193 lines
20 KiB
XML
193 lines
20 KiB
XML
<document id="065AB808FA68BDE83F63DDF933C8F364" ID-DOI="10.5281/zenodo.5340370" ID-ISSN="2345-7600" ID-Zenodo-Dep="5340370" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630186249002" checkinUser="diego" docAuthor="Dan, Polhemus, John, Polhemus & Sites, Robert" docDate="2008" docId="039F87C7E57BFFCCFC5BFEAB32C2FC2F" docLanguage="en" docName="56rbz255-279.pdf" docOrigin="Raffles Bulletin of Zoology 56 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Cheirochela assamensis Hope 1841" docType="treatment" docVersion="4" lastPageNumber="260" masterDocId="FFA6FFBFE57FFFC9FFFAFF893329FFF6" masterDocTitle="A Revision Of The Indochinese Genera Cheirochela And Gestroiella Heteroptera Naucoridae And A Review Of The Tribe Cheirochelini" masterLastPageNumber="279" masterPageNumber="255" pageNumber="259" updateTime="1699179848367" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1DB01F2ADF0A923E0430AC4D28BF9E3B">A Revision Of The Indochinese Genera Cheirochela And Gestroiella Heteroptera Naucoridae And A Review Of The Tribe Cheirochelini</mods:title>
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<mods:namePart id="B76B503EB2EDA1853301D74078D12BEA">Dan</mods:namePart>
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<mods:namePart id="29F73B7CB76604F94D78EC3834DB05F4">Polhemus</mods:namePart>
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<mods:namePart id="0178947E2342A44FDED7458AAFC4843B">John</mods:namePart>
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<mods:namePart id="76059FAA962640787C7E28D23402A5B0">Polhemus</mods:namePart>
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<mods:name id="57E9A8F419E01DDC285D07B8D8A22845" type="personal">
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<mods:roleTerm id="0527E94322ECAC434173BD57C3C78DE3">Author</mods:roleTerm>
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<mods:namePart id="EE5EF763FADD49257A4B6D3F80827BF2">Sites, Robert</mods:namePart>
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<mods:typeOfResource id="D4586073ABB2DE0291B18BB8AC2425EB">text</mods:typeOfResource>
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<mods:title id="67A6D23C569F9B42D3F0B2EA02930FF1">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="1541981EAE2604868FB4EF875F5EA49B">
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<mods:date id="3FDEBFC8BE0A64068F72A891587CD664">2008</mods:date>
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<mods:number id="42214A2B055EA880CB50AB264CCE3F94">2008-08-31</mods:number>
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<mods:number id="F23AF5A3188BF678C88AC3CBD054205B">56</mods:number>
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<mods:number id="F2E824B8E30D87D1BBD6F1D04B031453">2</mods:number>
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<mods:start id="80EA0C8D912C92CE141CAA688EEB162D">255</mods:start>
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<treatment id="039F87C7E57BFFCCFC5BFEAB32C2FC2F" ID-DOI="http://doi.org/10.5281/zenodo.5416682" ID-Zenodo-Dep="5416682" LSID="urn:lsid:plazi:treatment:039F87C7E57BFFCCFC5BFEAB32C2FC2F" httpUri="http://treatment.plazi.org/id/039F87C7E57BFFCCFC5BFEAB32C2FC2F" lastPageId="5" lastPageNumber="260" pageId="4" pageNumber="259">
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<subSubSection id="C32C655AE57BFFCDFC5BFEAB3720FC6F" pageId="4" pageNumber="259" type="nomenclature">
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<paragraph id="8B8936D1E57BFFCDFC5BFEAB3641FECF" blockId="4.[929,1384,290,313]" box="[929,1384,290,313]" pageId="4" pageNumber="259">
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<heading id="D0C181BDE57BFFCDFC5BFEAB3641FECF" box="[929,1384,290,313]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="259" reason="3">
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<taxonomicName id="4C364D52E57BFFCDFC5BFEAB364DFECF" ID-CoL="TTYV" authority="Hope, 1841: 442" authorityName="Hope" authorityPageNumber="442" authorityYear="1841" box="[929,1380,290,313]" class="Insecta" family="Naucoridae" genus="Cheirochela" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="assamensis">
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<emphasis id="B942EAC3E57BFFCDFC5BFEAB378DFECF" box="[929,1188,290,313]" italics="true" pageId="4" pageNumber="259">Cheirochela assamensis</emphasis>
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Hope, 1841: 442
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</taxonomicName>
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.
