269 lines
37 KiB
XML
269 lines
37 KiB
XML
<document id="1216F4AB61381FAB240DC72A1B86C1CC" ID-DOI="10.5281/zenodo.174082" ID-GBIF-Dataset="3ec64760-8a98-4411-811e-919b308ed4a8" ID-ISSN="11755326" ID-Zenodo-Dep="174082" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459790225771" checkinUser="plazi" docAuthor="Golovatch, Sergei I. & Geoffroy, Jean-Jacques" docDate="2006" docId="039F87BBD50365006975FA84FBD2F89E" docLanguage="en" docName="zt01325p368.pdf" docOrigin="Zootaxa 1325" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Pacidesmus martensi Golovatch & Geoffroy, 2006, sp. n." docType="treatment" docVersion="8" lastPageNumber="367" masterDocId="FFA6FFC3D5026504687DFF88FFBFFFB4" masterDocTitle="Golovatch, with the description of a new troglobitic species from southern China (Diplopoda: Polydesmida: Polydesmidae)" masterLastPageNumber="368" masterPageNumber="363" pageNumber="364" updateTime="1698219128824" updateUser="plazi">
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<mods:mods id="C22329165D478A7582F26A884DB6CFED" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="08567AA05E87A3D24ECBF5F3950CE3D8">Golovatch, with the description of a new troglobitic species from southern China (Diplopoda: Polydesmida: Polydesmidae)</mods:title>
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<mods:name id="A455AB779A560B7579A94AC2A18AFB7B" type="personal">
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<mods:roleTerm id="3350A06B7A6CF3F1346CDCBA1A09FD40">Author</mods:roleTerm>
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<mods:namePart id="B337DF6A6DF5DB10CE8E8C69CCEE4129">Golovatch, Sergei I.</mods:namePart>
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<mods:name id="B2589C7A7B3E00B54E10545291E8512D" type="personal">
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<mods:roleTerm id="490CD4DD6D185E0C1B46F99CBFE0C61C">Author</mods:roleTerm>
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<mods:namePart id="92C973B581DEF87ADEC5E012234227DF">Geoffroy, Jean-Jacques</mods:namePart>
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<mods:typeOfResource id="9029DFF83F18ADA0114BDD7FF95D6B04">text</mods:typeOfResource>
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<mods:title id="F3DDD66F534DAEAA2408E0956CA337EB">Zootaxa</mods:title>
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<mods:part id="B6754EC3F1E1C19BD882DD544D5F2FE4">
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<mods:date id="5FE8C244FD369B57E80441E919FBF55E">2006</mods:date>
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<mods:detail id="D0C1F7684CB51A6C879857880CA9BFC0" type="volume">
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<mods:number id="4683844AECEDC36084E146067C231F8A">1325</mods:number>
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<mods:start id="77B729C88D0F4720325B4E205A382139">363</mods:start>
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<mods:end id="774E96E94416560210BC72187E1CD937">368</mods:end>
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<mods:classification id="7BE2A98DAD7552A24B446DDCA256AE83">journal article</mods:classification>
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<mods:identifier id="07ECFBBB397D45AB906DC22C01335C21" type="DOI">10.5281/zenodo.174082</mods:identifier>
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<mods:identifier id="97BCC44D5CD7A33263826D2F7C9B6FB8" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="591144E8C7114ADF8B0D98E1795CB26F" type="Zenodo-Dep">174082</mods:identifier>
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<treatment id="039F87BBD50365006975FA84FBD2F89E" ID-DOI="http://doi.org/10.5281/zenodo.6262505" ID-GBIF-Taxon="119350228" ID-Zenodo-Dep="6262505" LSID="urn:lsid:plazi:treatment:039F87BBD50365006975FA84FBD2F89E" httpUri="http://treatment.plazi.org/id/039F87BBD50365006975FA84FBD2F89E" lastPageId="4" lastPageNumber="367" pageId="1" pageNumber="364">
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<subSubSection id="C32C6526D50365056975FA84FEC2FAFD" pageId="1" pageNumber="364" type="nomenclature">
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<paragraph id="8B8936ADD50365056975FA84FDEDFA92" blockId="1.[264,594,1292,1353]" box="[264,594,1292,1318]" pageId="1" pageNumber="364">
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<heading id="D0C181C1D50365056975FA84FDEDFA92" bold="true" box="[264,594,1292,1318]" fontSize="11" level="1" pageId="1" pageNumber="364" reason="1">
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<emphasis id="B942EABFD50365056975FA84FDEDFA92" bold="true" box="[264,594,1292,1318]" pageId="1" pageNumber="364">
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<taxonomicName id="4C364D2ED50365056975FA84FDB7FA92" ID-CoL="4BW3V" box="[264,520,1292,1318]" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="364" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
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<emphasis id="B942EABFD50365056975FA84FDB7FA92" bold="true" box="[264,520,1292,1318]" italics="true" pageId="1" pageNumber="364">Pacidesmus martensi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27157C4D50365056A6DFA84FDEDFA92" box="[528,594,1292,1318]" pageId="1" pageNumber="364" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8936ADD50365056975FAA7FEC2FAFD" blockId="1.[264,594,1292,1353]" box="[264,381,1327,1353]" pageId="1" pageNumber="364">
