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<document id="59CD9E282B79B4D9DEB10A012ED348D0" ID-CLB-Dataset="46816" ID-DOI="10.11646/zootaxa.4078.1.11" ID-GBIF-Dataset="4649cba5-233d-4010-b0f4-062b26ac299f" ID-ISSN="1175-5326" ID-Zenodo-Dep="263491" ID-ZooBank="584BBD6C-265A-4F23-9634-B441DCDE533D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460360667020" checkinUser="plazi" docAuthor="Souto, Paula Malaquias, Da-Silva, Elidiomar Ribeiro, Nessimian, Jorge Luiz &amp; Gonçalves, Inês Corrêa" docDate="2016" docId="039F879FC32EAA1AFF6257BE7FA8FF4D" docLanguage="en" docName="zootaxa.4078.1.11.pdf" docOrigin="Zootaxa 4078 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ulmeritoides" docType="key" docVersion="9" lastPageNumber="136" masterDocId="FFA6FFE7C326AA13FFF555687A33FFAA" masterDocTitle="Two new species of Ulmeritoides Traver (Ephemeroptera: Leptophlebiidae) from Southeastern Brazil" masterLastPageNumber="136" masterPageNumber="127" pageNumber="135" updateTime="1698294420992" updateUser="ExternalLinkService">
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<mods:title id="5B786AAC19FAFE70A667B5CE0CB1B18D">Two new species of Ulmeritoides Traver (Ephemeroptera: Leptophlebiidae) from Southeastern Brazil</mods:title>
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<mods:namePart id="9C7434C0DE147950573A98480C78B2C0">Souto, Paula Malaquias</mods:namePart>
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<mods:namePart id="E685E8B0C9FE04DB741A9DFEA2E807C1">Nessimian, Jorge Luiz</mods:namePart>
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<paragraph id="8B893689C32EAA1BFF6257BE7EECFD5A" blockId="8.[151,1247,726,752]" box="[151,1247,726,752]" pageId="8" pageNumber="135">
<heading id="D0C181E5C32EAA1BFF6257BE7EECFD5A" bold="true" box="[151,1247,726,752]" fontSize="11" level="1" pageId="8" pageNumber="135" reason="1">
<emphasis id="B942EA9BC32EAA1BFF6257BE7EECFD5A" bold="true" box="[151,1247,726,752]" pageId="8" pageNumber="135">
Key to the genera
<taxonomicName id="4C364D0AC32EAA1BFE8E57BE7825FD5A" ID-CoL="84MH" ID-ENA="1921401" box="[379,534,726,752]" class="Insecta" family="Leptophlebiidae" genus="Ulmeritoides" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B942EA9BC32EAA1BFE8E57BE7825FD5A" bold="true" box="[379,534,726,752]" italics="true" pageId="8" pageNumber="135">Ulmeritoides</emphasis>
</taxonomicName>
and
<taxonomicName id="4C364D0AC32EAA1BFDA657BE78F8FD5A" box="[595,715,726,752]" class="Insecta" family="Leptophlebiidae" genus="Ulmeritus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B942EA9BC32EAA1BFDA657BE78F8FD5A" bold="true" box="[595,715,726,752]" italics="true" pageId="8" pageNumber="135">Ulmeritus</emphasis>
</taxonomicName>
(modified from
<bibRefCitation id="EFA74B78C32EAA1BFC6257BE7EE4FD5A" author="Salles" box="[919,1239,726,752]" pageId="8" pageNumber="135" refString="Salles, F. F. &amp; Dominguez, E. (2012) Systematics and Phylogeny of Ulmeritus - Ulmeritoides revisited (Ephemeroptera: Leptophlebiidae). Zootaxa, 3571, 49 - 65." type="journal article" year="2012">Salles &amp; Domínguez 2012</bibRefCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8B893689C32EAA1BFF6256757F4FFCF0" blockId="8.[151,1436,797,858]" pageId="8" pageNumber="135">
The key to male imagos of
<taxonomicName id="4C364D0AC32EAA1BFE395675786DFC9C" box="[460,606,797,822]" class="Insecta" family="Leptophlebiidae" genus="Ulmeritoides" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B942EA9BC32EAA1BFE395675786DFC9C" box="[460,606,797,822]" italics="true" pageId="8" pageNumber="135">Ulmeritoides</emphasis>
</taxonomicName>
and
<taxonomicName id="4C364D0AC32EAA1BFD6056757936FC9C" box="[661,773,797,822]" class="Insecta" family="Leptophlebiidae" genus="Ulmeritus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B942EA9BC32EAA1BFD6056757936FC9C" box="[661,773,797,822]" italics="true" pageId="8" pageNumber="135">Ulmeritus</emphasis>
</taxonomicName>
presented by
<bibRefCitation id="EFA74B78C32EAA1BFC5656757EECFC9C" author="Salles" box="[931,1247,797,822]" pageId="8" pageNumber="135" refString="Salles, F. F. &amp; Dominguez, E. (2012) Systematics and Phylogeny of Ulmeritus - Ulmeritoides revisited (Ephemeroptera: Leptophlebiidae). Zootaxa, 3571, 49 - 65." type="journal article" year="2012">Salles &amp; Domínguez (2012)</bibRefCitation>
was modified to include the two new species here described and
<taxonomicName id="4C364D0AC32EAA1BFD4456297F3BFCF0" authority="Gama Neto &amp; Hamada, 2014" authorityName="Gama Neto &amp; Hamada" authorityYear="2014" box="[689,1288,833,858]" class="Insecta" family="Leptophlebiidae" genus="Ulmeritoides" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="135" phylum="Arthropoda" rank="species" species="passorum">
<emphasis id="B942EA9BC32EAA1BFD4456297984FCF3" box="[689,951,833,858]" italics="true" pageId="8" pageNumber="135">Ulmeritoides passorum</emphasis>
<bibRefCitation id="EFA74B78C32EAA1BFC4B562A7F3BFCF0" author="Gama" box="[958,1288,833,858]" pageId="8" pageNumber="135" refString="Gama Neto, J. L. &amp; Hamada, N. (2014) Leptophlebiidae (Ephemeroptera) of the Serra do Tepequem, Roraima State, Brazil: new records and description of two new species. Zootaxa, 3900 (2), 279 - 286. http: // dx. doi. org / 10.11646 / zootaxa. 3900.2.8" type="journal article" year="2014">Gama Neto &amp; Hamada, 2014</bibRefCitation>
</taxonomicName>
, as below.
</paragraph>
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<paragraph id="8B893689C32EAA1BFF6256E07FAFFC11" blockId="8.[151,1436,904,2011]" pageId="8" pageNumber="135">
<keyLead id="30C795BCC32EAA1BFF6256E07FAFFC11" pageId="8" pageNumber="135">
1. Vein Sc of hind wings 9/10 length of wing; membrane of wings with dark maculae at cross veins (fig.
<quantity id="4CCE9B6CC32EAA1BFB4E56E07ED2FC34" box="[1211,1249,904,926]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="135" unit="in" value="2.0">2 in</quantity>
<bibRefCitation id="EFA74B78C32EAA1BFB1256E07FA3FC34" author="Dominguez" box="[1255,1424,904,926]" pageId="8" pageNumber="135" refString="Dominguez, E. (1991) The status of the genus Ulmeritus (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) and related taxa. In: Alba-Tercedor, J. &amp; Sanchez-Ortega, A. Overview and strategies of Ephemeroptera and Plecoptera. Sandhill Crane Press, Gainesville, pp. 157 - 167." type="book chapter" year="1991">Domínguez 1991</bibRefCitation>
); apex of each penis round, lobe with ventral digitiform projection (fig.
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<bibRefCitation id="EFA74B78C32EAA1BFC5956CD7E60FC11" author="Dominguez" box="[940,1107,933,955]" pageId="8" pageNumber="135" refString="Dominguez, E. (1991) The status of the genus Ulmeritus (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) and related taxa. In: Alba-Tercedor, J. &amp; Sanchez-Ortega, A. Overview and strategies of Ephemeroptera and Plecoptera. Sandhill Crane Press, Gainesville, pp. 157 - 167." type="book chapter" year="1991">Domínguez 1991</bibRefCitation>
)................ 2
<taxonomicName id="4C364D0AC32EAA1BFAC956CD7FAFFC11" ID-CoL="84MJ" box="[1340,1436,933,955]" class="Insecta" family="Leptophlebiidae" genus="Ulmeritus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B942EA9BC32EAA1BFAC956CD7FAFFC11" box="[1340,1436,933,955]" italics="true" pageId="8" pageNumber="135">Ulmeritus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B893689C32EAA1BFF6256AB7FA8FC5C" blockId="8.[151,1436,904,2011]" pageId="8" pageNumber="135">
<keyLead id="30C795BCC32EAA1BFF6256AB7FA8FC5C" pageId="8" pageNumber="135">
1. Vein Sc of hind wings 8/10 or less length of wing (
<figureCitation id="130D2A0CC32EAA1BFD3356AB7937FC73" box="[710,772,963,985]" captionStart="FIGURES 5 9" captionStartId="2.[151,264,1912,1935]" captionTargetBox="[257,1327,603,1873]" captionTargetId="figure@2.[239,1347,590,1889]" captionTargetPageId="2" captionText="FIGURES 5 9. Ulmeritoides angelus sp. nov., male imago. 5. fore wing; 6. hind wing; 7. hind wing (enlarged); 8. genitalia (ventral view); 9. penis lobe (ventral view)." httpUri="https://zenodo.org/record/263493/files/figure.png" pageId="8" pageNumber="135">Figs 6</figureCitation>
,
<figureCitation id="130D2A0CC32EAA1BFCFB56AB791BFC73" box="[782,808,963,985]" captionStart="FIGURES 15 18" captionStartId="5.[151,264,1105,1128]" captionTargetBox="[151,1436,193,1084]" captionTargetId="figure@5.[151,1436,193,1085]" captionTargetPageId="5" captionText="FIGURES 15 18. Ulmeritoides angelus sp. nov., imago and subimago. 15. male subimago; 16. male imago; 17. female subimago; 18. female imago. Photos by Frederico F. Salles." httpUri="https://zenodo.org/record/263495/files/figure.png" pageId="8" pageNumber="135">17</figureCitation>
); membrane of wings without color pattern, except in costal area; penis lobes without digitiform projections (
<figureCitation id="130D2A0CC32EAA1BFD755688788EFC5C" box="[640,701,992,1014]" captionStart="FIGURES 5 9" captionStartId="2.[151,264,1912,1935]" captionTargetBox="[257,1327,603,1873]" captionTargetId="figure@2.[239,1347,590,1889]" captionTargetPageId="2" captionText="FIGURES 5 9. Ulmeritoides angelus sp. nov., male imago. 5. fore wing; 6. hind wing; 7. hind wing (enlarged); 8. genitalia (ventral view); 9. penis lobe (ventral view)." httpUri="https://zenodo.org/record/263493/files/figure.png" pageId="8" pageNumber="135">Figs 8</figureCitation>
,
<figureCitation id="130D2A0CC32EAA1BFD3D568878D2FC5C" box="[712,737,992,1014]" captionStart="FIGURES 19 21" captionStartId="7.[151,264,1659,1682]" captionTargetBox="[151,1436,193,1638]" captionTargetId="figure@7.[151,1436,193,1638]" captionTargetPageId="7" captionText="FIGURES 19 21. SEM photographs of eggs of Ulmeritoides species. 19. U. angelus sp. nov., general aspect; 20 21. U. angelus sp. nov., attachments structures (20) and chorionic sculpturing showing punctation and tubercules (21)." httpUri="https://zenodo.org/record/263496/files/figure.png" pageId="8" pageNumber="135">19</figureCitation>
)............................................. 4
<taxonomicName id="4C364D0AC32EAA1BFAEB56887FA8FC5C" ID-CoL="84MH" box="[1310,1435,992,1014]" class="Insecta" family="Leptophlebiidae" genus="Ulmeritoides" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B942EA9BC32EAA1BFAEB56887FA8FC5C" box="[1310,1435,992,1014]" italics="true" pageId="8" pageNumber="135">Ulmeritoides</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C22E2CC32EAA1BFF6256957FA8FB9B" pageId="8" pageNumber="135">
<paragraph id="8B893689C32EAA1BFF6256957FAFFBB9" blockId="8.[151,1436,904,2011]" box="[151,1436,1021,1043]" pageId="8" pageNumber="135">
<keyLead id="30C795BCC32EAA1BFF6256957FAFFBB9" box="[151,1436,1021,1043]" pageId="8" pageNumber="135">
2(1). Cross veins of hind wings few (around 20); maculae only around subcostal cross veins............
<taxonomicName id="4C364D0AC32EAA1BFB4F56957FAFFBB9" ID-CoL="7DFZD" box="[1210,1436,1021,1043]" class="Insecta" family="Leptophlebiidae" genus="Ulmeritus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="135" phylum="Arthropoda" rank="species" species="saopaulensis">
<emphasis id="B942EA9BC32EAA1BFB4F56957FAFFBB9" box="[1210,1436,1021,1043]" italics="true" pageId="8" pageNumber="135">Ulmeritus saopaulensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B893689C32EAA1BFF6251737FA8FB9B" blockId="8.[151,1436,904,2011]" box="[151,1435,1051,1073]" pageId="8" pageNumber="135">
<keyLead id="30C795BCC32EAA1BFF6251737FA8FB9B" box="[151,1435,1051,1073]" pageId="8" pageNumber="135">2. Cross veins of hind wings numerous (33 to 50); maculae around all cross veins..................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C22E2CC32EAA1BFF6251507FAFFB0C" pageId="8" pageNumber="135">
<paragraph id="8B893689C32EAA1BFF6251507FAFFBC1" blockId="8.[151,1436,904,2011]" pageId="8" pageNumber="135">
<keyLead id="30C795BCC32EAA1BFF6251507FAFFBC1" pageId="8" pageNumber="135">
3(2). Maculae of fore wings forming bands (fig. 199A in
<bibRefCitation id="EFA74B78C32EAA1BFD1D515079E3FBE4" author="Dominguez" box="[744,976,1080,1102]" pageId="8" pageNumber="135" refString="Dominguez, E., Molineri, C., Pescador, M. L., Hubbard, M. D. &amp; Nieto, C. (2006) Ephemeroptera of South America. Aquatic Biodiversity of Latin America. Pensoft, Sofia-Moscow, 646 pp." type="book" year="2006">
Domínguez
<emphasis id="B942EA9BC32EAA1BFC94515179BDFBE4" box="[865,910,1080,1102]" italics="true" pageId="8" pageNumber="135">et al</emphasis>
. 2006
</bibRefCitation>
); ventral prolongation of penes short, approxi- mately as long as wide (fig. 199C
<bibRefCitation id="EFA74B78C32EAA1BFDDC513D7935FBC1" author="Dominguez" box="[553,774,1109,1131]" pageId="8" pageNumber="135" refString="Dominguez, E., Molineri, C., Pescador, M. L., Hubbard, M. D. &amp; Nieto, C. (2006) Ephemeroptera of South America. Aquatic Biodiversity of Latin America. Pensoft, Sofia-Moscow, 646 pp." type="book" year="2006">
Domínguez
<emphasis id="B942EA9BC32EAA1BFD6A513E78E3FBC1" box="[671,720,1109,1131]" italics="true" pageId="8" pageNumber="135">et al.</emphasis>
2006
</bibRefCitation>
)......................................
<taxonomicName id="4C364D0AC32EAA1BFB28513D7FAFFBC1" ID-CoL="7DFZG" box="[1245,1436,1109,1131]" class="Insecta" family="Leptophlebiidae" genus="Ulmeritus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="135" phylum="Arthropoda" rank="species" species="balteatus">
<emphasis id="B942EA9BC32EAA1BFB28513D7FAFFBC1" box="[1245,1436,1109,1131]" italics="true" pageId="8" pageNumber="135">Ulmeritus balteatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B893689C32EAA1BFF62511B7FAFFB0C" blockId="8.[151,1436,904,2011]" pageId="8" pageNumber="135">
<keyLead id="30C795BCC32EAA1BFF62511B7FAFFB0C" pageId="8" pageNumber="135">
3. Maculae of fore wings not forming bands (fig. 199E in
<bibRefCitation id="EFA74B78C32EAA1BFD08511B79ECFB23" author="Dominguez" box="[765,991,1139,1161]" pageId="8" pageNumber="135" refString="Dominguez, E., Molineri, C., Pescador, M. L., Hubbard, M. D. &amp; Nieto, C. (2006) Ephemeroptera of South America. Aquatic Biodiversity of Latin America. Pensoft, Sofia-Moscow, 646 pp." type="book" year="2006">
Domínguez
<emphasis id="B942EA9BC32EAA1BFC81511C7996FB23" box="[884,933,1139,1161]" italics="true" pageId="8" pageNumber="135">et al.</emphasis>
2006
</bibRefCitation>
); ventral prolongation of penes long, approxi- mately twice as long as wide (fig. 199G in
<bibRefCitation id="EFA74B78C32EAA1BFD8851F87969FB0C" author="Dominguez" box="[637,858,1168,1190]" pageId="8" pageNumber="135" refString="Dominguez, E., Molineri, C., Pescador, M. L., Hubbard, M. D. &amp; Nieto, C. (2006) Ephemeroptera of South America. Aquatic Biodiversity of Latin America. Pensoft, Sofia-Moscow, 646 pp." type="book" year="2006">
Domínguez
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2006
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)..............................
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<emphasis id="B942EA9BC32EAA1BFB2651F87FAFFB0C" box="[1235,1436,1168,1190]" italics="true" pageId="8" pageNumber="135">Ulmeritus carbonelli</emphasis>
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<keyLead id="30C795BCC32EAA1BFF6251C57FA8FB69" box="[151,1435,1197,1219]" pageId="8" pageNumber="135">4(1). Fore wings with costal and subcostal areas tinged with brown.................................................. 5</keyLead>
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<keyLead id="30C795BCC32EAA1BFF6251A37FA8FB4B" box="[151,1435,1227,1249]" pageId="8" pageNumber="135">4. Fore wings with costal and subcostal areas hyaline.......................................................... 12</keyLead>
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5(4). Penis lobes with apex rounded (
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; see also figs
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<keyLead id="30C795BCC32EAA1BFF62506D7FAFFAB1" box="[151,1436,1285,1307]" pageId="8" pageNumber="135">
5. Penis lobes with apex straight (figs 8, 19 in
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) or slightly concave (
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)................ 9
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6(5). Penis lobes with a ventral keel (figs
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)............................................... 7
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<keyLead id="30C795BCC32EAA1BFF6250287FAFFAFC" box="[151,1436,1344,1366]" pageId="8" pageNumber="135">
6. Penis lobes without a ventral keel and with a central spine (
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)........................
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<emphasis id="B942EA9BC32EAA1BFB8E50287F7AFAFC" box="[1147,1353,1344,1366]" italics="true" pageId="8" pageNumber="135">Ulmeritoides angelus</emphasis>
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<emphasis id="B942EA9BC32EAA1BFABA50287FAFFAFC" bold="true" box="[1359,1436,1344,1366]" pageId="8" pageNumber="135">sp. nov.</emphasis>
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<keyLead id="30C795BCC32EAA1BFF6250367FA8FAD9" box="[151,1435,1373,1395]" pageId="8" pageNumber="135">
7 (6). Ventral keel of penis lobe located near apex (figs 9FG in
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)......................
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<emphasis id="B942EA9BC32EAA1BFB2650357FA8FAD9" box="[1235,1435,1373,1395]" italics="true" pageId="8" pageNumber="135">Ulmeritoides tifferae</emphasis>
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<keyLead id="30C795BCC32EAA1BFF6250137FA8FA3B" box="[151,1435,1403,1425]" pageId="8" pageNumber="135">
7. Ventral keel of penis lobe located on median length (figs.
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)............................ 8
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<keyLead id="30C795BCC32EAA1BFF6250F07FAFFA04" box="[151,1436,1432,1454]" pageId="8" pageNumber="135">
8(7). Abdomen translucent yellowish brown, lacking pattern of pale spots or dark bands...............
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<emphasis id="B942EA9BC32EAA1BFB5A50F07FAFFA04" box="[1199,1436,1432,1454]" italics="true" pageId="8" pageNumber="135">Ulmeritoides chavarriae</emphasis>
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<keyLead id="30C795BCC32EAA1BFF6250DD7FAFFA61" box="[151,1436,1461,1483]" pageId="8" pageNumber="135">
8. Abdominal terga orange-brown with pale basal spots and dark brown apical transverse spots...........
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<emphasis id="B942EA9BC32EAA1BFB1550DD7FAFFA61" box="[1248,1436,1461,1483]" italics="true" pageId="8" pageNumber="135">Ulmeritoides acosa</emphasis>
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9(5). Basal broad region of forceps almost 1/4 length of segment I (
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, also fig. 196N in
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Domínguez
<emphasis id="B942EA9BC32EAA1BFB5450BC7EE1FA43" box="[1185,1234,1491,1513]" italics="true" pageId="8" pageNumber="135">et al.</emphasis>
2006
</bibRefCitation>
)......... 10
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9. Basal broad region of forceps about 1/6 length of segment I (figs 7, 18 in
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).................. 11
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<keyLead id="30C795BCC32EAA1BFF6253657FAFF9EB" pageId="8" pageNumber="135">
10(9). Abdominal terga brown shaded with dark brown, with posterior margins darker and median line dark brown (
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).....................................................................................
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<emphasis id="B942EA9BC32EAA1BFB7653437F7AF9EB" box="[1155,1353,1579,1601]" italics="true" pageId="8" pageNumber="135">Ulmeritoides tamoio</emphasis>
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<emphasis id="B942EA9BC32EAA1BFABA53437FAFF9EB" bold="true" box="[1359,1436,1579,1601]" pageId="8" pageNumber="135">sp. nov.</emphasis>
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<keyLead id="30C795BCC32EAA1BFF6253207FAFF9D1" pageId="8" pageNumber="135">
10. Abdominal terga yellowish-brown, with posterior margins darker, median line absent (fig. 196P in
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Domínguez
<emphasis id="B942EA9BC32EAA1BFAFE53217F0FF9F4" box="[1291,1340,1608,1630]" italics="true" pageId="8" pageNumber="135">et al.</emphasis>
2006
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).......................................................................................
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<emphasis id="B942EA9BC32EAA1BFB21530D7FAFF9D1" box="[1236,1436,1637,1659]" italics="true" pageId="8" pageNumber="135">Ulmeritoides huitoto</emphasis>
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<keyLead id="30C795BCC32EAA1BFF6253EB7FAFF933" box="[151,1436,1667,1689]" pageId="8" pageNumber="135">
11(9). Penes with apex ending in an inner acute pointed projection (figs 8FG in
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).......
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<emphasis id="B942EA9BC32EAA1BFB5153EB7FAFF933" box="[1188,1436,1667,1689]" italics="true" pageId="8" pageNumber="135">Ulmeritoides misionensis</emphasis>
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<keyLead id="30C795BCC32EAA1BFF6253C87FAFF91C" box="[151,1436,1696,1718]" pageId="8" pageNumber="135">
11. Inner projection of penes rounded, outer corner with a small ventral spine (fig. 7C in
<bibRefCitation id="EFA74B78C32EAA1BFBE453C87E8BF91C" author="Dominguez" box="[1041,1208,1696,1718]" pageId="8" pageNumber="135" refString="Dominguez, E. (1995) Cladistic analysis of the Ulmeritus - Ulmeritoides group (Ephemeroptera, Leptophlebiidae), with descriptions of five new species of Ulmeritoides. Journal of the New York Entomological Society, 103, 15 - 38." type="journal article" year="1995">Domínguez 1995</bibRefCitation>
).
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<emphasis id="B942EA9BC32EAA1BFB3C53C87FAFF91C" box="[1225,1436,1696,1718]" italics="true" pageId="8" pageNumber="135">Ulmeritoides haarupi</emphasis>
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<keyLead id="30C795BCC32EAA1BFF6253D57FA8F979" box="[151,1435,1725,1747]" pageId="8" pageNumber="135">12(4). Apex of penis lobes rounded............................................................................ 13</keyLead>
</paragraph>
<paragraph id="8B893689C32EAA1BFF6253B37FA8F95B" blockId="8.[151,1436,904,2011]" box="[151,1435,1755,1777]" pageId="8" pageNumber="135">
<keyLead id="30C795BCC32EAA1BFF6253B37FA8F95B" box="[151,1435,1755,1777]" pageId="8" pageNumber="135">12. Apex of penis lobes not rounded......................................................................... 14</keyLead>
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<paragraph id="8B893689C32EAA1BFF6253907FA5F8A4" blockId="8.[151,1436,904,2011]" box="[151,1430,1784,1806]" pageId="8" pageNumber="135">
<keyLead id="30C795BCC32EAA1BFF6253907FA5F8A4" box="[151,1430,1784,1806]" pageId="8" pageNumber="135">
13(12). Each penis lobe cylindrical without a ventral keel (
<figureCitation id="130D2A0CC32EAA1BFD3F53907930F8A4" box="[714,771,1784,1806]" captionStart="FIGURES 1 4" captionStartId="1.[151,264,1893,1916]" captionTargetBox="[381,1203,231,1868]" captionTargetId="figure@1.[376,1211,231,1869]" captionTargetPageId="1" captionText="FIGURES 1 4. Male imagos of Ulmeritoides species. 1 2. U. angelus sp. nov. in dorsal (1) and lateral (2) views; 3 4. U. tamoio sp. nov. in dorsal (3) and lateral (4) views." httpUri="https://zenodo.org/record/263492/files/figure.png" pageId="8" pageNumber="135">Fig. 4</figureCitation>
F in
<bibRefCitation id="EFA74B78C32EAA1BFCDA539079E5F8A4" author="Dominguez" box="[815,982,1784,1806]" pageId="8" pageNumber="135" refString="Dominguez, E. (1995) Cladistic analysis of the Ulmeritus - Ulmeritoides group (Ephemeroptera, Leptophlebiidae), with descriptions of five new species of Ulmeritoides. Journal of the New York Entomological Society, 103, 15 - 38." type="journal article" year="1995">Domínguez 1995</bibRefCitation>
).................................. 17
</keyLead>
</paragraph>
<paragraph id="8B893689C32EAA1BFF62527D7FA8F881" blockId="8.[151,1436,904,2011]" box="[151,1435,1813,1835]" pageId="8" pageNumber="135">
<keyLead id="30C795BCC32EAA1BFF62527D7FA8F881" box="[151,1435,1813,1835]" pageId="8" pageNumber="135">
13. Each penis lobe flattened and with a ventral keel (
<figureCitation id="130D2A0CC32EAA1BFD35527D7937F881" box="[704,772,1813,1835]" captionStart="FIGURES 10 14" captionStartId="3.[151,264,1686,1709]" captionTargetBox="[263,1315,206,1650]" captionTargetId="figure@3.[254,1332,193,1665]" captionTargetPageId="3" captionText="FIGURES 10 14. Ulmeritoides tamoio sp. nov., male imago. 10. fore wing; 11. hind wing; 12. hind wing (enlarged); 13. genitalia (ventral view); 14. penis lobe (ventral view)." httpUri="https://zenodo.org/record/263494/files/figure.png" pageId="8" pageNumber="135">Fig. 12</figureCitation>
G in
<bibRefCitation id="EFA74B78C32EAA1BFCC1527D79E8F881" author="Dominguez" box="[820,987,1813,1835]" pageId="8" pageNumber="135" refString="Dominguez, E. (1995) Cladistic analysis of the Ulmeritus - Ulmeritoides group (Ephemeroptera, Leptophlebiidae), with descriptions of five new species of Ulmeritoides. Journal of the New York Entomological Society, 103, 15 - 38." type="journal article" year="1995">Domínguez 1995</bibRefCitation>
)................