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</heading>
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</paragraph>
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<paragraph id="8B8936D1E57BFFCDFCBFFEE83720FC6F" blockId="4.[837,1476,353,921]" pageId="4" pageNumber="259">
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<emphasis id="B942EAC3E57BFFCDFCBFFEE83746FE8F" box="[837,1135,353,377]" italics="true" pageId="4" pageNumber="259">
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<emphasis id="B942EAC3E57BFFCDFCBFFEE83741FE8F" bold="true" box="[837,1128,353,377]" italics="true" pageId="4" pageNumber="259">Supplemental redescription</emphasis>
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.
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</emphasis>
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<emphasis id="B942EAC3E57BFFCDFB89FEE83644FE8F" bold="true" box="[1139,1389,353,377]" pageId="4" pageNumber="259">– Brachypterous male:</emphasis>
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General colouration dark brown, with dark yellow markings on head, pronotum, hemelytra, and lateral abdomen; head dark yellow with dark, posteriorly opening V-shaped mark centrally on frons, posterior margin broadly dark, eyes black; pronotum narrowly margined with dark yellow laterally and posteriorly; hemelytra with claval suture and basal embolar area dark yellow; visible sections of abdominal tergites pale to medium brown, narrowly margined laterally with dark yellow. Overall form elongate-ovate (
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<figureCitation id="130D2A54E57BFFCDFBD9FD0B3749FD6F" box="[1059,1120,642,665]" captionStart="Figs" captionStartId="5.[115,159,2065,2086]" captionTargetBox="[118,1452,1214,2007]" captionTargetId="figure-16@5.[85,1470,1181,2031]" captionTargetPageId="5" captionText="Figs. 3–6. Cheirochela assamensis Hope, holotype male, structural details: 3, Dorsal habitus; 4, Left antenna, dorsal view; 5, Terminal abdomen, ventral view; 6, Medial processes on abdominal ventrites IV and V, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5340376" httpUri="https://zenodo.org/record/5340376/files/figure.png" pageId="4" pageNumber="259">Fig. 3</figureCitation>
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), size small for genus, body length 21.90, maximum width (across embolia) 11.90; head length 4.00, width 4.80, anterior/posterior interocular 3.30/3.00, eye length 2.00, width 1.00; pronotum length (midline) 2.50, width 8.60; scutellum length 3.10, width 5.00. Hemelytra brachypterous, venation obscure, corium finely and shallowly rugose, membrane rudiments strongly narrowed posteriorly, tips rounded, reaching to posterior margin of sixth visible abdominal tergite.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C655AE57BFFCCFCBFFC4833B4FCEF" lastPageId="5" lastPageNumber="260" pageId="4" pageNumber="259" type="discussion">
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<paragraph id="8B8936D1E57BFFCDFCBFFC48372FFBCF" blockId="4.[837,1476,961,1081]" pageId="4" pageNumber="259">
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<emphasis id="B942EAC3E57BFFCDFCBFFC4830F4FC2F" bold="true" box="[837,989,961,985]" italics="true" pageId="4" pageNumber="259">Discussion. –</emphasis>
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The
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<typeStatus id="548D8873E57BFFCDFBE1FC4B3753FC2F" box="[1051,1146,962,985]" pageId="4" pageNumber="259" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="4C364D52E57BFFCDFB5FFC4B366BFC2F" authorityName="Hope" authorityYear="1841" box="[1189,1346,962,985]" class="Insecta" family="Naucoridae" genus="Cheirochela" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="assamensis">
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<emphasis id="B942EAC3E57BFFCDFB5FFC4B366BFC2F" box="[1189,1346,962,985]" italics="true" pageId="4" pageNumber="259">C. assamensis</emphasis>
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</taxonomicName>
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came from an unspecified locality in
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<collectingRegion id="49F2F833E57BFFCDFB92FC6B379DFC0F" box="[1128,1204,994,1017]" country="India" name="Assam" pageId="4" pageNumber="259">Assam</collectingRegion>
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, northeastern
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<collectingCountry id="F3217641E57BFFCDFAABFC6B36A5FC0F" box="[1361,1420,994,1017]" name="India" pageId="4" pageNumber="259">India</collectingCountry>
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. The specimen is in the Hope Museum in Oxford, and bears the label “Type 437.”