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<figureCitation id="130D2A28D50365056975FAA7FEC2FAFD" box="[264,381,1327,1353]" captionStart-0="FIGURES 1 – 8" captionStart-1="FIGURES 9 – 12" captionStartId-0="3.[264,383,1374,1398]" captionStartId-1="4.[264,383,1572,1596]" captionTargetBox-0="[441,1147,296,1354]" captionTargetBox-1="[408,1192,513,1559]" captionTargetId-0="figure@3.[439,1148,296,1362]" captionTargetId-1="figure@4.[389,1199,497,1579]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1 – 8. Pacidesmus martensi sp. n., ɗ holotype. 1 – 2 Anterior body part, dorsal and lateral views, respectively; 3 – 4 Midbody segments (3 dorsal, 4 lateral view); 5 Segment 8, dorsal view; 6 – 7 Caudal body part (6 dorsal, 7 lateral view); 8 Right gonopod, lateral view. Scale bars 1.0 mm (Fig. 8 not to scale)." captionText-1="FIGURES 9 – 12. Pacidesmus martensi sp. n., ɗ holotype. 9 Antenna; 10 Leg 9; 11 – 12 Left gonopod (11 mesal, 12 lateral). Scale bar 0.3 mm." httpUri-0="https://zenodo.org/record/174083/files/figure.png" httpUri-1="https://zenodo.org/record/174084/files/figure.png" pageId="1" pageNumber="364">Figs 1–12</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C6526D50365056975FAF9FCEBF99F" pageId="1" pageNumber="364" type="materials_examined">
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<paragraph id="8B8936ADD50365056975FAF9FCEBF99F" blockId="1.[264,1323,1393,1899]" pageId="1" pageNumber="364">
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<emphasis id="B942EABFD50365056975FAF9FE49FA3F" bold="true" box="[264,502,1393,1419]" pageId="1" pageNumber="364">
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<emphasis id="B942EABFD50365056975FAF9FE50FA3F" bold="true" box="[264,495,1393,1419]" italics="true" pageId="1" pageNumber="364">Material examined</emphasis>
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.
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</emphasis>
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ɗ
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<typeStatus id="548D880FD50365056A63FAF9FD3BFA3F" box="[542,644,1393,1419]" pageId="1" pageNumber="364" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="ED27AE68D50365056AEBFAF9FD4DFA3F" LSID="urn:lsid:biocol.org:col:34988" box="[662,754,1393,1419]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="364">MNHN</collectionCode>
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<collectionCode id="ED27AE68D50365056A81FAF9FCA2FA3F" box="[764,797,1393,1419]" pageId="1" pageNumber="364">JC</collectionCode>
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306),
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<collectingCountry id="F321763DD50365056B1CFAF9FC15FA3F" box="[865,938,1393,1419]" name="China" pageId="1" pageNumber="364">China</collectingCountry>
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, Guizhou Prov., Qianxi County, Hong Lin Town, Ishui Luo Dong Cave (= Waterfall Cave),
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<date id="FF88106DD50365056BB9FA11FBF3FA07" box="[964,1100,1433,1459]" pageId="1" pageNumber="364" value="2003-11-16">16.XI.2003</date>
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, leg. L. Latella. —
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<typeStatus id="548D880FD50365056975FA49FEC0FA6F" box="[264,383,1473,1499]" pageId="1" pageNumber="364" type="paratype">Paratypes</typeStatus>
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: 2 ɗ, 4 Ψ,
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<specimenCount id="9D30FD24D50365056986FA49FD82FA6F" box="[507,573,1473,1499]" pageId="1" pageNumber="364" type="generic">1 juv.</specimenCount>
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(
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<collectionCode id="ED27AE68D50365056A36FA49FD18FA6F" LSID="urn:lsid:biocol.org:col:34988" box="[587,679,1473,1499]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="364">MNHN</collectionCode>
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<collectionCode id="ED27AE68D50365056AD2FA49FD6FFA6F" box="[687,720,1473,1499]" pageId="1" pageNumber="364">JC</collectionCode>
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306), 1 ɗ, 1 Ψ (
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<collectionCode id="ED27AE68D50365056BF9FA49FC5AFA6F" box="[900,997,1473,1499]" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="1" pageNumber="364">ZMUM</collectionCode>
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), 1 ɗ, 1 Ψ (BDBFUL), 1 ɗ, 1 Ψ (
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<collectionCode id="ED27AE68D50365056932FA61FE15F9B7" box="[335,426,1513,1539]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="1" pageNumber="364">ZMUC</collectionCode>
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), Guizhou Prov., Dafang County, Yangzhamba Village, Hei Dong Cave (terrestrial),
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<date id="FF88106DD503650569E7F999FE4DF99F" box="[410,498,1553,1579]" pageId="1" pageNumber="364" value="2004-02">II.2004</date>
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, leg. S. Prevorčnik & B. Sket.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C6526D50365056945F9B1FC37F917" pageId="1" pageNumber="364" type="etymology">
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<paragraph id="8B8936ADD50365056945F9B1FC37F917" blockId="1.[264,1323,1393,1899]" pageId="1" pageNumber="364">
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<emphasis id="B942EABFD50365056945F9B1FE02F9E7" bold="true" box="[312,445,1593,1619]" pageId="1" pageNumber="364">
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<emphasis id="B942EABFD50365056945F9B1FE08F9E7" bold="true" box="[312,439,1593,1619]" italics="true" pageId="1" pageNumber="364">Etymology</emphasis>
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.
