971,1995]" captionTargetBox="[157,1430,309,1948]" captionTargetId="figure-37@2.[157,1430,309,1948]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.4417283" httpUri="https://zenodo.org/record/4417283/files/figure.png" pageId="8" pageNumber="69">Figs. 1</figureCitation>
,
<figureCitation id="133A2A32A969196CAF8FFA9647F4614D" box="[241,255,1374,1400]" captionStart="FIGURE 6" captionStartId="9.[151,250,1979,2003]" captionTargetBox="[170,1426,470,1941]" captionTargetId="figure-121@9.[151,1435,453,1956]" captionTargetPageId="9" captionText="FIGURE 6. Hydrochus bucapitus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417295" httpUri="https://zenodo.org/record/4417295/files/figure.png" pageId="8" pageNumber="69">6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA969196CAFE9FA6E4686623C" pageId="8" pageNumber="69" type="materials_examined">
<paragraph id="8BBE36B7A969196CAFE9FA6E4686623C" blockId="8.[151,1437,1446,2013]" pageId="8" pageNumber="69">
<emphasis id="B975EAA5A969196CAFE9FA6E464161F5" bold="true" box="[151,330,1446,1472]" pageId="8" pageNumber="69">Type Material.</emphasis>
<materialsCitation id="3B693CEAA969196CAE2DFA6E44CE61D1" ID-GBIF-Occurrence="3015416368" collectingDate="1979-11-26" collectingDateMax="1979-11-30" collectingDateMin="1979-11-26" collectionCode="NMNH" collectorName="J. B. Heppner" country="Peru" elevation="290" location="Rio Tambopata Res" pageId="8" pageNumber="69" specimenCount="1" stateProvince="Madre de Dios" typeStatus="holotype">
<emphasis id="B975EAA5A969196CAE2DFA6E451561F5" bold="true" box="[339,542,1446,1472]" pageId="8" pageNumber="69">
<typeStatus id="54BA8815A969196CAE2DFA6E46CB61F5" box="[339,448,1446,1472]" pageId="8" pageNumber="69" type="holotype">Holotype</typeStatus>
(male):
</emphasis>
<collectingCountry id="F3167627A969196CAD4FFA6E457461F5" box="[561,639,1446,1472]" name="Peru" pageId="8" pageNumber="69">PERU</collectingCountry>
:
<collectingRegion id="49C5F855A969196CADF5FA6E443261F5" box="[651,825,1446,1472]" country="Peru" name="Madre de Dios" pageId="8" pageNumber="69">Madre de Dios</collectingRegion>
;
<location id="8EDE606CA969196CAC38FA6E432261F4" LSID="urn:lsid:plazi:treatment:03A887A1A969196EAFE9FAF244616528:8EDE606CA969196CAC38FA6E432261F4" box="[838,1065,1446,1473]" country="Peru" name="Rio Tambopata Res" pageId="8" pageNumber="69" stateProvince="Madre de Dios">Rio Tambopata Res</location>
; 30 air km SW Pto. Maldonado,
<quantity id="4CF99B52A969196CAFE9FA0247EB61D0" box="[151,224,1482,1509]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="8" pageNumber="69" unit="m" value="290.0">
<elevation id="002CD184A969196CAFE9FA0247EB61D0" box="[151,224,1482,1509]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="8" pageNumber="69" unit="m" value="290.0">290 m</elevation>
</quantity>
,
<date id="FFBF1077A969196CAF95FA02468461D1" box="[235,399,1482,1508]" pageId="8" pageNumber="69" value="1979-11-26" valueMax="1979-11-30" valueMin="1979-11-26">
<collectingDate id="EFFBE99FA969196CAF95FA02468461D1" box="[235,399,1482,1508]" pageId="8" pageNumber="69" value="1979-11-26" valueMax="1979-11-30" valueMin="1979-11-26">2630 xi 1979</collectingDate>
</date>
,
<collectorName id="26F45361A969196CAEE7FA03453F61D0" box="[409,564,1483,1509]" pageId="8" pageNumber="69">J. B. Heppner</collectorName>
, subtropical moist forest” (
<collectionCode id="ED10AE72A969196CAC1FFA0244B661D1" box="[865,957,1482,1508]" country="USA" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="69" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B693CEAA969196CACB1FA024682623C" ID-GBIF-Occurrence="3015416374" collectingDate="1979-11-05" collectingDateMax="1979-11-30" collectingDateMin="1979-11-26" collectionCode="NMNH" collectorName="J. B. Heppner" country="Peru" elevation="290" location="Rio Tambopata Res" pageId="8" pageNumber="69" specimenCount="1" stateProvince="Madre de Dios" typeStatus="paratype">
<emphasis id="B975EAA5A969196CACB1FA02434E61D1" bold="true" box="[975,1093,1482,1508]" pageId="8" pageNumber="69">
<typeStatus id="54BA8815A969196CACB1FA02434B61D1" box="[975,1088,1482,1508]" pageId="8" pageNumber="69" type="paratype">Paratype</typeStatus>
:
</emphasis>
Same data as holotype, except
<date id="FFBF1077A969196CAFE9FA26460A623D" box="[151,257,1518,1545]" pageId="8" pageNumber="69" value="1979-11-05">
<collectingDate id="EFFBE99FA969196CAFE9FA26460A623D" box="[151,257,1518,1545]" pageId="8" pageNumber="69" value="1979-11-05">5 xi 1979</collectingDate>
</date>
(1
<collectionCode id="ED10AE72A969196CAE58FA27468A623C" box="[294,385,1519,1545]" country="USA" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="69" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA969196CAFB9F9DA43B06241" pageId="8" pageNumber="69" type="diagnosis">
<paragraph id="8BBE36B7A969196CAFB9F9DA43B06241" blockId="8.[151,1437,1446,2013]" pageId="8" pageNumber="69">
<emphasis id="B975EAA5A969196CAFB9F9DA46DF6219" bold="true" box="[199,468,1554,1580]" pageId="8" pageNumber="69">Differential Diagnosis.</emphasis>
Differentiated from other Peruvian species by the combination of the moderate body size (ca.