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</paragraph>
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<caption id="DF496659E57BFFCDFCBEF7983756F7B5" ID-DOI="http://doi.org/10.5281/zenodo.5340374" ID-Zenodo-Dep="5340374" httpUri="https://zenodo.org/record/5340374/files/figure.png" pageId="4" pageNumber="259" startId="4.[836,871,2065,2086]" targetBox="[844,1475,1140,2016]" targetPageId="4">
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<paragraph id="8B8936D1E57BFFCDFCBEF7983756F7B5" blockId="4.[836,1475,2065,2115]" pageId="4" pageNumber="259">
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Fig. 2.
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<taxonomicName id="4C364D52E57BFFCDFC76F79B366BF7D0" authority="Polhemus & Polhemus." authorityName="Dan & Polhemus & John & Polhemus & Sites" authorityYear="2008" box="[908,1346,2065,2087]" class="Insecta" family="Naucoridae" genus="Cheirochela" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="tonkina">
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<emphasis id="B942EAC3E57BFFCDFC76F79B377BF7D1" box="[908,1106,2066,2087]" italics="true" pageId="4" pageNumber="259">Cheirochela tonkina</emphasis>
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Polhemus & Polhemus.
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</taxonomicName>
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Male, dorsal habitus. Drawn by Young Sohn.
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</paragraph>
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</caption>
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<paragraph id="8B8936D1E57AFFCCFF89FF4B3292FD6F" blockId="5.[115,754,194,665]" pageId="5" pageNumber="260">
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This is the smallest species in the genus, with the two examples seen having body lengths of 21.0 and
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<quantity id="4CCE9B34E57AFFCCFD75FF6B31DBFF0F" box="[655,754,226,249]" metricMagnitude="-2" metricUnit="m" metricValue="2.12" pageId="5" pageNumber="260" unit="mm" value="21.2">21.2 mm</quantity>
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respectively. This species is similar to
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<taxonomicName id="4C364D52E57AFFCCFDDCFE8B31FBFEEF" authorityName="Montandon" authorityYear="1897" box="[550,722,258,281]" class="Insecta" family="Naucoridae" genus="Cheirochela" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="birmaniensis">
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<emphasis id="B942EAC3E57AFFCCFDDCFE8B31FBFEEF" box="[550,722,258,281]" italics="true" pageId="5" pageNumber="260">C. birmaniensis</emphasis>
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</taxonomicName>
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in possessing well developed angulate tubercles posteromedially on abdominal sternites IV and V, but differs from that latter species in that these tubercles project primarily downward rather than being produced posteriorly over the bases of the following tergites (
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<figureCitation id="130D2A54E57AFFCCFEBFFE2B329BFE4F" box="[325,434,418,441]" captionStart="Figs" captionStartId="5.[115,159,2065,2086]" captionTargetBox="[118,1452,1214,2007]" captionTargetId="figure-16@5.[85,1470,1181,2031]" captionTargetPageId="5" captionText="Figs. 3–6. Cheirochela assamensis Hope, holotype male, structural details: 3, Dorsal habitus; 4, Left antenna, dorsal view; 5, Terminal abdomen, ventral view; 6, Medial processes on abdominal ventrites IV and V, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5340376" httpUri="https://zenodo.org/record/5340376/files/figure.png" pageId="5" pageNumber="260">Figs. 5, 6</figureCitation>
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). The male parameres of the two species are also differently formed (
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<figureCitation id="130D2A54E57AFFCCFDE6FE4B31A9FE2F" box="[540,640,450,473]" captionStart="Figs" captionStartId="6.[138,182,2065,2086]" captionTargetBox="[145,1490,188,2025]" captionTargetId="figure-14@6.[112,1498,155,2045]" captionTargetPageId="6" captionText="Figs. 7–11. Cheirochela species, male right paramere: 7, C. assamensis, holotype; 8, C. feana, paratype. 9. C. birmaniensis, holotype; 10, C. tonkina, paratype; 11, C. thailandana, paratype." figureDoi="http://doi.org/10.5281/zenodo.5340378" httpUri="https://zenodo.org/record/5340378/files/figure.png" pageId="5" pageNumber="260">Figs. 7, 8</figureCitation>
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), with that of
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<taxonomicName id="4C364D52E57AFFCCFF68FE6B3202FE0F" authorityName="Hope" authorityYear="1841" box="[146,299,482,505]" class="Insecta" family="Naucoridae" genus="Cheirochela" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="assamensis">
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<emphasis id="B942EAC3E57AFFCCFF68FE6B3202FE0F" box="[146,299,482,505]" italics="true" pageId="5" pageNumber="260">C. assamensis</emphasis>
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</taxonomicName>