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</emphasis>
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We take pleasure in dedicating this species to Prof. Dr Jochen Martens (Mainz University), a devoted student and collector of soil arthropods, birds and other animals in various parts of the world, including
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<collectingCountry id="F321763DD50365056B47F901FC3BF917" box="[826,900,1673,1699]" name="China" pageId="1" pageNumber="364">China</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C6526D50365056945F939FCB3F8AF" pageId="1" pageNumber="364" type="diagnosis">
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<paragraph id="8B8936ADD50365056945F939FCB3F8AF" blockId="1.[264,1323,1393,1899]" pageId="1" pageNumber="364">
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<emphasis id="B942EABFD50365056945F939FE07F97F" bold="true" box="[312,440,1713,1739]" italics="true" pageId="1" pageNumber="364">Diagnosis.</emphasis>
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Differs from congeners by the strongly elevated paraterga with narrowly rounded caudal corners, the poorly developed accessory seminal chamber, and the shape of the gonopod acropodite (see Key below).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C6526D50365076945F8A1FC57F9E0" lastPageId="3" lastPageNumber="366" pageId="1" pageNumber="364" type="description">
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<paragraph id="8B8936ADD50365066945F8A1FD1BFEEA" blockId="1.[264,1323,1393,1899]" lastBlockId="2.[264,1324,283,1870]" lastPageId="2" lastPageNumber="365" pageId="1" pageNumber="364">
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<emphasis id="B942EABFD50365056945F8A1FE74F8F7" bold="true" box="[312,459,1833,1859]" pageId="1" pageNumber="364">
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<emphasis id="B942EABFD50365056945F8A1FE7BF8F7" bold="true" box="[312,452,1833,1859]" italics="true" pageId="1" pageNumber="364">Description</emphasis>
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.
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</emphasis>
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Length of
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<typeStatus id="548D880FD50365056A2CF8A1FD08F8F7" box="[593,695,1833,1859]" pageId="1" pageNumber="364" type="holotype">holotype</typeStatus>
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about
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<quantity id="4CCE9B48D50365056B76F8A1FCDDF8F7" box="[779,866,1833,1859]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="364" unit="mm" value="21.0">21 mm</quantity>
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, width of midbody pro and metazona 1.3 and
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<quantity id="4CCE9B48D5036505691AF8D9FE7AF8DF" box="[359,453,1873,1899]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="364" unit="mm" value="2.1">2.1 mm</quantity>
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, respectively.
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<typeStatus id="548D880FD50365056A16F8D9FD62F8DF" box="[619,733,1873,1899]" pageId="1" pageNumber="364" type="paratype">Paratypes</typeStatus>
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<quantity id="4CCE9B48D50365056A98F8D9FCD8F8DF" box="[741,871,1873,1899]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.2" metricValueMin="1.7" pageId="1" pageNumber="364" unit="mm" value="19.5" valueMax="22.0" valueMin="17.0">17–22 mm</quantity>
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, width of midbody pro and metazona 1.3–1.8 and
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<quantity id="4CCE9B48D500650669DFFE94FD89FE82" box="[418,566,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.8" metricValueMin="2.1" pageId="2" pageNumber="365" unit="mm" value="2.45" valueMax="2.8" valueMin="2.1">2.1–2.8 mm</quantity>
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; ɗ usually slightly smaller than Ψ. Coloration entirely pallid, whitish, with pinkish tint in places.
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</paragraph>
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<paragraph id="8B8936ADD50065066945FEE4FEDEFD92" blockId="2.[264,1324,283,1870]" pageId="2" pageNumber="365">
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Head usual, densely setose; genae roundly subquadrate; epicranial groove evident but shallow. Antennae (
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<figureCitation id="130D2A28D5006506698EFE1CFDE0FE1A" box="[499,607,404,430]" captionStart="FIGURES 1 – 8" captionStartId="3.[264,383,1374,1398]" captionTargetBox="[441,1147,296,1354]" captionTargetId="figure@3.[439,1148,296,1362]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Pacidesmus martensi sp. n., ɗ holotype. 1 – 2 Anterior body part, dorsal and lateral views, respectively; 3 – 4 Midbody segments (3 dorsal, 4 lateral view); 5 Segment 8, dorsal view; 6 – 7 Caudal body part (6 dorsal, 7 lateral view); 8 Right gonopod, lateral view. Scale bars 1.0 mm (Fig. 8 not to scale)." httpUri="https://zenodo.org/record/174083/files/figure.png" pageId="2" pageNumber="365">Figs 1–2</figureCitation>
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,
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<figureCitation id="130D2A28D50065066A10FE1CFDC3FE1A" box="[621,636,404,430]" captionStart="FIGURES 9 – 12" captionStartId="4.[264,383,1572,1596]" captionTargetBox="[408,1192,513,1559]" captionTargetId="figure@4.[389,1199,497,1579]" captionTargetPageId="4" captionText="FIGURES 9 – 12. Pacidesmus martensi sp. n., ɗ holotype. 9 Antenna; 10 Leg 9; 11 – 12 Left gonopod (11 mesal, 12 lateral). Scale bar 0.3 mm." httpUri="https://zenodo.org/record/174084/files/figure.png" pageId="2" pageNumber="365">9</figureCitation>
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) long and slender, weakly clavate; antennomeres 5 and 6 each with a distodorsal field of minute bacilliform sensilla; antennomere 7 dorsally with a small, but evident, claviform, transparent tubercle at base and a flat tubercle in distal quarter.
