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<document id="45D0CF1DE0F68C9E5642D6495D05304A" ID="10.11646/zootaxa.4139.3.2" ID-DOI="10.11646/zootaxa.4139.3.2" ID-GBIF-Dataset="51358935-98b0-4d89-a47f-e4341171e5b9" ID-ISSN="1175-5326" ID-Zenodo-Dep="262765" ID-ZooBank="DCF4220B-4A73-407C-837C-54DEAE29F435" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1469091585085" checkinUser="plazi" docAuthor="Last, Peter R., Naylor, Gavin J. P. &amp; Manjaji-Matsumoto, B. Mabel" docDate="2016" docId="03A7CA1D8565145FE7C6FD37FBC602DC" docLanguage="en" docName="zootaxa.4139.3.2.pdf" docOrigin="Zootaxa 4139 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fluvitrygon Last, Naylor &amp; Manjaji-Matsumoto, 2016, gen. nov." docType="treatment" docVersion="9" lastPageNumber="360" masterDocId="FF9EB265856A1450E751FF83FFDE0766" masterDocTitle="A revised classification of the family Dasyatidae (Chondrichthyes: Myliobatiformes) based on new morphological and molecular insights" masterLastPageNumber="368" masterPageNumber="345" pageNumber="360" updateTime="1698365466156" updateUser="plazi">
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<mods:title id="BEBF110EF3CCE8691A29819A467CC838">A revised classification of the family Dasyatidae (Chondrichthyes: Myliobatiformes) based on new morphological and molecular insights</mods:title>
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<mods:namePart id="DB0B9E1F6D430629A0455B9C8770846B">Last, Peter R.</mods:namePart>
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<mods:namePart id="C42B08FD44A7F66EB0F81EBBC5E44F05">Naylor, Gavin J. P.</mods:namePart>
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<subSubSection id="C31428808565145FE7C6FD37FE2C05A8" box="[151,498,692,718]" pageId="15" pageNumber="360" type="nomenclature">
<paragraph id="8BB17B0B8565145FE7C6FD37FE2C05A8" blockId="15.[151,498,692,718]" box="[151,498,692,718]" pageId="15" pageNumber="360">
<heading id="D0F9CC678565145FE7C6FD37FE2C05A8" bold="true" box="[151,498,692,718]" fontSize="11" level="1" pageId="15" pageNumber="360" reason="1">
<emphasis id="B97AA7198565145FE7C6FD37FE2C05A8" bold="true" box="[151,498,692,718]" pageId="15" pageNumber="360">
Genus
<taxonomicName id="4C0E00888565145FE7BCFD37FEA205A8" ID-CoL="6352R" ID-ENA="2338057" box="[237,380,692,718]" class="Elasmobranchii" family="Dasyatidae" genus="Fluvitrygon" kingdom="Animalia" order="Myliobatiformes" pageId="15" pageNumber="360" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B97AA7198565145FE7BCFD37FEA205A8" bold="true" box="[237,380,692,718]" italics="true" pageId="15" pageNumber="360">Fluvitrygon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2491A628565145FE6D5FD37FE2C05A8" box="[388,498,692,718]" pageId="15" pageNumber="360" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C31428808565145FE7C6FD78FC4503D8" pageId="15" pageNumber="360" type="materials_examined">
<paragraph id="8BB17B0B8565145FE7C6FD78FC650476" blockId="15.[151,955,762,784]" box="[151,955,762,784]" pageId="15" pageNumber="360">
<typeStatus id="54B5C5A98565145FE7C6FD78FF170476" box="[151,201,763,784]" pageId="15" pageNumber="360">Type</typeStatus>
species
<taxonomicName id="4C0E00888565145FE64EFD78FCD60476" authority="Compagno &amp; Roberts, 1982" authorityName="Compagno &amp; Roberts" authorityYear="1982" box="[287,776,762,784]" class="Elasmobranchii" family="Dasyatidae" genus="Himantura" kingdom="Animalia" order="Myliobatiformes" pageId="15" pageNumber="360" phylum="Chordata" rank="species" species="signifer">
<emphasis id="B97AA7198565145FE64EFD78FE3F0476" box="[287,481,762,784]" italics="true" pageId="15" pageNumber="360">Himantura signifer</emphasis>
<bibRefCitation id="EF9F06FA8565145FE6B9FD79FCD60476" author="Compagno" box="[488,776,762,784]" pageId="15" pageNumber="360" refString="Compagno, L. J. V. &amp; Roberts, T. R. (1982) Freshwater stingrays (Dasyatidae) of southeast Asia and New Guinea, with description of a new species of Himantura and reports of unidentified species. Environmental Biology of Fishes, 7 (4), 321 - 339. http: // dx. doi. org / 10.1007 / BF 00005567" type="journal article" year="1982">Compagno &amp; Roberts, 1982</bibRefCitation>
</taxonomicName>
; newly proposed.