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having a rounded projection on the basal half. Although Distant (1906) considered
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<taxonomicName id="4C364D52E57AFFCCFDBEFD8B31DBFDEF" authorityName="Montandon" authorityYear="1897" box="[580,754,514,537]" class="Insecta" family="Naucoridae" genus="Cheirochela" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="birmaniensis">
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<emphasis id="B942EAC3E57AFFCCFDBEFD8B31DBFDEF" box="[580,754,514,537]" italics="true" pageId="5" pageNumber="260">C. birmaniensis</emphasis>
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</taxonomicName>
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to be a synonym of
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<taxonomicName id="4C364D52E57AFFCCFEB7FDAB32CFFDCF" authorityName="Hope" authorityYear="1841" box="[333,486,546,569]" class="Insecta" family="Naucoridae" genus="Cheirochela" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="assamensis">
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<emphasis id="B942EAC3E57AFFCCFEB7FDAB32CFFDCF" box="[333,486,546,569]" italics="true" pageId="5" pageNumber="260">C. assamensis</emphasis>
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</taxonomicName>
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, based on a study of the
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<typeStatus id="548D8873E57AFFCCFF89FDCB33F2FDAF" box="[115,219,578,601]" pageId="5" pageNumber="260" type="holotype">holotypes</typeStatus>
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of both species, we feel that the characters noted above serve to separate the two species and therefore reject Distant’s proposed synonymy.
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</paragraph>
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<paragraph id="8B8936D1E57AFFCCFF89FD4B33B4FCEF" blockId="5.[115,754,706,793]" pageId="5" pageNumber="260">
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This species has so far been collected only from the upper Brahmaputra drainage basin in far northeastern
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<collectingCountry id="F3217641E57AFFCCFD85FD6B319FFD0F" box="[639,694,738,761]" name="India" pageId="5" pageNumber="260">India</collectingCountry>
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(
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<figureCitation id="130D2A54E57AFFCCFD3CFD6B33B8FCEF" captionStart="Fig" captionStartId="8.[836,870,2065,2086]" captionTargetBox="[836,1475,1238,2053]" captionTargetId="figure-11@8.[836,1475,1238,2053]" captionTargetPageId="8" captionText="Fig.16. Distributions of Cheirochela species in Indochina:squares = C. birmaniensis; diamonds = C. assamensis; circles = C. feana." figureDoi="http://doi.org/10.5281/zenodo.5340382" httpUri="https://zenodo.org/record/5340382/files/figure.png" pageId="5" pageNumber="260">Fig. 16</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C655AE57AFFCCFF89FCC832C2FC2F" pageId="5" pageNumber="260" type="materials_examined">
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<paragraph id="8B8936D1E57AFFCCFF89FCC832C2FC2F" blockId="5.[115,754,833,985]" pageId="5" pageNumber="260">
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<emphasis id="B942EAC3E57AFFCCFF89FCC83246FCAF" bold="true" box="[115,367,833,857]" italics="true" pageId="5" pageNumber="260">Material examined. –</emphasis>
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<materialsCitation id="3B5E3C8CE57AFFCCFE80FCCB316FFC8F" collectionCode="HEMO" country="India" location="Assam" pageId="5" pageNumber="260" specimenCount="1" stateProvince="Assam" typeStatus="holotype">
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<collectingCountry id="F3217641E57AFFCCFE80FCCB32E2FCAF" box="[378,459,834,857]" name="India" pageId="5" pageNumber="260">INDIA</collectingCountry>
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,
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<emphasis id="B942EAC3E57AFFCCFE20FCC83101FCAF" bold="true" box="[474,552,833,857]" pageId="5" pageNumber="260">
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<collectingRegion id="49F2F833E57AFFCCFE20FCC83101FCAF" box="[474,552,833,857]" country="India" name="Assam" pageId="5" pageNumber="260">Assam</collectingRegion>
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</emphasis>
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: 1 brachypterous male, no locality data (
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<typeStatus id="548D8873E57AFFCCFE82FCEB32F2FC8F" box="[376,475,866,889]" pageId="5" pageNumber="260" type="holotype">holotype</typeStatus>
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,
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<collectionCode id="ED27AE14E57AFFCCFE10FCEB3117FC8F" box="[490,574,866,889]" pageId="5" pageNumber="260">HEMO</collectionCode>
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||
)
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</materialsCitation>
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.