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</paragraph>
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<paragraph id="8B8936ADD50065066945FDBCFD29FB42" blockId="2.[264,1324,283,1870]" pageId="2" pageNumber="365">
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Body polydesmoid, with 20 segments. Collum subequal in width to head and segments 2 and 3, with three indistinct transverse rows of very flat setigerous bosses; setae of front row slightly longer than others; paraterga broadly rounded, slightly elevated up to level of dorsum, devoid of evident indentations. Body parallelsided on segments 7–17, thereafter abruptly tapering caudally. Paraterga of segment 2 elevated, not surpassing rear tergal contour. Subsequent paraterga slightly projecting caudad beyond rear tergal contour, evidently elevated above dorsum until about segment 10, onward about level to dorsum until segment 19, evidently below level of dorsum only on segment 19 (
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||
<figureCitation id="130D2A28D50065066C25FCC4FB7BFCD2" box="[1112,1220,844,870]" captionStart="FIGURES 1 – 8" captionStartId="3.[264,383,1374,1398]" captionTargetBox="[441,1147,296,1354]" captionTargetId="figure@3.[439,1148,296,1362]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Pacidesmus martensi sp. n., ɗ holotype. 1 – 2 Anterior body part, dorsal and lateral views, respectively; 3 – 4 Midbody segments (3 dorsal, 4 lateral view); 5 Segment 8, dorsal view; 6 – 7 Caudal body part (6 dorsal, 7 lateral view); 8 Right gonopod, lateral view. Scale bars 1.0 mm (Fig. 8 not to scale)." httpUri="https://zenodo.org/record/174083/files/figure.png" pageId="2" pageNumber="365">Figs 1–7</figureCitation>
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||
). Caudal corners of all paraterga more or less narrowly rounded, never pointed. Tergal surface smooth, with three transverse setigerous rows of very flat bosses, forming a typical pattern (
|
||
<figureCitation id="130D2A28D5006506696CFC4CFEDFFC6A" box="[273,352,964,990]" captionStart="FIGURES 1 – 8" captionStartId="3.[264,383,1374,1398]" captionTargetBox="[441,1147,296,1354]" captionTargetId="figure@3.[439,1148,296,1362]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Pacidesmus martensi sp. n., ɗ holotype. 1 – 2 Anterior body part, dorsal and lateral views, respectively; 3 – 4 Midbody segments (3 dorsal, 4 lateral view); 5 Segment 8, dorsal view; 6 – 7 Caudal body part (6 dorsal, 7 lateral view); 8 Right gonopod, lateral view. Scale bars 1.0 mm (Fig. 8 not to scale)." httpUri="https://zenodo.org/record/174083/files/figure.png" pageId="2" pageNumber="365">Fig. 5</figureCitation>
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); setae minute, very short. Prozona very finely shagreened, metazona below paraterga microgranulate. Structure between pro and metazona nearly smooth, only sides with extremely faint transverse striolation. Limbus microdenticulate. Porebearing paraterga with four, poreless paraterga with three, minute lateral incisions. Pore formula normal, ozopores lying entirely dorsally, at bottom of a lanceolate groove near lateral margin between two caudolateral incisions (
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<figureCitation id="130D2A28D50065066B54FB04FC21FB12" box="[809,926,1164,1190]" captionStart="FIGURES 1 – 8" captionStartId="3.[264,383,1374,1398]" captionTargetBox="[441,1147,296,1354]" captionTargetId="figure@3.[439,1148,296,1362]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Pacidesmus martensi sp. n., ɗ holotype. 1 – 2 Anterior body part, dorsal and lateral views, respectively; 3 – 4 Midbody segments (3 dorsal, 4 lateral view); 5 Segment 8, dorsal view; 6 – 7 Caudal body part (6 dorsal, 7 lateral view); 8 Right gonopod, lateral view. Scale bars 1.0 mm (Fig. 8 not to scale)." httpUri="https://zenodo.org/record/174083/files/figure.png" pageId="2" pageNumber="365">Figs 3, 5</figureCitation>
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). Epiproct coniform (
|
||
<figureCitation id="130D2A28D50065066CD0FB04FAA2FB12" box="[1197,1309,1164,1190]" captionStart="FIGURES 1 – 8" captionStartId="3.[264,383,1374,1398]" captionTargetBox="[441,1147,296,1354]" captionTargetId="figure@3.[439,1148,296,1362]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Pacidesmus martensi sp. n., ɗ holotype. 1 – 2 Anterior body part, dorsal and lateral views, respectively; 3 – 4 Midbody segments (3 dorsal, 4 lateral view); 5 Segment 8, dorsal view; 6 – 7 Caudal body part (6 dorsal, 7 lateral view); 8 Right gonopod, lateral view. Scale bars 1.0 mm (Fig. 8 not to scale)." httpUri="https://zenodo.org/record/174083/files/figure.png" pageId="2" pageNumber="365">Figs 6–7</figureCitation>
|
||
), barely surpassing paraprocts caudally. Hypoproct distinctly trapeziform, with a pair of long setae at caudolateral corners.