<taxonomicName id="4C364D0AC32EAA1BFB5E527D7FA8F881" ID-CoL="7DG99" box="[1195,1435,1813,1835]" class="Insecta" family="Leptophlebiidae" genus="Ulmeritoides" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="135" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="B942EA9BC32EAA1BFB5E527D7FA8F881" box="[1195,1435,1813,1835]" italics="true" pageId="8" pageNumber="135">Ulmeritoides guanacaste</emphasis>
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</paragraph>
</keyStep>
<keyStep id="30C22E2CC32EAA1BFF62525B7FA8F8CC" pageId="8" pageNumber="135">
<paragraph id="8B893689C32EAA1BFF62525B7FA8F8E3" blockId="8.[151,1436,904,2011]" box="[151,1435,1843,1865]" pageId="8" pageNumber="135">
<keyLead id="30C795BCC32EAA1BFF62525B7FA8F8E3" box="[151,1435,1843,1865]" pageId="8" pageNumber="135">
14(12). Penes lobes with several small apical spines (
<figureCitation id="130D2A0CC32EAA1BFD6B525B78E5F8E3" box="[670,726,1843,1865]" captionStart="FIGURES 5 9" captionStartId="2.[151,264,1912,1935]" captionTargetBox="[257,1327,603,1873]" captionTargetId="figure@2.[239,1347,590,1889]" captionTargetPageId="2" captionText="FIGURES 5 9. Ulmeritoides angelus sp. nov., male imago. 5. fore wing; 6. hind wing; 7. hind wing (enlarged); 8. genitalia (ventral view); 9. penis lobe (ventral view)." httpUri="https://zenodo.org/record/263493/files/figure.png" pageId="8" pageNumber="135">Fig. 6</figureCitation>
D in
<bibRefCitation id="EFA74B78C32EAA1BFCF3525B799EF8E3" author="Dominguez" box="[774,941,1843,1865]" pageId="8" pageNumber="135" refString="Dominguez, E. (1995) Cladistic analysis of the Ulmeritus - Ulmeritoides group (Ephemeroptera, Leptophlebiidae), with descriptions of five new species of Ulmeritoides. Journal of the New York Entomological Society, 103, 15 - 38." type="journal article" year="1995">Domínguez 1995</bibRefCitation>
)...................
<taxonomicName id="4C364D0AC32EAA1BFB51525B7FA8F8E3" ID-CoL="7DG8V" box="[1188,1435,1843,1865]" class="Insecta" family="Leptophlebiidae" genus="Ulmeritoides" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="135" phylum="Arthropoda" rank="species" species="spinulipenis">
<emphasis id="B942EA9BC32EAA1BFB51525B7FA8F8E3" box="[1188,1435,1843,1865]" italics="true" pageId="8" pageNumber="135">Ulmeritoides spinulipenis</emphasis>
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</paragraph>
<paragraph id="8B893689C32EAA1BFF6252387FA8F8CC" blockId="8.[151,1436,904,2011]" box="[151,1435,1872,1894]" pageId="8" pageNumber="135">
<keyLead id="30C795BCC32EAA1BFF6252387FA8F8CC" box="[151,1435,1872,1894]" pageId="8" pageNumber="135">14. Penis lobes with inner apical margin ending in an acute projection.............................................. 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C22E2CC32EAA1BFF6252057FA8F80B" pageId="8" pageNumber="135">
<paragraph id="8B893689C32EAA1BFF6252057FA8F829" blockId="8.[151,1436,904,2011]" box="[151,1435,1901,1923]" pageId="8" pageNumber="135">
<keyLead id="30C795BCC32EAA1BFF6252057FA8F829" box="[151,1435,1901,1923]" pageId="8" pageNumber="135">
15(14). Abdominal color pattern with conspicuous lighter rounded marks..........................
<taxonomicName id="4C364D0AC32EAA1BFB6D52057FA8F829" ID-CoL="7DFZH" box="[1176,1435,1901,1923]" class="Insecta" family="Leptophlebiidae" genus="Ulmeritoides" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="135" phylum="Arthropoda" rank="species" species="uruguayensis">
<emphasis id="B942EA9BC32EAA1BFB6D52057FA8F829" box="[1176,1435,1901,1923]" italics="true" pageId="8" pageNumber="135">Ulmeritoides uruguayensis</emphasis>
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</paragraph>
<paragraph id="8B893689C32EAA1BFF6252E37FA8F80B" blockId="8.[151,1436,904,2011]" box="[151,1435,1931,1953]" pageId="8" pageNumber="135">
<keyLead id="30C795BCC32EAA1BFF6252E37FA8F80B" box="[151,1435,1931,1953]" pageId="8" pageNumber="135">15. Abdominal color pattern never as above................................................................... 16</keyLead>
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<keyStep id="30C22E2CC32EAA1BFF6252C07FA8F871" pageId="8" pageNumber="135">
<paragraph id="8B893689C32EAA1BFF6252C07FA8F814" blockId="8.[151,1436,904,2011]" box="[151,1435,1960,1982]" pageId="8" pageNumber="135">
<keyLead id="30C795BCC32EAA1BFF6252C07FA8F814" box="[151,1435,1960,1982]" pageId="8" pageNumber="135">
16(15). Bulla not surrounded with black mark (fig. 5a in
<bibRefCitation id="EFA74B78C32EAA1BFD3552C079F3F814" author="Salles" box="[704,960,1960,1982]" pageId="8" pageNumber="135" refString="Salles, F. F. &amp; Dominguez, E. (2012) Systematics and Phylogeny of Ulmeritus - Ulmeritoides revisited (Ephemeroptera: Leptophlebiidae). Zootaxa, 3571, 49 - 65." type="journal article" year="2012">Salles &amp; Domínguez 2012</bibRefCitation>
)....................
<taxonomicName id="4C364D0AC32EAA1BFB4E52C07FA8F814" ID-CoL="7DFWX" box="[1211,1435,1960,1982]" class="Insecta" family="Leptophlebiidae" genus="Ulmeritoides" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="135" phylum="Arthropoda" rank="species" species="araponga">
<emphasis id="B942EA9BC32EAA1BFB4E52C07FA8F814" box="[1211,1435,1960,1982]" italics="true" pageId="8" pageNumber="135">Ulmeritoides araponga</emphasis>
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</paragraph>
<paragraph id="8B893689C32EAA1BFF6252AD7FA8F871" blockId="8.[151,1436,904,2011]" box="[151,1435,1989,2011]" pageId="8" pageNumber="135">
<keyLead id="30C795BCC32EAA1BFF6252AD7FA8F871" box="[151,1435,1989,2011]" pageId="8" pageNumber="135">
16. Bulla surrounded with black mark (fig. 16a in
<bibRefCitation id="EFA74B78C32EAA1BFD6B52AD79ADF871" author="Salles" box="[670,926,1989,2011]" pageId="8" pageNumber="135" refString="Salles, F. F. &amp; Dominguez, E. (2012) Systematics and Phylogeny of Ulmeritus - Ulmeritoides revisited (Ephemeroptera: Leptophlebiidae). Zootaxa, 3571, 49 - 65." type="journal article" year="2012">Salles &amp; Domínguez 2012</bibRefCitation>
)......................
<taxonomicName id="4C364D0AC32EAA1BFB5B52AD7FA8F871" ID-CoL="7DFWV" box="[1198,1435,1989,2011]" class="Insecta" family="Leptophlebiidae" genus="Ulmeritoides" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="135" phylum="Arthropoda" rank="species" species="nigribullae">
<emphasis id="B942EA9BC32EAA1BFB5B52AD7FA8F871" box="[1198,1435,1989,2011]" italics="true" pageId="8" pageNumber="135">Ulmeritoides nigribullae</emphasis>
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<keyStep id="30C22E2CC32FAA1AFF6255FF7FA8FF4D" pageId="9" pageNumber="136">
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<keyLead id="30C795BCC32FAA1AFF6255FF7FA8FF07" box="[151,1435,151,173]" pageId="9" pageNumber="136">
17(13). Each penis lobe with a lateral groove (fig. 4F in
<bibRefCitation id="EFA74B78C32FAA1AFD4755FF796AFF07" author="Dominguez" box="[690,857,151,173]" pageId="9" pageNumber="136" refString="Dominguez, E. (1995) Cladistic analysis of the Ulmeritus - Ulmeritoides group (Ephemeroptera, Leptophlebiidae), with descriptions of five new species of Ulmeritoides. Journal of the New York Entomological Society, 103, 15 - 38." type="journal article" year="1995">Domínguez 1995</bibRefCitation>
)...........................
<taxonomicName id="4C364D0AC32FAA1AFB4455FF7FA8FF07" ID-CoL="7DG8X" box="[1201,1435,151,173]" class="Insecta" family="Leptophlebiidae" genus="Ulmeritoides" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="136" phylum="Arthropoda" rank="species" species="flavopedes">
<emphasis id="B942EA9BC32FAA1AFB4455FF7FA8FF07" box="[1201,1435,151,173]" italics="true" pageId="9" pageNumber="136">Ulmeritoides flavopedes</emphasis>
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<keyLead id="30C795BCC32FAA1AFF6255DC7FA8FF4D" pageId="9" pageNumber="136">
17. Penes lobes without lateral groove, apex of posterior scutal protuberance with a yellowish transversal line (fig.
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)......................................................................
<taxonomicName id="4C364D0AC32FAA1AFB4E55B97FA8FF4D" box="[1211,1435,209,231]" class="Insecta" family="Leptophlebiidae" genus="Ulmeritoides" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="136" phylum="Arthropoda" rank="species" species="passorum">
<emphasis id="B942EA9BC32FAA1AFB4E55B97FA8FF4D" box="[1211,1435,209,231]" italics="true" pageId="9" pageNumber="136">Ulmeritoides passorum</emphasis>
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