</paragraph>
<paragraph id="8BB17B0B8565145FE7C6FCBFFC4503D8" blockId="15.[151,1437,828,1466]" pageId="15" pageNumber="360">
<emphasis id="B97AA7198565145FE7C6FCBFFED10433" bold="true" box="[151,271,828,853]" pageId="15" pageNumber="360">Definition</emphasis>
. Small dasyatids (adults typically to ~
<quantity id="4CF6D6EE8565145FE59DFCBDFCCA0433" box="[716,788,830,854]" metricMagnitude="-1" metricUnit="m" metricValue="3.7" pageId="15" pageNumber="360" unit="cm" value="37.0">37 cm</quantity>
DW) characterised by the following: depressed, oval to almost circular disc with pectoral-fin apex broadly rounded; snout acutely angular to obtuse, short or long (2.23.7 times combined orbit and spiracle length); eye very small or small and protruding slightly; nasal curtain broadly skirt shaped; mouth narrow to medium-sized with 28 oral papillae; tail rather short or long, whip-like (length 1.3 3.8 times DW), its base narrow and oval to almost circular in cross section; pelvic fins medium-sized, almost entirely concealed by disc; dorsal fold and ventral folds absent; caudal sting rather forward on tail (distance from pectoral-fin insertion to caudal-sting base 2.33.2 times interspiracular width); 02 pearl-shaped mid-scapular thorns, no other scapular thorns; denticle band well developed with edge diffuse to sharply defined, skin on rest of disc typically naked; median thorns on tail weak or absent, small denticles usually on posterior tail in adults; dorsal coloration plain or with strong pattern; ventral surface white, disc margin sometimes dark edged; posterior tail plain or with pattern; estuarine/freshwater, IndoMalay Archipelago.
</paragraph>
</subSubSection>
<subSubSection id="C31428808565145FE796FB4AFE330228" pageId="15" pageNumber="360" type="etymology">
<paragraph id="8BB17B0B8565145FE796FB4AFD800260" blockId="15.[151,1437,828,1466]" pageId="15" pageNumber="360">
<emphasis id="B97AA7198565145FE796FB4AFE980387" bold="true" box="[199,326,1225,1249]" pageId="15" pageNumber="360">Etymology</emphasis>
. Combination of the Latin
<emphasis id="B97AA7198565145FE5F0FB4AFD320384" box="[673,748,1225,1250]" italics="true" pageId="15" pageNumber="360">fluvius</emphasis>
(river) and Greek
<taxonomicName id="4C0E00888565145FE48CFB49FBF90387" box="[989,1063,1226,1249]" class="Elasmobranchii" family="Dasyatidae" genus="Trygon" higherTaxonomySource="CoL" kingdom="Animalia" order="Myliobatiformes" pageId="15" pageNumber="362" phylum="Chordata" rank="genus">
<emphasis id="B97AA7198565145FE48CFB49FBF90387" box="[989,1063,1226,1249]" italics="true" pageId="15" pageNumber="360">trygon</emphasis>
</taxonomicName>
(stingray) reflects the riverine distribution of all members of the genus.
</paragraph>
<paragraph id="8BB17B0B8565145FE796FA92FE330228" blockId="15.[151,1437,828,1466]" pageId="15" pageNumber="360">
<emphasis id="B97AA7198565145FE796FA92FEC0024C" bold="true" box="[199,286,1297,1322]" pageId="15" pageNumber="360">Species</emphasis>
.