|
||
</paragraph>
|
||
<paragraph id="8B8936ADD50065066945FA8CFC3DFADA" blockId="2.[264,1324,283,1870]" pageId="2" pageNumber="365">
|
||
A distinct pleurosternal flapshaped crest present only on segment 2 (
|
||
<figureCitation id="130D2A28D50065066C23FA8CFB1AFAAA" box="[1118,1189,1284,1310]" captionStart="FIGURES 1 – 8" captionStartId="3.[264,383,1374,1398]" captionTargetBox="[441,1147,296,1354]" captionTargetId="figure@3.[439,1148,296,1362]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Pacidesmus martensi sp. n., ɗ holotype. 1 – 2 Anterior body part, dorsal and lateral views, respectively; 3 – 4 Midbody segments (3 dorsal, 4 lateral view); 5 Segment 8, dorsal view; 6 – 7 Caudal body part (6 dorsal, 7 lateral view); 8 Right gonopod, lateral view. Scale bars 1.0 mm (Fig. 8 not to scale)." httpUri="https://zenodo.org/record/174083/files/figure.png" pageId="2" pageNumber="365">Fig. 2</figureCitation>
|
||
), thereafter totally wanting. Sterna evidently setose, but without modifications. Gonopod aperture obcordate, with a small, rounded shelf caudomedially.
|
||
</paragraph>
|
||
<paragraph id="8B8936ADD50065066945FAF4FCAEFA52" blockId="2.[264,1324,283,1870]" pageId="2" pageNumber="365">
|
||
Legs very long and slender (
|
||
<figureCitation id="130D2A28D50065066AF6FAF4FCAAFA22" box="[651,789,1404,1430]" captionStart="FIGURES 1 – 8" captionStartId="3.[264,383,1374,1398]" captionTargetBox="[441,1147,296,1354]" captionTargetId="figure@3.[439,1148,296,1362]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Pacidesmus martensi sp. n., ɗ holotype. 1 – 2 Anterior body part, dorsal and lateral views, respectively; 3 – 4 Midbody segments (3 dorsal, 4 lateral view); 5 Segment 8, dorsal view; 6 – 7 Caudal body part (6 dorsal, 7 lateral view); 8 Right gonopod, lateral view. Scale bars 1.0 mm (Fig. 8 not to scale)." httpUri="https://zenodo.org/record/174083/files/figure.png" pageId="2" pageNumber="365">Figs 2, 4, 7</figureCitation>
|
||
,
|
||
<figureCitation id="130D2A28D50065066B59FAF4FCFBFA22" box="[804,836,1404,1430]" captionStart="FIGURES 9 – 12" captionStartId="4.[264,383,1572,1596]" captionTargetBox="[408,1192,513,1559]" captionTargetId="figure@4.[389,1199,497,1579]" captionTargetPageId="4" captionText="FIGURES 9 – 12. Pacidesmus martensi sp. n., ɗ holotype. 9 Antenna; 10 Leg 9; 11 – 12 Left gonopod (11 mesal, 12 lateral). Scale bar 0.3 mm." httpUri="https://zenodo.org/record/174084/files/figure.png" pageId="2" pageNumber="365">10</figureCitation>
|
||
), especially so towards caudal body end, nearly twice as long as body height; telopodite devoid of sphaerotrichomes; claw very long and slender, distinctly curved (
|
||
<figureCitation id="130D2A28D50065066AD1FA44FCBBFA52" box="[684,772,1484,1510]" captionStart="FIGURES 9 – 12" captionStartId="4.[264,383,1572,1596]" captionTargetBox="[408,1192,513,1559]" captionTargetId="figure@4.[389,1199,497,1579]" captionTargetPageId="4" captionText="FIGURES 9 – 12. Pacidesmus martensi sp. n., ɗ holotype. 9 Antenna; 10 Leg 9; 11 – 12 Left gonopod (11 mesal, 12 lateral). Scale bar 0.3 mm." httpUri="https://zenodo.org/record/174084/files/figure.png" pageId="2" pageNumber="365">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8936ADD50065066945FA7CFB05F8FA" blockId="2.[264,1324,283,1870]" pageId="2" pageNumber="365">
|
||
Gonopods (
|
||