<quantity id="4CF99B52A969196CAF82F9FE466E6264" box="[252,357,1590,1617]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="69" unit="mm" value="2.5">2.50 mm</quantity>
), the rather large head (width greater than width or length of pronotum), and the male genitalia (
<figureCitation id="133A2A32A969196CAFDEF99247EF6241" box="[160,228,1626,1652]" captionStart="FIGURE 6" captionStartId="9.[151,250,1979,2003]" captionTargetBox="[170,1426,470,1941]" captionTargetId="figure-121@9.[151,1435,453,1956]" captionTargetPageId="9" captionText="FIGURE 6. Hydrochus bucapitus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417295" httpUri="https://zenodo.org/record/4417295/files/figure.png" pageId="8" pageNumber="69">Fig. 6</figureCitation>
; described below). Completely reliable determinations will include dissection of males.
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA969196EAFB9F9B647DC64CC" lastPageId="10" lastPageNumber="71" pageId="8" pageNumber="69" type="description">
<paragraph id="8BBE36B7A969196CAFB9F9B647E96345" blockId="8.[151,1437,1446,2013]" pageId="8" pageNumber="69">
<emphasis id="B975EAA5A969196CAFB9F9B6465A62AD" bold="true" box="[199,337,1662,1688]" pageId="8" pageNumber="69">Description</emphasis>
. Habitus as illustrated (
<figureCitation id="133A2A32A969196CAD23F9B645A962AC" box="[605,674,1662,1689]" captionStart="FIGURE 6" captionStartId="9.[151,250,1979,2003]" captionTargetBox="[170,1426,470,1941]" captionTargetId="figure-121@9.[151,1435,453,1956]" captionTargetPageId="9" captionText="FIGURE 6. Hydrochus bucapitus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417295" httpUri="https://zenodo.org/record/4417295/files/figure.png" pageId="8" pageNumber="69">Fig. 6</figureCitation>
). Size:
<typeStatus id="54BA8815A969196CAD89F9B6445262AD" box="[759,857,1662,1688]" pageId="8" pageNumber="69" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 2.50/0.83; head width 0.62; pronotum l/w 0.58/0.55; PA 0.55; PB 0.42; elytra 1.60/0.83. Body size moderate (ca.
<quantity id="4CF99B52A969196CAFE9F90E460962D4" box="[151,258,1734,1761]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="69" unit="mm" value="2.5">2.50 mm</quantity>
), very narrow, except head relatively wide. Dorsum dark brown with slight iridescence, elytra without black spots. Dorsum with numerous microsetae. Legs testaceous, with tibiofemoral joints and basal ½ of femora darker. Punctation on elytra ca. 24x that of pronotum. Elytra interstriae narrow walls to ca. 0.5x strial puncture diameter. Interstria 9
<superScript id="7C749BFFA969196CAEFEF8FA46866375" attach="left" box="[384,397,1842,1856]" fontSize="6" pageId="8" pageNumber="69">th</superScript>
more convex than others, overhanging 10
<superScript id="7C749BFFA969196CAC1AF8FA447A6375" attach="left" box="[868,881,1842,1856]" fontSize="6" pageId="8" pageNumber="69">th</superScript>
interstria. Usual area of callus on 5
<superScript id="7C749BFFA969196CAB81F8FA42076375" attach="left" box="[1279,1292,1842,1856]" fontSize="6" pageId="8" pageNumber="69">th</superScript>
very weakly raised.
</paragraph>
<paragraph id="8BBE36B7A969196CAFB9F8B3456163E8" blockId="8.[151,1437,1446,2013]" pageId="8" pageNumber="69">Pronotum very slightly longer than wide, widest at anterior margin, narrowed at base, sides very weakly arcuate, almost straight, smooth; anterior margin arcuate; depressions moderately deep, coarsely densely punctate, reliefs between depressions less punctate.</paragraph>
<paragraph id="8BBE36B7A968196DAFB9FF5F430664E0" blockId="9.[151,1436,150,429]" pageId="9" pageNumber="70">Elytra with spaces between strial punctures narrow walls to ca. 0.5x their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin without angulation.</paragraph>
<paragraph id="8BBE36B7A968196DAFB9FF1746C56528" blockId="9.[151,1436,150,429]" pageId="9" pageNumber="70">Ventral characters: Mentum moderately densely finely punctate, with large, deep median fovea. Submentum with two large deep foveae.</paragraph>
<paragraph id="8BBE36B7A968196DAFB9FEEF44DE6598" blockId="9.[151,1436,150,429]" pageId="9" pageNumber="70">
Male genitalia (
<figureCitation id="133A2A32A968196DAE02FEEF46CA6577" box="[380,449,295,322]" captionStart="FIGURE 6" captionStartId="9.[151,250,1979,2003]" captionTargetBox="[170,1426,470,1941]" captionTargetId="figure-121@9.[151,1435,453,1956]" captionTargetPageId="9" captionText="FIGURE 6. Hydrochus bucapitus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417295" httpUri="https://zenodo.org/record/4417295/files/figure.png" pageId="9" pageNumber="70">Fig. 6</figureCitation>
) general characters: genitalia very small, very slender, length ca. 6x width; parameres pointed in dorsal/ventral views and lateral view, only slightly longer than aedeagus, lacking plsa; aedeagus narrowly lanceolate, agw at about midlength; basal piece slightly shorter than parameres, width ca. equal in dorsal/ventral and lateral views, orifice sclerotized, inverted V-shape in dorsal/ventral views.
</paragraph>
<caption id="DF7E663FA968196DAFE9F87344C363E6" ID-DOI="http://doi.org/10.5281/zenodo.4417295" ID-Zenodo-Dep="4417295" box="[151,968,1979,2003]" httpUri="https://zenodo.org/record/4417295/files/figure.png" pageId="9" pageNumber="70" startId="9.[151,250,1979,2003]" targetBox="[170,1426,470,1941]" targetPageId="9">
<paragraph id="8BBE36B7A968196DAFE9F87344C363E6" blockId="9.[151,968,1979,2003]" box="[151,968,1979,2003]" pageId="9" pageNumber="70">
<emphasis id="B975EAA5A968196DAFE9F873461863E6" bold="true" box="[151,275,1979,2003]" pageId="9" pageNumber="70">FIGURE 6.</emphasis>
<taxonomicName id="4C014D34A968196DAE67F87346FA63E6" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[281,497,1979,2003]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="species" species="bucapitus" status="sp. nov.">
<emphasis id="B975EAA5A968196DAE67F87346FA63E6" box="[281,497,1979,2003]" italics="true" pageId="9" pageNumber="70">Hydrochus bucapitus</emphasis>
</taxonomicName>
<emphasis id="B975EAA5A968196DAE89F873453B63E6" bold="true" box="[503,560,1979,2003]" pageId="9" pageNumber="70">
<taxonomicNameLabel id="A24657DEA968196DAE89F873453B63E6" box="[503,560,1979,2003]" pageId="9" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, habitus and male genitalia of holotype.
</paragraph>
</caption>
<paragraph id="8BBE36B7A96B196EAFB9FF5F442B6484" blockId="10.[151,1436,151,285]" box="[199,800,151,177]" pageId="10" pageNumber="71">Dorsal surface: adbl large; pdmm approximating alm.</paragraph>
<paragraph id="8BBE36B7A96B196EAFB9FF7347DC64CC" blockId="10.[151,1436,151,285]" pageId="10" pageNumber="71">Ventral surface: pvmm over middle ½ overlapping aedeagus, margins very narrowly separated one from the other.</paragraph>
</subSubSection>
<subSubSection id="C31B653CA96B196EAFB9FECB44616528" box="[199,874,259,285]" pageId="10" pageNumber="71" type="etymology">
<paragraph id="8BBE36B7A96B196EAFB9FECB44616528" blockId="10.[151,1436,151,285]" box="[199,874,259,285]" pageId="10" pageNumber="71">
<emphasis id="B975EAA5A96B196EAFB9FECB46456528" bold="true" box="[199,334,259,285]" pageId="10" pageNumber="71">Etymology.</emphasis>
Named in reference to the relatively large head.
</paragraph>
</subSubSection>
</treatment>
</document>