<taxonomicName id="4C0E00888565145FE665FA91FC91024C" authority="Vidthayanon &amp; Roberts, 2006" authorityName="Vidthayanon &amp; Roberts" authorityYear="2006" box="[308,847,1297,1322]" class="Elasmobranchii" family="Dasyatidae" genus="Fluvitrygon" kingdom="Animalia" order="Myliobatiformes" pageId="15" pageNumber="360" phylum="Chordata" rank="species" species="kittipongi">
<emphasis id="B97AA7198565145FE665FA91FE1C024C" box="[308,450,1297,1322]" italics="true" pageId="15" pageNumber="360">F. kittipongi</emphasis>
(Vidthayanon &amp; Roberts, 2006)
</taxonomicName>
,
<taxonomicName id="4C0E00888565145FE433FA91FB09024C" authority="Sauvage, 1878" authorityName="Sauvage" authorityYear="1878" box="[866,1239,1297,1322]" class="Elasmobranchii" family="Dasyatidae" genus="Fluvitrygon" kingdom="Animalia" order="Myliobatiformes" pageId="15" pageNumber="360" phylum="Chordata" rank="species" species="oxyrhyncha">
<emphasis id="B97AA7198565145FE433FA91FBD9024C" box="[866,1031,1297,1322]" italics="true" pageId="15" pageNumber="360">F. oxyrhyncha</emphasis>
(Sauvage, 1878)
</taxonomicName>
, and
<taxonomicName id="4C0E00888565145FE272FA91FE360228" authority="Compagno &amp; Roberts, 1982" authorityName="Compagno &amp; Roberts" authorityYear="1982" class="Elasmobranchii" family="Dasyatidae" genus="Himantura" kingdom="Animalia" order="Myliobatiformes" pageId="15" pageNumber="360" phylum="Chordata" rank="species" species="signifer">
<emphasis id="B97AA7198565145FE272FA91FA45024C" box="[1315,1435,1298,1322]" italics="true" pageId="15" pageNumber="360">F. signifer</emphasis>
(
<bibRefCitation id="EF9F06FA8565145FE7CFFAB5FE3E0228" author="Compagno" box="[158,480,1333,1358]" pageId="15" pageNumber="360" refString="Compagno, L. J. V. &amp; Roberts, T. R. (1982) Freshwater stingrays (Dasyatidae) of southeast Asia and New Guinea, with description of a new species of Himantura and reports of unidentified species. Environmental Biology of Fishes, 7 (4), 321 - 339. http: // dx. doi. org / 10.1007 / BF 00005567" type="journal article" year="1982">Compagno &amp; Roberts, 1982</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C31428808565145FE796FADAFBC602DC" pageId="15" pageNumber="360" type="discussion">
<paragraph id="8BB17B0B8565145FE796FADAFBC602DC" blockId="15.[151,1437,828,1466]" pageId="15" pageNumber="360">
<emphasis id="B97AA7198565145FE796FADAFEEA0217" bold="true" box="[199,308,1369,1393]" pageId="15" pageNumber="360">Remarks</emphasis>
. Newly erected, morphologically similar genus consisting of three small species previously placed in
<taxonomicName id="4C0E00888565145FE7C6FAFDFECC02F3" box="[151,274,1406,1429]" class="Elasmobranchii" family="Dasyatidae" genus="Himantura" kingdom="Animalia" order="Myliobatiformes" pageId="15" pageNumber="360" phylum="Chordata" rank="genus">
<emphasis id="B97AA7198565145FE7C6FAFDFECC02F3" box="[151,274,1406,1429]" italics="true" pageId="15" pageNumber="360">Himantura</emphasis>
</taxonomicName>
. These freshwater/estuarine species are sister to
<taxonomicName id="4C0E00888565145FE463FAFDFC6902F3" box="[818,951,1406,1429]" class="Elasmobranchii" family="Dasyatidae" genus="Brevitrygon" kingdom="Animalia" order="Myliobatiformes" pageId="15" pageNumber="359" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B97AA7198565145FE463FAFDFC6902F3" box="[818,951,1406,1429]" italics="true" pageId="15" pageNumber="360">Brevitrygon</emphasis>
</taxonomicName>
(
<figureCitation id="1335678E8565145FE497FAFEFBD702F0" box="[966,1033,1405,1430]" captionStart="FIGURE 3" captionStartId="5.[151,250,1873,1895]" captionText="FIGURE 3. Dasyatidae (subfamily Urogymninae). Topology and branch lengths resulting from a constrained Maximum" pageId="15" pageNumber="360">Fig. 3</figureCitation>
), but differ in tail morphology from members of that marine group (having a longer, narrower-based, whip-like tail).
</paragraph>
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</treatment>
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