<figureCitation id="130D2A28D500650669B7FA7CFD9FF9BA" box="[458,544,1524,1550]" captionStart="FIGURES 1 – 8" captionStartId="3.[264,383,1374,1398]" captionTargetBox="[441,1147,296,1354]" captionTargetId="figure@3.[439,1148,296,1362]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Pacidesmus martensi sp. n., ɗ holotype. 1 – 2 Anterior body part, dorsal and lateral views, respectively; 3 – 4 Midbody segments (3 dorsal, 4 lateral view); 5 Segment 8, dorsal view; 6 – 7 Caudal body part (6 dorsal, 7 lateral view); 8 Right gonopod, lateral view. Scale bars 1.0 mm (Fig. 8 not to scale)." httpUri="https://zenodo.org/record/174083/files/figure.png" pageId="2" pageNumber="365">Figs 8</figureCitation>
|
||
,
|
||
<figureCitation id="130D2A28D50065066A45FA7CFD3EF9BA" box="[568,641,1524,1550]" captionStart="FIGURES 9 – 12" captionStartId="4.[264,383,1572,1596]" captionTargetBox="[408,1192,513,1559]" captionTargetId="figure@4.[389,1199,497,1579]" captionTargetPageId="4" captionText="FIGURES 9 – 12. Pacidesmus martensi sp. n., ɗ holotype. 9 Antenna; 10 Leg 9; 11 – 12 Left gonopod (11 mesal, 12 lateral). Scale bar 0.3 mm." httpUri="https://zenodo.org/record/174084/files/figure.png" pageId="2" pageNumber="365">11–12</figureCitation>
|
||
) with typical, enlarged, subtriangular, medially fully coalesced coxae, each coxa bearing 2–3 macrosetae ventrally and a cannula dorsomedially. Telopodite elongate, relatively simple; acropodite (= branch lying distally of recurvature point of seminal groove,
|
||
<emphasis id="B942EABFD50065066A4CF9E4FDF1F931" box="[561,590,1644,1669]" italics="true" pageId="2" pageNumber="365">cf.</emphasis>
|
||
<bibRefCitation id="EFA74B5CD50065066A2BF9E4FCA9F932" author="Golovatch" box="[598,790,1644,1670]" pageId="2" pageNumber="368" refString="Golovatch, S. I. (1991) The millipede family Polydesmidae in Southeast Asia, with notes on phylogeny (Diplopoda: Polydesmida). Steenstrupia, 17 (4), 141 - 159." type="journal article" year="1991">Golovatch 1991</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA74B5CD50065066B5FF9E4FBA6F932" author="Djursvoll" box="[802,1049,1644,1670]" pageId="2" pageNumber="368" refString="Djursvoll, P., Golovatch, S. I., Johanson, K. A. & Meidell, B. (2001) Phylogenetic relationships within Polydesmus sensu lato (Diplopoda: Polydesmidae). Fragmenta Faunistica, 43 (Suppl.), 37 - 59 (for 2000)." type="journal article" year="2001">
|
||
Djursvoll
|
||
<emphasis id="B942EABFD50065066BE4F9E4FC6BF931" box="[921,980,1644,1669]" italics="true" pageId="2" pageNumber="365">et al.</emphasis>
|
||
2001
|
||
</bibRefCitation>
|
||
) split into two unequal parts: a longer, slender, distally curved endomere (
|
||
<emphasis id="B942EABFD50065066B28F91CFCDBF91A" bold="true" box="[853,868,1684,1710]" pageId="2" pageNumber="365">a</emphasis>
|
||
) and a shorter, lobuliform exomere (
|
||
<emphasis id="B942EABFD50065066D6CF91CFAA6F91A" bold="true" box="[1297,1305,1684,1710]" pageId="2" pageNumber="365">l</emphasis>
|
||
). Endomere with a small denticle (
|
||
<emphasis id="B942EABFD50065066AF3F934FD21F962" bold="true" box="[654,670,1724,1750]" pageId="2" pageNumber="365">d</emphasis>
|
||
) frontally at base. Region near pulvillus, and that lying distolaterally of recurvature point of seminal groove, conspicuously papillatespinulate, often with another small but clear denticle distally on mesal face. Accessory seminal chamber very small, inconspicuous. Orifice on pulvillus surrounded by spinules.
|
||
</paragraph>
|
||
<caption id="DF496625D50165076975FAD6FE60FA6B" httpUri="https://zenodo.org/record/174083/files/figure.png" pageId="3" pageNumber="366" targetBox="[441,1147,296,1354]" targetPageId="3">
|
||
<paragraph id="8B8936ADD50165076975FAD6FE60FA6B" blockId="3.[264,1323,1372,1503]" pageId="3" pageNumber="366">
|
||
<emphasis id="B942EABFD50165076975FAD6FE13FAC2" bold="true" box="[264,428,1374,1398]" pageId="3" pageNumber="366">FIGURES 1–8</emphasis>
|
||
.