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<materialsCitation id="3B5E3C8CE57AFFCCFDA9FCE832CFFC2F" collectingDate="1996-05-09" collectingDateMax="1996-05-17" collectingDateMin="1996-05-09" collectionCode="NHMW" collectorName="Nokrek N. P. & E. Jendek & O. Sausa" country="India" elevation="1250" location="West Garo Hills" longitude="25.493334" pageId="5" pageNumber="260" specimenCode="WGS84" specimenCount="1" stateProvince="Meghalaya" typeStatus="holotype">
|
||
<emphasis id="B942EAC3E57AFFCCFDA9FCE831FBFC8F" bold="true" box="[595,722,865,889]" pageId="5" pageNumber="260">
|
||
<collectingRegion id="49F2F833E57AFFCCFDA9FCE831FBFC8F" box="[595,722,865,889]" country="India" name="Meghalaya" pageId="5" pageNumber="260">Meghalaya</collectingRegion>
|
||
</emphasis>
|
||
: 1 brachypterous male,
|
||
<location id="8EE9600AE57AFFCCFEADFC0B3125FC6F" LSID="urn:lsid:plazi:treatment:039F87C7E57BFFCCFC5BFEAB32C2FC2F:8EE9600AE57AFFCCFEADFC0B3125FC6F" box="[343,524,898,921]" country="India" longitude="25.493334" name="West Garo Hills" pageId="5" pageNumber="260" stateProvince="Meghalaya">West Garo Hills</location>
|
||
,
|
||
<collectorName id="26C35307E57AFFCCFDE0FC0B3187FC6F" box="[538,686,898,921]" pageId="5" pageNumber="260">Nokrek N. P.</collectorName>
|
||
,
|
||
<date id="FF881011E57AFFCCFD46FC0B33F6FC4F" pageId="5" pageNumber="260" value="1996-05-09" valueMax="1996-05-17" valueMin="1996-05-09">
|
||
<collectingDate id="EFCCE9F9E57AFFCCFD46FC0B33F6FC4F" pageId="5" pageNumber="260" value="1996-05-09" valueMax="1996-05-17" valueMin="1996-05-09">9–17 May 1996</collectingDate>
|
||
</date>
|
||
, 950–
|
||
<quantity id="4CCE9B34E57AFFCCFEE4FC2B3244FC4F" box="[286,365,930,953]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="5" pageNumber="260" unit="m" value="1250.0">
|
||
<elevation id="001BD1E2E57AFFCCFEE4FC2B3244FC4F" box="[286,365,930,953]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="5" pageNumber="260" unit="m" value="1250.0">1250 m</elevation>
|
||
</quantity>
|
||
, 90°19.5'
|
||
<geoCoordinate id="EE025016E57AFFCCFE28FC2B3110FC4F" box="[466,569,930,953]" degrees="25" direction="east" minutes="29.6" orientation="longitude" pageId="5" pageNumber="260" precision="92" value="25.493334">25°29.6'E</geoCoordinate>
|
||
(
|
||
<specimenCode id="DB909EAAE57AFFCCFDBFFC2B31B5FC4F" box="[581,668,930,953]" collectionCode="WGS" pageId="5" pageNumber="260">WGS84</specimenCode>
|
||
datum),
|
||
<collectorName id="26C35307E57AFFCCFF89FC4B33F5FC2F" box="[115,220,962,985]" pageId="5" pageNumber="260">E. Jendek</collectorName>
|
||
and
|
||
<collectorName id="26C35307E57AFFCCFEE8FC4B325BFC2F" box="[274,370,962,985]" pageId="5" pageNumber="260">O. Sausa</collectorName>
|
||
(
|
||
<collectionCode id="ED27AE14E57AFFCCFE78FC4B32C9FC2F" box="[386,480,962,985]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="260" type="Museum">NHMW</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |