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<document id="BBF180A695809C0B063CF8AA70DBDC44" ID-CLB-Dataset="32201" ID-DOI="10.11646/zootaxa.4317.3.4" ID-GBIF-Dataset="ca0d653d-32de-4c75-a95d-d9a4690d6fda" ID-ISSN="1175-5326" ID-Zenodo-Dep="884766" ID-ZooBank="5Fd492A4-D41D-4F37-A121-Ffa680E7E778" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1504596385874" checkinUser="plazi" docAuthor="Fery, Hans" docDate="2017" docId="039C87CE0F30FF97FF1F64F8CB3550C8" docLanguage="en" docName="zootaxa.4317.3.4.pdf" docOrigin="Zootaxa 4317 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coelambus Thomson 1860" docType="treatment" docVersion="7" lastPageNumber="521" masterDocId="FFA5FFB60F23FF81FF886361C8305140" masterDocTitle="A new classification of the tribe Hygrotini Portevin, 1929 (Coleoptera: Dytiscidae: Hydroporinae)" masterLastPageNumber="529" masterPageNumber="499" pageNumber="518" updateTime="1698467863718" updateUser="ExternalLinkService">
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<mods:title id="8735D7EC6BBCC8773F6BB4F95B3D6FD5">A new classification of the tribe Hygrotini Portevin, 1929 (Coleoptera: Dytiscidae: Hydroporinae)</mods:title>
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<treatment id="039C87CE0F30FF97FF1F64F8CB3550C8" ID-DOI="http://doi.org/10.5281/zenodo.6026853" ID-GBIF-Taxon="133183477" ID-Zenodo-Dep="6026853" LSID="urn:lsid:plazi:treatment:039C87CE0F30FF97FF1F64F8CB3550C8" httpUri="http://treatment.plazi.org/id/039C87CE0F30FF97FF1F64F8CB3550C8" lastPageId="22" lastPageNumber="521" pageId="19" pageNumber="518">
<subSubSection id="C32F65530F30FF92FF1F64F8CA5456F3" box="[151,612,1944,1971]" pageId="19" pageNumber="518" type="nomenclature">
<paragraph id="8B8A36D80F30FF92FF1F64F8CA5456F3" blockId="19.[151,612,1944,1971]" box="[151,612,1944,1971]" pageId="19" pageNumber="518">
<heading id="D0C281B40F30FF92FF1F64F8CA5456F3" bold="true" box="[151,612,1944,1971]" fontSize="11" level="1" pageId="19" pageNumber="518" reason="1">
<emphasis id="B941EACA0F30FF92FF1F64F8CA5456F3" bold="true" box="[151,612,1944,1971]" pageId="19" pageNumber="518">
Subgenus
<taxonomicName id="4C354D5B0F30FF92FE9F64F9CA5456F3" authority="Thomson, 1860" authorityName="Thomson" authorityYear="1860" box="[279,612,1944,1971]" class="Insecta" family="Dytiscidae" genus="Coelambus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="518" phylum="Arthropoda" rank="genus">
<emphasis id="B941EACA0F30FF92FE9F64F9C9AD56F2" bold="true" box="[279,413,1944,1970]" italics="true" pageId="19" pageNumber="518">Coelambus</emphasis>
<bibRefCitation id="EFA44B290F30FF92FE2D64F8CA5456F3" author="Thomson" box="[421,612,1945,1971]" pageId="19" pageNumber="518" refString="Thomson, C. G. (1860) Skandinaviens Coleoptera, synoptiskt bearbetade. Vol. II. Lundbergska Boktryckeriet, Lund, 304 pp." type="book" year="1860">Thomson, 1860</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32F65530F30FF94FF1F6481C8D95717" lastPageId="21" lastPageNumber="520" pageId="19" pageNumber="518" type="materials_examined">
<paragraph id="8B8A36D80F30FF92FF1F6481CCF956B6" blockId="19.[151,1225,2016,2038]" box="[151,1225,2016,2038]" pageId="19" pageNumber="518">
<emphasis id="B941EACA0F30FF92FF1F6481C92C56B6" bold="true" box="[151,284,2016,2038]" pageId="19" pageNumber="518">Type species</emphasis>
:
<taxonomicName id="4C354D5B0F30FF92FEA26480CAF256B6" authority="Fabricius, 1787: 193" authorityName="Fabricius" authorityPageNumber="193" authorityYear="1787" box="[298,706,2016,2038]" class="Insecta" family="Dytiscidae" genus="Dytiscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="518" phylum="Arthropoda" rank="species" species="confluens">
<emphasis id="B941EACA0F30FF92FEA26480C9D856B6" box="[298,488,2016,2038]" italics="true" pageId="19" pageNumber="518">Dytiscus confluens</emphasis>
<bibRefCitation id="EFA44B290F30FF92FE676481CAA056B6" author="Fabricius" box="[495,656,2016,2038]" pageId="19" pageNumber="518" refString="Fabricius, J. C. (1787) Mantissa Insectorum sistens species nuper detectas adiectis characteribus genericis, differentiis specificis, emendationibus, observationibus. Vol. 1 &amp; Vol. 2. C. G. Proft, Hafniae, xx + 348 + 382 pp." type="book" year="1787">Fabricius, 1787</bibRefCitation>
: 193
</taxonomicName>
, by subsequent designation of
<bibRefCitation id="EFA44B290F30FF92FC716481CCF556B6" author="Zaitzev" box="[1017,1221,2016,2038]" pageId="19" pageNumber="518" refString="Zaitzev, F. A. [Ph.] (1953) Nasekomye zhestkokrylye. Plavuntsovye i vertyachki. Fauna SSSR, New Series, 4 (58), 1 - 376. [in Russian]" type="journal article" year="1953">Zaitzev (1953: 129)</bibRefCitation>
.
</paragraph>
<caption id="DF4A66500F37FF95FF1F6419C9F256AA" httpUri="https://zenodo.org/record/884784/files/figure.png" pageId="20" pageNumber="519" targetBox="[151,1436,193,1890]" targetPageId="20">
<paragraph id="8B8A36D80F37FF95FF1F6419C9F256AA" blockId="20.[151,1436,1912,2026]" pageId="20" pageNumber="519">
<emphasis id="B941EACA0F37FF95FF1F6419C96456CE" bold="true" box="[151,340,1912,1934]" pageId="20" pageNumber="519">FIGURES 41–48.</emphasis>
Head with details of clypeal rim of (41)
<taxonomicName id="4C354D5B0F37FF95FD7D6418CC8156CE" baseAuthorityName="Schaller" baseAuthorityYear="1783" box="[757,1201,1912,1934]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="519" phylum="Arthropoda" rank="species" species="impressopunctatus" subGenus="Leptolambus">
<emphasis id="B941EACA0F37FF95FD7D6418CB6256CE" box="[757,850,1913,1934]" italics="true" pageId="20" pageNumber="519">Hygrotus</emphasis>
(
<emphasis id="B941EACA0F37FF95FCEA6419CBD556CE" box="[866,997,1912,1934]" italics="true" pageId="20" pageNumber="519">Leptolambus</emphasis>
)
<emphasis id="B941EACA0F37FF95FC7C6418CC8156CE" box="[1012,1201,1913,1934]" italics="true" pageId="20" pageNumber="519">impressopunctatus</emphasis>
</taxonomicName>
, (42)
<taxonomicName id="4C354D5B0F37FF95FB666418C8E856EC" authorityName="Aube" authorityYear="1838" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="519" phylum="Arthropoda" rank="species" species="solieri" subGenus="Hyphoporus">
<emphasis id="B941EACA0F37FF95FB666418CD3056CE" box="[1262,1280,1913,1934]" italics="true" pageId="20" pageNumber="519">H</emphasis>
. (
<emphasis id="B941EACA0F37FF95FA9E6419CDA356CE" box="[1302,1427,1912,1934]" italics="true" pageId="20" pageNumber="519">Hyphoporus</emphasis>
)
<emphasis id="B941EACA0F37FF95FF1F64F7C8E856EC" box="[151,216,1942,1964]" italics="true" pageId="20" pageNumber="519">solieri</emphasis>
</taxonomicName>
, (43)
<taxonomicName id="4C354D5B0F37FF95FE9064F9C9C456ED" authorityName="Aube" authorityYear="1838" box="[280,500,1942,1965]" class="Insecta" family="Dytiscidae" genus="Hyphydrus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="519" phylum="Arthropoda" rank="species" species="guineensis">
<emphasis id="B941EACA0F37FF95FE9064F9C91A56ED" box="[280,298,1944,1965]" italics="true" pageId="20" pageNumber="519">H</emphasis>
. (s. str.)
<emphasis id="B941EACA0F37FF95FE0164F9C9C456ED" box="[393,500,1944,1965]" italics="true" pageId="20" pageNumber="519">guineensis</emphasis>
</taxonomicName>
, (44)
<taxonomicName id="4C354D5B0F37FF95FDBD64F9CB2056EC" authorityName="Fabricius" authorityYear="1777" box="[565,784,1942,1965]" class="Insecta" family="Dytiscidae" genus="Dytiscus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="519" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B941EACA0F37FF95FDBD64F9CA7756ED" box="[565,583,1944,1965]" italics="true" pageId="20" pageNumber="519">H</emphasis>
. (s. str.)
<emphasis id="B941EACA0F37FF95FD2E64F7CB2056EC" box="[678,784,1942,1964]" italics="true" pageId="20" pageNumber="519">inaequalis</emphasis>
</taxonomicName>
, (45) idem in lateral view, (46)
<taxonomicName id="4C354D5B0F37FF95FBD764F9CD5C56EC" authorityName="Fabricius" authorityYear="1787" box="[1119,1388,1942,1965]" class="Insecta" family="Dytiscidae" genus="Dytiscus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="519" phylum="Arthropoda" rank="species" species="confluens" subGenus="Coelambus">
<emphasis id="B941EACA0F37FF95FBD764F9CC4156ED" box="[1119,1137,1944,1965]" italics="true" pageId="20" pageNumber="519">H</emphasis>
. (
<emphasis id="B941EACA0F37FF95FB0164F7CCCB56EC" box="[1161,1275,1942,1964]" italics="true" pageId="20" pageNumber="519">Coelambus</emphasis>
)
<emphasis id="B941EACA0F37FF95FA8364F7CD5C56EC" box="[1291,1388,1942,1964]" italics="true" pageId="20" pageNumber="519">confluens</emphasis>
</taxonomicName>
(the lighter area before the anterior border of the clypeus is the &quot;clypeal stripe&quot;; see Appendix), (47)
<taxonomicName id="4C354D5B0F37FF95FBF864D7CDAC568B" baseAuthorityName="Crotch" baseAuthorityYear="1874" box="[1136,1436,1973,1995]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="519" phylum="Arthropoda" rank="species" species="masculinus" subGenus="Leptolambus">
<emphasis id="B941EACA0F37FF95FBF864D7CCB2568B" box="[1136,1154,1974,1995]" italics="true" pageId="20" pageNumber="519">H</emphasis>
. (
<emphasis id="B941EACA0F37FF95FB1064D4CD2B568B" box="[1176,1307,1973,1995]" italics="true" pageId="20" pageNumber="519">Leptolambus</emphasis>
)
<emphasis id="B941EACA0F37FF95FAA264D4CDAC568B" box="[1322,1436,1973,1995]" italics="true" pageId="20" pageNumber="519">masculinus</emphasis>
</taxonomicName>
and (48) idem in lateral view.
</paragraph>
</caption>
<caption id="DF4A66500F36FF94FF1F6620CC4954B0" httpUri="https://zenodo.org/record/884786/files/figure.png" pageId="21" pageNumber="520" targetBox="[191,1436,208,1316]" targetPageId="21">
<paragraph id="8B8A36D80F36FF94FF1F6620CC4954B0" blockId="21.[151,1436,1345,1520]" pageId="21" pageNumber="520">
<emphasis id="B941EACA0F36FF94FF1F6620C9855417" bold="true" box="[151,437,1345,1367]" pageId="21" pageNumber="520">FIGURES 49–55. (49–53):</emphasis>
Epipleuron with oblique epipleural carina and genicular fossa of: (49)
<taxonomicName id="4C354D5B0F36FF94FB1C6620CDA65417" authorityName="Young &amp; Wolfe" authorityYear="1984" box="[1172,1430,1345,1367]" class="Insecta" family="Dytiscidae" genus="Clemnius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="520" phylum="Arthropoda" rank="species" species="berneri">
<emphasis id="B941EACA0F36FF94FB1C6620CCC35417" box="[1172,1267,1345,1367]" italics="true" pageId="21" pageNumber="520">Clemnius</emphasis>
(s. str.)
<emphasis id="B941EACA0F36FF94FAC36620CDA65417" box="[1355,1430,1345,1367]" italics="true" pageId="21" pageNumber="520">berneri</emphasis>
</taxonomicName>
, (50)
<taxonomicName id="4C354D5B0F36FF94FF4F6600CABF5435" authority="polonicus" authorityName="polonicus (Aube" authorityYear="1842" box="[199,655,1375,1398]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="520" phylum="Arthropoda" rank="subSpecies" species="polonicus" subGenus="Leptolambus" subSpecies="polonicus">
<emphasis id="B941EACA0F36FF94FF4F6600C9145436" box="[199,292,1377,1398]" italics="true" pageId="21" pageNumber="520">Hygrotus</emphasis>
(
<emphasis id="B941EACA0F36FF94FEBB663EC9865435" box="[307,438,1375,1397]" italics="true" pageId="21" pageNumber="520">Leptolambus</emphasis>
)
<emphasis id="B941EACA0F36FF94FE4C663ECA175435" box="[452,551,1375,1397]" italics="true" pageId="21" pageNumber="520">polonicus</emphasis>
<emphasis id="B941EACA0F36FF94FDA5663ECABF5435" box="[557,655,1375,1397]" italics="true" pageId="21" pageNumber="520">polonicus</emphasis>
</taxonomicName>
, (51)
<taxonomicName id="4C354D5B0F36FF94FD446600CB8F5436" authorityName="" baseAuthorityName="Wehncke" baseAuthorityYear="1875" box="[716,959,1375,1398]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="520" phylum="Arthropoda" rank="species" species="caspius" subGenus="Coelambus">
<emphasis id="B941EACA0F36FF94FD446600CAEE5436" box="[716,734,1377,1398]" italics="true" pageId="21" pageNumber="520">H</emphasis>
. (
<emphasis id="B941EACA0F36FF94FD7B663ECB555435" box="[755,869,1375,1397]" italics="true" pageId="21" pageNumber="520">Coelambus</emphasis>
)
<emphasis id="B941EACA0F36FF94FCFB6600CB8F5436" box="[883,959,1377,1398]" italics="true" pageId="21" pageNumber="520">caspius</emphasis>
</taxonomicName>
, (52)
<taxonomicName id="4C354D5B0F36FF94FC746600CD415436" baseAuthorityName="Schaller" baseAuthorityYear="1783" box="[1020,1393,1375,1398]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="520" phylum="Arthropoda" rank="species" species="impressopunctatus" subGenus="Leptolambus">
<emphasis id="B941EACA0F36FF94FC746600CC255436" box="[1020,1045,1377,1398]" italics="true" pageId="21" pageNumber="520">H.</emphasis>
(
<emphasis id="B941EACA0F36FF94FBAB663ECC965435" box="[1059,1190,1375,1397]" italics="true" pageId="21" pageNumber="520">Leptolambus</emphasis>
)
<emphasis id="B941EACA0F36FF94FB3C6600CD415436" box="[1204,1393,1377,1398]" italics="true" pageId="21" pageNumber="520">impressopunctatus</emphasis>
</taxonomicName>
and (53)
<taxonomicName id="4C354D5B0F36FF94FF41661FC9BD54D4" authorityName="Foster" authorityYear="1992" box="[201,397,1406,1428]" class="Insecta" family="Dytiscidae" genus="Rhithrodytes" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="520" phylum="Arthropoda" rank="species" species="agnus">
<emphasis id="B941EACA0F36FF94FF41661FC9BD54D4" box="[201,397,1406,1428]" italics="true" pageId="21" pageNumber="520">Rhithrodytes agnus</emphasis>
</taxonomicName>
; epipleural carina (c) and inner margin of epipleuron (m) including angle mentioned in diagnoses of subgenera.
<emphasis id="B941EACA0F36FF94FE8366FDC95454F2" bold="true" box="[267,356,1436,1458]" pageId="21" pageNumber="520">(54–55):</emphasis>
Elytral margin in lateral view of: (54)
<taxonomicName id="4C354D5B0F36FF94FD7E66FFCBDC54F3" authorityName="" baseAuthorityName="Wehncke" baseAuthorityYear="1875" box="[758,1004,1436,1459]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="520" phylum="Arthropoda" rank="species" species="caspius" subGenus="Coelambus">
<emphasis id="B941EACA0F36FF94FD7E66FFCB3854F3" box="[758,776,1438,1459]" italics="true" pageId="21" pageNumber="520">H</emphasis>
. (
<emphasis id="B941EACA0F36FF94FC9766FDCBA154F2" box="[799,913,1436,1458]" italics="true" pageId="21" pageNumber="520">Coelambus</emphasis>
)
<emphasis id="B941EACA0F36FF94FC2866FFCBDC54F3" box="[928,1004,1438,1459]" italics="true" pageId="21" pageNumber="520">caspius</emphasis>
</taxonomicName>
(strongly ascending to shoulder) and (55)
<taxonomicName id="4C354D5B0F36FF94FF1F66DAC9A85491" authorityName="Young &amp; Wolfe" authorityYear="1984" box="[151,408,1467,1489]" class="Insecta" family="Dytiscidae" genus="Clemnius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="520" phylum="Arthropoda" rank="species" species="berneri">
<emphasis id="B941EACA0F36FF94FF1F66DAC8C75491" box="[151,247,1467,1489]" italics="true" pageId="21" pageNumber="520">Clemnius</emphasis>
(s. str.)
<emphasis id="B941EACA0F36FF94FEC566DAC9A85491" box="[333,408,1467,1489]" italics="true" pageId="21" pageNumber="520">berneri</emphasis>
</taxonomicName>
(moderately ascending to shoulder) (Figs 52 and 53 are reproduced from Fery 2013 and from Fery 2016 with the permissions of F. Gusenleitner, Linz, Austria, and G. Foster, Ayr, UK, respectively).
</paragraph>
</caption>
<paragraph id="8B8A36D80F36FF94FF1F657BC8D95717" blockId="21.[151,1437,1562,2019]" pageId="21" pageNumber="520">
The subgenus
<taxonomicName id="4C354D5B0F36FF94FEB3657BC9895773" authorityName="Thomson" authorityYear="1860" box="[315,441,1562,1587]" class="Insecta" family="Dytiscidae" genus="Coelambus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="520" phylum="Arthropoda" rank="genus">
<emphasis id="B941EACA0F36FF94FEB3657BC9895773" box="[315,441,1562,1587]" italics="true" pageId="21" pageNumber="520">Coelambus</emphasis>
</taxonomicName>
contains eight Palaearctic and one Nearctic species (
<taxonomicName id="4C354D5B0F36FF94FB9D657AC8D55717" authority="(Fall, 1919))" authorityName="" authorityYear="1919" baseAuthorityName="Fall" baseAuthorityYear="1919" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="520" phylum="Arthropoda" rank="species" species="punctilineatus" subGenus="Coelambus">
<emphasis id="B941EACA0F36FF94FB9D657ACC4C5772" box="[1045,1148,1563,1586]" italics="true" pageId="21" pageNumber="520">Hygrotus</emphasis>
(
<emphasis id="B941EACA0F36FF94FB06657ACC975772" box="[1166,1191,1563,1586]" italics="true" pageId="21" pageNumber="520">C.</emphasis>
)
<emphasis id="B941EACA0F36FF94FB31657BCD685773" box="[1209,1368,1562,1587]" italics="true" pageId="21" pageNumber="520">punctilineatus</emphasis>
<bibRefCitation id="EFA44B290F36FF94FAE8657BC8D55717" author="Fall" pageId="21" pageNumber="520" refString="Fall, H. C. (1919) The North American species of Coelambus. Privately printed, Mount Vernon, New York, 20 pp." type="book" year="1919">(Fall, 1919))</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C32F65530F36FF97FF4F6503C91851B8" lastPageId="22" lastPageNumber="521" pageId="21" pageNumber="520" type="diagnosis">
<paragraph id="8B8A36D80F36FF94FF4F6503CC8A56DB" blockId="21.[151,1437,1562,2019]" pageId="21" pageNumber="520">
<emphasis id="B941EACA0F36FF94FF4F6503C971573B" bold="true" box="[199,321,1634,1659]" pageId="21" pageNumber="520">Diagnosis.</emphasis>
Body shape moderately to elongate oval (TL/MW ca. 1.7–2.0); small to medium sized species (TL 3.0–
<quantity id="4CCD9B3D0F36FF94FF4065E7C91257DF" box="[200,290,1670,1695]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="21" pageNumber="520" unit="mm" value="4.5">4.5 mm</quantity>
) (see
<figureCitation id="130E2A5D0F36FF94FEE065E7C99C57DF" box="[360,428,1670,1695]" captionStart="FIGURES 7 – 12" captionStartId="15.[151,264,1834,1856]" captionTargetBox="[155,1428,213,1778]" captionTargetId="figure@15.[151,1436,193,1813]" captionTargetPageId="15" captionText="FIGURES 7 – 12. Habitus of: (7) Clemnius (Cyclopius) acaroides, (8) Hygrotus (s. str.) inaequalis, (9) Hygrotus (Coelambus) confluens, (10) Clemnius. (s. str.) decoratus, (11) Clemnius (s. str.) laccophilinus and (12) Clemnius (s. str.) berneri (male holotype and labels)." httpUri="https://zenodo.org/record/884776/files/figure.png" pageId="21" pageNumber="520">Fig. 9</figureCitation>
for
<taxonomicName id="4C354D5B0F36FF94FE5765E6CB3857DF" authorityName="Fabricius" authorityYear="1787" box="[479,776,1670,1695]" class="Insecta" family="Dytiscidae" genus="Dytiscus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="520" phylum="Arthropoda" rank="species" species="confluens" subGenus="Coelambus">
<emphasis id="B941EACA0F36FF94FE5765E6C9CB57DE" box="[479,507,1671,1694]" italics="true" pageId="21" pageNumber="520">H.</emphasis>
(
<emphasis id="B941EACA0F36FF94FD8465E7CABA57DF" box="[524,650,1670,1695]" italics="true" pageId="21" pageNumber="520">Coelambus</emphasis>
)
<emphasis id="B941EACA0F36FF94FD1565E7CB3857DF" box="[669,776,1670,1695]" italics="true" pageId="21" pageNumber="520">confluens</emphasis>
</taxonomicName>
). Head with anterior clypeal margin truncate and slightly emarginated, border not produced forwards; bead absent. Antennomeres simple, not broadened. Elytra with margin in lateral view strongly ascending to shoulder (see
<figureCitation id="130E2A5D0F36FF94FC8F65AFCB6E57A7" box="[775,862,1742,1767]" captionStart="FIGURES 49 – 55" captionStartId="21.[151,264,1345,1367]" captionTargetBox="[191,1436,208,1316]" captionTargetId="figure@21.[151,1436,193,1323]" captionTargetPageId="21" captionText="FIGURES 49 – 55. (49 – 53): Epipleuron with oblique epipleural carina and genicular fossa of: (49) Clemnius (s. str.) berneri, (50) Hygrotus (Leptolambus) polonicus polonicus, (51) H. (Coelambus) caspius, (52) H. (Leptolambus) impressopunctatus and (53) Rhithrodytes agnus; epipleural carina (c) and inner margin of epipleuron (m) including angle mentioned in diagnoses of subgenera. (54 – 55): Elytral margin in lateral view of: (54) H. (Coelambus) caspius (strongly ascending to shoulder) and (55) Clemnius (s. str.) berneri (moderately ascending to shoulder) (Figs 52 and 53 are reproduced from Fery 2013 and from Fery 2016 with the permissions of F. Gusenleitner, Linz, Austria, and G. Foster, Ayr, UK, respectively)." httpUri="https://zenodo.org/record/884786/files/figure.png" pageId="21" pageNumber="520">Fig. 54</figureCitation>
for
<taxonomicName id="4C354D5B0F36FF94FC1265AECDA757A7" authority="(Wehncke, 1875))" authorityName="" authorityYear="1875" baseAuthorityName="Wehncke" baseAuthorityYear="1875" box="[922,1431,1742,1767]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="520" phylum="Arthropoda" rank="species" species="caspius" subGenus="Coelambus">
<emphasis id="B941EACA0F36FF94FC1265AECB9E57A6" box="[922,942,1743,1766]" italics="true" pageId="21" pageNumber="520">H</emphasis>
. (
<emphasis id="B941EACA0F36FF94FC4465AFCC7A57A7" box="[972,1098,1742,1767]" italics="true" pageId="21" pageNumber="520">Coelambus</emphasis>
)
<emphasis id="B941EACA0F36FF94FBEA65AECC8657A6" box="[1122,1206,1743,1766]" italics="true" pageId="21" pageNumber="520">caspius</emphasis>
(Wehncke, 1875))
</taxonomicName>
; epipleuron comparably narrow, narrower than mesotibia distally; carina meeting inner margin of epipleuron forming a comparably wide angle (&lt;ca. 150°; similar to
<figureCitation id="130E2A5D0F36FF94FC9F6477CB5A566F" box="[791,874,1814,1839]" captionStart="FIGURES 49 – 55" captionStartId="21.[151,264,1345,1367]" captionTargetBox="[191,1436,208,1316]" captionTargetId="figure@21.[151,1436,193,1323]" captionTargetPageId="21" captionText="FIGURES 49 – 55. (49 – 53): Epipleuron with oblique epipleural carina and genicular fossa of: (49) Clemnius (s. str.) berneri, (50) Hygrotus (Leptolambus) polonicus polonicus, (51) H. (Coelambus) caspius, (52) H. (Leptolambus) impressopunctatus and (53) Rhithrodytes agnus; epipleural carina (c) and inner margin of epipleuron (m) including angle mentioned in diagnoses of subgenera. (54 – 55): Elytral margin in lateral view of: (54) H. (Coelambus) caspius (strongly ascending to shoulder) and (55) Clemnius (s. str.) berneri (moderately ascending to shoulder) (Figs 52 and 53 are reproduced from Fery 2013 and from Fery 2016 with the permissions of F. Gusenleitner, Linz, Austria, and G. Foster, Ayr, UK, respectively)." httpUri="https://zenodo.org/record/884786/files/figure.png" pageId="21" pageNumber="520">Fig. 51</figureCitation>
). Elytra light yellowish (more yellowish brown in
<taxonomicName id="4C354D5B0F36FF94FF1F645ACAD45613" authority="Hajek, Fery &amp; Erman, 2005" authorityName="Hajek, Fery &amp; Erman" authorityYear="2005" box="[151,740,1850,1875]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="520" phylum="Arthropoda" rank="species" species="ahmeti" subGenus="Coelambus">
<emphasis id="B941EACA0F36FF94FF1F645AC8835612" box="[151,179,1851,1874]" italics="true" pageId="21" pageNumber="520">H.</emphasis>
(
<emphasis id="B941EACA0F36FF94FF4B645BC9715613" box="[195,321,1850,1875]" italics="true" pageId="21" pageNumber="520">Coelambus</emphasis>
)
<emphasis id="B941EACA0F36FF94FED9645BC9AE5613" box="[337,414,1850,1875]" italics="true" pageId="21" pageNumber="520">ahmeti</emphasis>
Hájek, Fery &amp; Erman, 2005
</taxonomicName>
), distinctly vittate (in some species/individuals vittae strongly reduced in anterior half). Venter usually black, but females of some species (e.g.
<taxonomicName id="4C354D5B0F36FF94FBA5643ECD695637" baseAuthorityName="Aube" baseAuthorityYear="1850" box="[1069,1369,1886,1911]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="520" phylum="Arthropoda" rank="species" species="pallidulus" subGenus="Coelambus">
<emphasis id="B941EACA0F36FF94FBA5643ECC795636" box="[1069,1097,1887,1910]" italics="true" pageId="21" pageNumber="520">H.</emphasis>
(
<emphasis id="B941EACA0F36FF94FBD2643FCCE85637" box="[1114,1240,1886,1911]" italics="true" pageId="21" pageNumber="520">Coelambus</emphasis>
)
<emphasis id="B941EACA0F36FF94FB61643FCD695637" box="[1257,1369,1886,1911]" italics="true" pageId="21" pageNumber="520">pallidulus</emphasis>
</taxonomicName>
) with abdomen at least in part yellow or brownish. Last abdominal ventrite without deep depression.
</paragraph>
<paragraph id="8B8A36D80F36FF97FF4F64C7C91851B8" blockId="21.[151,1437,1562,2019]" lastBlockId="22.[151,1436,151,392]" lastPageId="22" lastPageNumber="521" pageId="21" pageNumber="520">
Aedeagus with median lobe elongate, very slender (distal half very thin in lateral view), symmetric or almost symmetric (e.g. in
<taxonomicName id="4C354D5B0F36FF94FEE164AACBB056A3" authority="(Ahrens, 1833)" authorityName="Ahrens" authorityYear="1833" baseAuthorityName="Ahrens" baseAuthorityYear="1833" box="[361,896,1994,2019]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="520" phylum="Arthropoda" rank="species" species="enneagrammus" subGenus="Coelambus">
<emphasis id="B941EACA0F36FF94FEE164AAC9B556A2" box="[361,389,1995,2018]" italics="true" pageId="21" pageNumber="520">H.</emphasis>
(
<emphasis id="B941EACA0F36FF94FE1C64ABCA2256A3" box="[404,530,1994,2019]" italics="true" pageId="21" pageNumber="520">Coelambus</emphasis>
)
<emphasis id="B941EACA0F36FF94FDAB64AACAFE56A2" box="[547,718,1995,2018]" italics="true" pageId="21" pageNumber="520">enneagrammus</emphasis>
(Ahrens, 1833)
</taxonomicName>
, and
<taxonomicName id="4C354D5B0F36FF94FC3364AACCD056A3" authorityName="Fabricius" authorityYear="1787" box="[955,1248,1994,2019]" class="Insecta" family="Dytiscidae" genus="Dytiscus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="520" phylum="Arthropoda" rank="species" species="confluens" subGenus="Coelambus">
<emphasis id="B941EACA0F36FF94FC3364AACBE756A2" box="[955,983,1995,2018]" italics="true" pageId="21" pageNumber="520">H.</emphasis>
(
<emphasis id="B941EACA0F36FF94FC6F64ABCC5556A3" box="[999,1125,1994,2019]" italics="true" pageId="21" pageNumber="520">Coelambus</emphasis>
)
<emphasis id="B941EACA0F36FF94FBFD64ABCCD056A3" box="[1141,1248,1994,2019]" italics="true" pageId="21" pageNumber="520">confluens</emphasis>
</taxonomicName>
; see
<figureCitation id="130E2A5D0F36FF97FA9164ABC88751F0" captionStart="FIGURES 19 – 28" captionStartId="17.[151,264,1527,1549]" captionTargetBox="[158,1432,199,1502]" captionTargetId="figure@17.[151,1436,193,1506]" captionTargetPageId="17" captionText="FIGURES 19 – 28. Median lobe in ventral and lateral view of: (19) Clemnius (Cyclopius) acaroides, (20) Clemnius (s. str.) berneri, (21) Clemnius (s. str.) decoratus, (22) Hygrotus (s. str.) inaequalis, (23) H. (Coelambus) enneagrammus, (24) H. (Coelambus) confluens, (25) H. (Leptolambus) parallellogrammus, (26) H. (Leptolambus) impressopunctatus, (27) H. (s. str.) guineensis and (28) H. (Hyphoporus) tonkinensis (Fig. 28 is a modified reproduction of figs 3 and 4 in Brancucci &amp; Biström 2013)." httpUri="https://zenodo.org/record/884780/files/figure.png" lastPageId="22" lastPageNumber="521" pageId="21" pageNumber="520">Figs 23 and 24</figureCitation>
). Parameres with condylar process rather long and forming an almost right angle with distal part (
<figureCitation id="130E2A5D0F35FF97FA8763F6C8835194" captionStart="FIGURES 29 – 40" captionStartId="18.[151,264,1665,1687]" captionTargetBox="[151,1430,221,1637]" captionTargetId="figure@18.[144,1443,193,1643]" captionTargetPageId="18" captionText="FIGURES 29 – 40. Left paramere of: (29) Clemnius (Cyclopius) acaroides, (30) Clemnius (s. str.) berneri, (31) Clemnius (s. str.) decoratus, (32) Hygrotus (s. str.) inaequalis, (33) H. (Leptolambus) fresnedai, (34) H. (Leptolambus) impressopunctatus, (35) H. (Leptolambus) parallellogrammus, (36) H. (Leptolambus) nubilus, (37) H. (Coelambus) enneagrammus, (38) H. (Coelambus) confluens, (39) H. (s. str.) guineensis and (40) H. (Hyphoporus) tonkinensis (Fig. 40 is a modified reproduction of fig. 5 in Brancucci &amp; Biström 2013)." httpUri="https://zenodo.org/record/884782/files/figure.png" pageId="22" pageNumber="521">Figs. 37 and 38</figureCitation>
for
<taxonomicName id="4C354D5B0F35FF97FF6263DCCA675194" baseAuthorityName="Ahrens" baseAuthorityYear="1833" box="[234,599,188,213]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="521" phylum="Arthropoda" rank="species" species="enneagrammus" subGenus="Coelambus">
<emphasis id="B941EACA0F35FF97FF6263DCC9355194" box="[234,261,189,212]" italics="true" pageId="22" pageNumber="521">H.</emphasis>
(
<emphasis id="B941EACA0F35FF97FE9163DDC9A75195" box="[281,407,188,213]" italics="true" pageId="22" pageNumber="521">Coelambus</emphasis>
)
<emphasis id="B941EACA0F35FF97FE2463DCCA675194" box="[428,599,189,212]" italics="true" pageId="22" pageNumber="521">enneagrammus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C354D5B0F35FF97FD1D63DCCBF25195" authorityName="Fabricius" authorityYear="1787" box="[661,962,188,213]" class="Insecta" family="Dytiscidae" genus="Dytiscus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="521" phylum="Arthropoda" rank="species" species="confluens" subGenus="Coelambus">
<emphasis id="B941EACA0F35FF97FD1D63DCCA815194" box="[661,689,189,212]" italics="true" pageId="22" pageNumber="521">H.</emphasis>
(
<emphasis id="B941EACA0F35FF97FD4C63DDCB725195" box="[708,834,188,213]" italics="true" pageId="22" pageNumber="521">Coelambus</emphasis>
)
<emphasis id="B941EACA0F35FF97FCDF63DDCBF25195" box="[855,962,188,213]" italics="true" pageId="22" pageNumber="521">confluens</emphasis>
</taxonomicName>
, respectively). Male metatarsal claws of equal length.
</paragraph>
</subSubSection>
<subSubSection id="C32F65530F35FF97FF4F6265CB3550C8" pageId="22" pageNumber="521" type="distribution">
<paragraph id="8B8A36D80F35FF97FF4F6265C9765000" blockId="22.[151,1436,151,392]" pageId="22" pageNumber="521">
<emphasis id="B941EACA0F35FF97FF4F6265C950505D" bold="true" box="[199,352,260,285]" pageId="22" pageNumber="521">Distribution.</emphasis>
Mainly Palaearctic, with one species in
<collectingCountry id="F32276480F35FF97FCC26265CBAE505D" box="[842,926,260,285]" name="Canada" pageId="22" pageNumber="521">Canada</collectingCountry>
and northern
<collectingCountry id="F32276480F35FF97FBC26265CCB2505C" box="[1098,1154,260,284]" name="United States of America" pageId="22" pageNumber="521">USA</collectingCountry>
(
<taxonomicName id="4C354D5B0F35FF97FB106264C9065000" authorityName="" baseAuthorityName="Fall" baseAuthorityYear="1919" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="521" phylum="Arthropoda" rank="species" species="punctilineatus" subGenus="Coelambus">
<emphasis id="B941EACA0F35FF97FB106264CCCF505C" box="[1176,1279,261,284]" italics="true" pageId="22" pageNumber="521">Hygrotus</emphasis>
(
<emphasis id="B941EACA0F35FF97FA9C6265CDA2505D" box="[1300,1426,260,285]" italics="true" pageId="22" pageNumber="521">Coelambus</emphasis>
)
<emphasis id="B941EACA0F35FF97FF1F6246C9065000" box="[151,310,295,320]" italics="true" pageId="22" pageNumber="521">punctilineatus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B8A36D80F35FF97FF4F622DCB3550C8" blockId="22.[151,1436,151,392]" pageId="22" pageNumber="521">
<emphasis id="B941EACA0F35FF97FF4F622DC98A5025" bold="true" box="[199,442,332,357]" pageId="22" pageNumber="521">Main habitat types.</emphasis>
The subgenus includes species typical of both lentic and lotic habitats, with varied salinity tolerance going from freshwater to hypersaline.
</paragraph>
</subSubSection>
</treatment>
</document>
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