|
||
<taxonomicName id="4C364D2ED501650769C7FAD7FD22FAC2" box="[442,669,1375,1398]" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="366" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
|
||
<emphasis id="B942EABFD501650769C7FAD7FD22FAC2" box="[442,669,1375,1398]" italics="true" pageId="3" pageNumber="366">Pacidesmus martensi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27157C4D50165076AD9FAD6FD60FAC2" box="[676,735,1374,1398]" pageId="3" pageNumber="366" rank="species">
|
||
<emphasis id="B942EABFD50165076AD9FAD6FD60FAC2" bold="true" box="[676,735,1374,1398]" pageId="3" pageNumber="366">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, ɗ holotype. 1–2 Anterior body part, dorsal and lateral views, respectively; 3–4 Midbody segments (3 dorsal, 4 lateral view); 5 Segment 8, dorsal view; 6–7 Caudal body part (6 dorsal, 7 lateral view); 8 Right gonopod, lateral view. Scale bars 1.0 mm (Fig. 8 not to scale).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8936ADD50165076945F99AFC57F9E0" blockId="3.[264,1323,1554,1900]" pageId="3" pageNumber="366">Epigynal ridge low, slightly elevated and rounded only laterally. Vulvae usual, elongate, densely setose, with spiral canals at bottom of gutter.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32C6526D50165006945F9EBFBD2F89E" lastPageId="4" lastPageNumber="367" pageId="3" pageNumber="366" type="discussion">
|
||
<paragraph id="8B8936ADD50165006945F9EBFBCBFE62" blockId="3.[264,1323,1554,1900]" lastBlockId="4.[264,1323,284,470]" lastPageId="4" lastPageNumber="367" pageId="3" pageNumber="366">
|
||
<emphasis id="B942EABFD50165076945F9EBFE19F9C8" box="[312,422,1635,1660]" italics="true" pageId="3" pageNumber="366">Remarks.</emphasis>
|
||
Because this remarkable species shows several evident traits of troglomorphism, such as the completely unpigmented tegument and the elongate appendages, it can be considered as troglobitic with a fair degree of certainty. The cave Ishui Luo Dong supports two further diplopods, both presumably troglobitic and highly local:
|
||
<taxonomicName id="4C364D2ED5016507692CF88BFD40F8A8" authority="Mauries, 2005" authorityName="Mauries" authorityYear="2005" box="[337,767,1794,1820]" class="Diplopoda" family="Guizhousomatidae" genus="Guizhousoma" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="366" phylum="Arthropoda" rank="species" species="latellai">
|
||
<emphasis id="B942EABFD5016507692CF88BFDF2F8A8" box="[337,589,1795,1820]" italics="true" pageId="3" pageNumber="366">Guizhousoma latellai</emphasis>
|
||
<bibRefCitation id="EFA74B5CD50165076A28F88AFD40F8A8" author="Mauries" box="[597,767,1794,1820]" pageId="3" pageNumber="368" refString="Mauries, J. - P. (2005) Guizhousoma latellai gen. n., sp. n., de Chine continentale, type d'une nouvelle famille de la superfamille des Neoatractosomatoidea (Diplopoda: Chordeumatida). Arthropoda Selecta, 14 (1), 11 - 17." type="journal article" year="2005">Mauriès, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C364D2ED50165076B73F88AFC77F8A8" box="[782,968,1794,1820]" class="Diplopoda" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="366" phylum="Arthropoda" rank="order">Chordeumatida</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C364D2ED50165076BA9F88AFB0DF8A8" box="[980,1202,1794,1820]" class="Diplopoda" family="Guizhousomatidae" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="366" phylum="Arthropoda" rank="family">Guizhousomatidae</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="EFA74B5CD50165076CB4F88AFEFBF8F0" author="Mauries" pageId="3" pageNumber="368" refString="Mauries, J. - P. (2005) Guizhousoma latellai gen. n., sp. n., de Chine continentale, type d'une nouvelle famille de la superfamille des Neoatractosomatoidea (Diplopoda: Chordeumatida). Arthropoda Selecta, 14 (1), 11 - 17." type="journal article" year="2005">Mauriès 2005</bibRefCitation>
|
||
) and a
|
||
<taxonomicName id="4C364D2ED501650769A2F8A3FDDFF8F0" box="[479,608,1835,1860]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="366" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B942EABFD501650769A2F8A3FDDFF8F0" box="[479,608,1835,1860]" italics="true" pageId="3" pageNumber="366">Glyphiulus</emphasis>
|
||
</taxonomicName>
|
||
species from the
|
||
<emphasis id="B942EABFD50165076BF7F8A3FBB6F8F0" box="[906,1033,1835,1860]" italics="true" pageId="3" pageNumber="366">granulatus</emphasis>
|
||
group (
|
||
<taxonomicName id="4C364D2ED50165076C00F8A2FA99F8F0" box="[1149,1318,1834,1860]" class="Diplopoda" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="366" phylum="Arthropoda" rank="order">Spirostreptida</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C364D2ED50165076975F8DAFE00F8D8" box="[264,447,1874,1900]" class="Diplopoda" family="Cambalopsidae" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="366" phylum="Arthropoda" rank="family">Cambalopsidae</taxonomicName>
|
||
) (Golovatch
|
||
<emphasis id="B942EABFD50165076A1AF8DBFD1EF8D8" box="[615,673,1875,1900]" italics="true" pageId="3" pageNumber="366">et al</emphasis>
|
||
., in press). On the other hand, the cave Hei Dong harbours a tiny species of
|
||
<taxonomicName id="4C364D2ED50665006A30FE94FC78FE82" authority="(Polydesmida)" baseAuthorityName="Polydesmida" box="[589,967,284,310]" class="Diplopoda" family="Doratodesmidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="367" phylum="Arthropoda" rank="family">Doratodesmidae (Polydesmida)</taxonomicName>
|
||
, most probably troglobitic as well. The distance between the two caves being only about 20 airkm, both lying within the same karst massif situated between Dafang and Qianxi, it is hardly surprising that they contain the same species of
|
||
<taxonomicName id="4C364D2ED50665006A1CFE1CFD52FE19" box="[609,749,404,429]" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="367" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B942EABFD50665006A1CFE1CFD52FE19" box="[609,749,404,429]" italics="true" pageId="4" pageNumber="367">Pacidesmus</emphasis>
|
||
</taxonomicName>
|
||
. In addition,
|
||
<taxonomicName id="4C364D2ED50665006BE9FE1CFB2CFE19" box="[916,1171,404,429]" class="Diplopoda" family="Guizhousomatidae" genus="Guizhousoma" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="latellai">
|
||
<emphasis id="B942EABFD50665006BE9FE1CFB2CFE19" box="[916,1171,404,429]" italics="true" pageId="4" pageNumber="367">Guizhousoma latellai</emphasis>
|
||
</taxonomicName>
|
||
has hitherto been encountered in as many as four of the massif’s caves (
|
||
<bibRefCitation id="EFA74B5CD50665006BC2FE34FBD7FE62" author="Mauries" box="[959,1128,444,470]" pageId="4" pageNumber="368" refString="Mauries, J. - P. (2005) Guizhousoma latellai gen. n., sp. n., de Chine continentale, type d'une nouvelle famille de la superfamille des Neoatractosomatoidea (Diplopoda: Chordeumatida). Arthropoda Selecta, 14 (1), 11 - 17." type="journal article" year="2005">Mauriès 2005</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF496625D50665006975F9ACFCA3F9D5" httpUri="https://zenodo.org/record/174084/files/figure.png" pageId="4" pageNumber="367" targetBox="[408,1192,513,1559]" targetPageId="4">
|
||
<paragraph id="8B8936ADD50665006975F9ACFCA3F9D5" blockId="4.[264,1323,1570,1633]" pageId="4" pageNumber="367">
|
||
<emphasis id="B942EABFD50665006975F9ACFE7FF988" bold="true" box="[264,448,1572,1596]" pageId="4" pageNumber="367">FIGURES 9–12</emphasis>
|
||
.
|
||
<taxonomicName id="4C364D2ED506650069AEF9ACFD03F98F" box="[467,700,1572,1595]" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
|
||
<emphasis id="B942EABFD506650069AEF9ACFD03F98F" box="[467,700,1572,1595]" italics="true" pageId="4" pageNumber="367">Pacidesmus martensi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27157C4D50665006AB5F9ACFCB6F988" box="[712,777,1572,1596]" pageId="4" pageNumber="367" rank="species">
|
||
<emphasis id="B942EABFD50665006AB5F9ACFCB6F988" bold="true" box="[712,777,1572,1596]" pageId="4" pageNumber="367">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, ɗ holotype. 9 Antenna; 10 Leg 9; 11–12 Left gonopod (11 mesal, 12 lateral). Scale bar 0.3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8936ADD50665006945F910FBD2F89E" blockId="4.[264,1323,1688,1834]" pageId="4" pageNumber="367">
|
||
The discovery of
|
||
<taxonomicName id="4C364D2ED50665006A7BF910FD35F905" box="[518,650,1688,1713]" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
|
||
<emphasis id="B942EABFD50665006A7BF910FD35F905" box="[518,650,1688,1713]" italics="true" pageId="4" pageNumber="367">P. martensi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27157C4D50665006AECF911FD72F905" box="[657,717,1689,1713]" pageId="4" pageNumber="367" rank="species">
|
||
<emphasis id="B942EABFD50665006AECF911FD72F905" bold="true" box="[657,717,1689,1713]" pageId="4" pageNumber="367">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in Guizhou — a species very closely related to and sympatric with
|
||
<taxonomicName id="4C364D2ED506650069B8F948FDFDF96D" box="[453,578,1728,1753]" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="sinensis">
|
||
<emphasis id="B942EABFD506650069B8F948FDFDF96D" box="[453,578,1728,1753]" italics="true" pageId="4" pageNumber="367">P. sinensis</emphasis>
|
||
</taxonomicName>
|
||
, despite the latter’s poor original description (
|
||
<bibRefCitation id="EFA74B5CD50665006C08F948FABCF96E" author="Loksa" box="[1141,1283,1728,1754]" pageId="4" pageNumber="368" refString="Loksa, I. (1960) Einige neue Diplopoden- und Chilopodenarten aus chinesischen Hohlen. Acta Zoologica Academiae Scientiarum Hungaricae, 6 (1 - 2), 137 - 148." type="journal article" year="1960">Loksa 1960</bibRefCitation>
|
||
)— leaves no doubt whatever that they are congeneric. Therefore, the doubts expressed by
|
||
<bibRefCitation id="EFA74B5CD50665006975F898FE67F89E" author="Golovatch" box="[264,472,1808,1834]" pageId="4" pageNumber="368" refString="Golovatch, S. I. (1991) The millipede family Polydesmidae in Southeast Asia, with notes on phylogeny (Diplopoda: Polydesmida). Steenstrupia, 17 (4), 141 - 159." type="journal article" year="1991">Golovatch (1991)</bibRefCitation>
|
||
concerning the allocation of
|
||
<taxonomicName id="4C364D2ED50665006B53F898FC17F89D" box="[814,936,1808,1833]" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="sinensis">
|
||
<emphasis id="B942EABFD50665006B53F898FC17F89D" box="[814,936,1808,1833]" italics="true" pageId="4" pageNumber="367">P. sinensis</emphasis>
|
||
</taxonomicName>
|
||
no longer apply.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |