254 lines
35 KiB
XML
254 lines
35 KiB
XML
<document id="6151A4B35F95BF99EF54BA0541818382" ID-DOI="10.5281/zenodo.215242" ID-GBIF-Dataset="df0b1e64-69fb-475f-bf89-3df982ea1cd3" ID-ISSN="1175-5326" ID-Zenodo-Dep="215242" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460476121135" checkinUser="plazi" docAuthor="Hünefeld, Frank & Kristensen, Niels P." docDate="2012" docId="039C87981C0C1C531990F95D996BB721" docLanguage="en" docName="zt03281p068.pdf" docOrigin="Zootaxa 3281" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Heterobathmia megadecella Hünefeld & Kristensen, 2012, n. sp." docType="treatment" docVersion="9" lastPageNumber="66" masterDocId="FFA5FFE01C0D1C561907FFC39854B645" masterDocTitle="Two new heterobathmiid moth species with distinctive female genital configurations (Lepidoptera: Heterobathmiidae)" masterLastPageNumber="68" masterPageNumber="61" pageNumber="62" updateTime="1698306746004" updateUser="plazi">
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<mods:mods id="E2EB440C7A13400182E33BC648B11FC7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8CD42BC3A98DC64BC06E741140645B69">
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<mods:title id="50D46E33C2ECA179800312447C019EFE">Two new heterobathmiid moth species with distinctive female genital configurations (Lepidoptera: Heterobathmiidae)</mods:title>
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<mods:roleTerm id="F8CB70559CC65714877692D1A870CF35">Author</mods:roleTerm>
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<mods:namePart id="232AE53DA298AA6ED2A7B3FCEBDCE5FD">Hünefeld, Frank</mods:namePart>
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<mods:roleTerm id="D0B165DEFE7DBE12BFB2AD3F4D5831D8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9B79344837E7C8A597A077F7C63CCC53">Kristensen, Niels P.</mods:namePart>
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<mods:typeOfResource id="B027AD195AE28582DA5ECC9DE34E965E">text</mods:typeOfResource>
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<mods:title id="C4436B972955CCC4666F5B4EE9DA23C3">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="8FF0BC409D995A34FDC20B904BFC5FF3">
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<mods:date id="1B71B3949AE698D8BD19C250CF67B029">2012</mods:date>
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<mods:detail id="254C3C58A1F5C13C3E20F121AE0554A7" type="volume">
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<mods:number id="F1C45D93056FFB8F4BB38ED6DE8CE5C3">3281</mods:number>
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<mods:classification id="F9480C197ABF1885DC22279461CC063B">journal article</mods:classification>
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<mods:identifier id="F7A82EA247C0C0B55D594DAAA8B7F8F5" type="DOI">10.5281/zenodo.215242</mods:identifier>
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<mods:identifier id="0DE17A8A4EDBAE8947F566EDC1B7924E" type="GBIF-Dataset">df0b1e64-69fb-475f-bf89-3df982ea1cd3</mods:identifier>
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<mods:identifier id="99D7D83F8C5B8DB6853D7BB401C2180C" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="1D8626C9CCB869864C82F0310042CF4A" type="Zenodo-Dep">215242</mods:identifier>
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<treatment id="039C87981C0C1C531990F95D996BB721" ID-DOI="http://doi.org/10.5281/zenodo.6180623" ID-GBIF-Taxon="119516265" ID-Zenodo-Dep="6180623" LSID="urn:lsid:plazi:treatment:039C87981C0C1C531990F95D996BB721" httpUri="http://treatment.plazi.org/id/039C87981C0C1C531990F95D996BB721" lastPageId="5" lastPageNumber="66" pageId="1" pageNumber="62">
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<subSubSection id="C32F65051C0C1C571990F95D9A60B0FD" box="[151,564,1694,1720]" pageId="1" pageNumber="62" type="nomenclature">
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<paragraph id="8B8A368E1C0C1C571990F95D9A60B0FD" blockId="1.[151,564,1694,1720]" box="[151,564,1694,1720]" pageId="1" pageNumber="62">
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<heading id="D0C281E21C0C1C571990F95D9A60B0FD" bold="true" box="[151,564,1694,1720]" fontSize="11" level="1" pageId="1" pageNumber="62" reason="1">
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<emphasis id="B941EA9C1C0C1C571990F95D9A60B0FD" bold="true" box="[151,564,1694,1720]" pageId="1" pageNumber="62">
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<taxonomicName id="4C354D0D1C0C1C571990F95D99BFB0FD" box="[151,491,1694,1720]" class="Insecta" family="Heterobathmiidae" genus="Heterobathmia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="megadecella" status="sp. nov.">
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<emphasis id="B941EA9C1C0C1C571990F95D99BFB0FD" bold="true" box="[151,491,1694,1720]" italics="true" pageId="1" pageNumber="62">Heterobathmia megadecella</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27257E71C0C1C5718F5F95D9A60B0FD" box="[498,564,1694,1720]" pageId="1" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C32F65051C0C1C571990F9269D3DB1CB" pageId="1" pageNumber="62" type="materials_examined">
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<paragraph id="8B8A368E1C0C1C571990F9269D3DB1CB" blockId="1.[151,1436,1765,2005]" pageId="1" pageNumber="62">
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||
<emphasis id="B941EA9C1C0C1C571990F926992EB0BB" bold="true" box="[151,378,1765,1790]" pageId="1" pageNumber="62">Material examined</emphasis>
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:
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<emphasis id="B941EA9C1C0C1C57188EF92699A4B0B8" box="[393,496,1765,1789]" italics="true" pageId="1" pageNumber="62">
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<typeStatus id="548E882C1C0C1C57188EF92699A4B0B8" box="[393,496,1765,1789]" pageId="1" pageNumber="62" type="holotype">Holotype</typeStatus>
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</emphasis>
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3:
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<collectingCountry id="F322761E1C0C1C571B1BF9269A90B0B8" box="[540,708,1765,1789]" name="Argentina" pageId="1" pageNumber="62">ARGENTINA</collectingCountry>
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: Neuquen [Province] [sta.] 12, S. M. [San Martin] de los Andes, Piedra Trampul,
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<quantity id="4CCD9B6B1C0C1C57185BF8CA99ECB164" box="[348,440,1801,1825]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="62" unit="m" value="1000.0">1000 m</quantity>
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,
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<date id="FF8B104E1C0C1C5718CEF8CA9A6FB164" box="[457,571,1801,1825]" pageId="1" pageNumber="62" value="1981-10-15">15.x.1981</date>
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, Nielsen & Karsholt, genitalia slide
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<collectionCode id="ED24AE4B1C0C1C571AE8F8CA9C70B164" LSID="urn:lsid:biocol.org:col:12973" box="[1007,1060,1801,1825]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12973" name="École des Sciences de Niamey" pageId="1" pageNumber="62">ESN</collectionCode>
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2241.
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<emphasis id="B941EA9C1C0C1C571D7CF8C99CB8B164" box="[1147,1260,1802,1825]" italics="true" pageId="1" pageNumber="62">
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<typeStatus id="548E882C1C0C1C571D7CF8C99CB8B164" box="[1147,1260,1802,1825]" pageId="1" pageNumber="62" type="paratype">Paratypes</typeStatus>
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</emphasis>
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: Same data as
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<typeStatus id="548E882C1C0C1C571990F8EE98ADB100" box="[151,249,1837,1861]" pageId="1" pageNumber="62" type="holotype">holotype</typeStatus>
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(33);
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<collectingCountry id="F322761E1C0C1C571861F8EE99E9B100" box="[358,445,1837,1861]" name="Chile" pageId="1" pageNumber="62">CHILE</collectingCountry>
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: Osorno [Province] [sta.] 37, Parque Nacional Puyehue, Antillanca,
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<quantity id="4CCD9B6B1C0C1C571DF2F8EE9DC3B100" box="[1269,1431,1837,1861]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="1" pageNumber="62" unit="m" value="1200.0" valueMax="1300.0" valueMin="1100.0">1100–1300 m</quantity>
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,
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<date id="FF8B104E1C0C1C571990F8929945B12C" box="[151,273,1873,1897]" pageId="1" pageNumber="62" value="1981-11-14">14.xi.1981</date>
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, Nielsen & Karsholt (23; 1Ƥ, genitalia preparation NPK 1074). All in Natural History Museum of
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<collectingCountry id="F322761E1C0C1C571990F8B69957B1C8" box="[151,259,1909,1933]" name="Denmark" pageId="1" pageNumber="62">Denmark</collectingCountry>
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. Wing and body vestiture well preserved in
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<typeStatus id="548E882C1C0C1C571BE8F8B69B05B1C8" box="[751,849,1909,1933]" pageId="1" pageNumber="62" type="holotype">holotype</typeStatus>
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and female
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<typeStatus id="548E882C1C0C1C571ADDF8B59C69B1CB" box="[986,1085,1910,1934]" pageId="1" pageNumber="62" type="paratype">paratype</typeStatus>
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, poorly in other
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<typeStatus id="548E882C1C0C1C571DF1F8B59D31B1CB" box="[1270,1381,1910,1934]" pageId="1" pageNumber="62" type="paratype">paratypes</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32F65051C0C1C5419C0F85A9BF4B6B2" lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62" type="diagnosis">
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<paragraph id="8B8A368E1C0C1C5419C0F85A9BF4B6B2" blockId="1.[151,1436,1765,2005]" lastBlockId="2.[151,1436,151,247]" lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62">
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<emphasis id="B941EA9C1C0C1C5719C0F85A996EB1F7" bold="true" box="[199,314,1945,1970]" pageId="1" pageNumber="62">Diagnosis</emphasis>
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. Distinguishable from the other described
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<taxonomicName id="4C354D0D1C0C1C571A21F85A9B9BB1F4" box="[806,975,1945,1969]" class="Insecta" family="Heterobathmiidae" genus="Heterobathmia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
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<emphasis id="B941EA9C1C0C1C571A21F85A9B9BB1F4" box="[806,975,1945,1969]" italics="true" pageId="1" pageNumber="62">Heterobathmia</emphasis>
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</taxonomicName>
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species by the male’s prominent arched dorsum X sclerite, broad, trapezoid valves, and minute medial teguminal processes, and the female’s sclerotizations on the paramedian longitudinal elevations of the membranous lower surface of the terminal unit, as well as a conspicuous sclerotization in the genital chamber. Well-preserved specimens are distinctive by the very extensive suffusion of silvery-white scales on the forewing (
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<figureCitation id="130E2A0B1C0F1C541A3DFF1C9B2DB6B2" box="[826,889,223,247]" captionStart="FIGURE 1" captionStartId="2.[151,250,648,671]" captionTargetBox="[221,1365,305,615]" captionTargetId="figure@2.[221,1366,305,615]" captionTargetPageId="2" captionText="FIGURE 1. Heterobathmia habitus. A, H. megadecella male holotype; B, H. nielsenella, female holotype." httpUri="https://zenodo.org/record/215243/files/figure.png" pageId="2" pageNumber="63">Fig. 1</figureCitation>
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A).
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</paragraph>
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||
</subSubSection>
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||
<caption id="DF4A66061C0F1C541990FD4B9C85B4DB" box="[151,1233,648,671]" httpUri="https://zenodo.org/record/215243/files/figure.png" pageId="2" pageNumber="63" targetBox="[221,1365,305,615]" targetPageId="2">
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<paragraph id="8B8A368E1C0F1C541990FD4B9C85B4DB" blockId="2.[151,1233,648,671]" box="[151,1233,648,671]" pageId="2" pageNumber="63">
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<emphasis id="B941EA9C1C0F1C541990FD4B9947B4D8" bold="true" box="[151,275,648,671]" pageId="2" pageNumber="63">FIGURE 1.</emphasis>
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<taxonomicName id="4C354D0D1C0F1C54181EFD4B99E5B4DB" box="[281,433,648,670]" class="Insecta" family="Heterobathmiidae" genus="Heterobathmia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
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<emphasis id="B941EA9C1C0F1C54181EFD4B99E5B4DB" box="[281,433,648,670]" italics="true" pageId="2" pageNumber="63">Heterobathmia</emphasis>
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</taxonomicName>
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habitus. A,
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<taxonomicName id="4C354D0D1C0F1C541B2BFD4A9A9FB4DB" box="[556,715,648,670]" class="Insecta" family="Heterobathmiidae" genus="Heterobathmia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="megadecella">
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<emphasis id="B941EA9C1C0F1C541B2BFD4A9A9FB4DB" box="[556,715,648,670]" italics="true" pageId="2" pageNumber="63">H. megadecella</emphasis>
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</taxonomicName>
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male holotype; B,
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<taxonomicName id="4C354D0D1C0F1C541A8BFD4A9C4DB4DB" box="[908,1049,648,670]" class="Insecta" family="Heterobathmiidae" genus="Heterobathmia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="nielsenella">
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<emphasis id="B941EA9C1C0F1C541A8BFD4A9C4DB4DB" box="[908,1049,648,670]" italics="true" pageId="2" pageNumber="63">H. nielsenella</emphasis>
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</taxonomicName>
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, female holotype.
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</paragraph>
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</caption>
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<subSubSection id="C32F65051C0F1C5219C0FD099986B77A" lastPageId="4" lastPageNumber="65" pageId="2" pageNumber="63" type="description">
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<paragraph id="8B8A368E1C0F1C5419C0FD099B02B542" blockId="2.[151,1437,714,1855]" pageId="2" pageNumber="63">
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<emphasis id="B941EA9C1C0F1C5419C0FD099905B4A6" bold="true" box="[199,337,714,739]" pageId="2" pageNumber="63">Description</emphasis>
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:
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<taxonomicName id="4C354D0D1C0F1C541866FD0F9A46B4A6" box="[353,530,715,739]" class="Insecta" family="Heterobathmiidae" genus="Heterobathmia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="megadecella">
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<emphasis id="B941EA9C1C0F1C541866FD0F9A46B4A6" box="[353,530,715,739]" italics="true" pageId="2" pageNumber="63">H. megadecella</emphasis>
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</taxonomicName>
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is generally larger than the previously named
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<taxonomicName id="4C354D0D1C0F1C541D22FD089C9AB4A6" box="[1061,1230,715,739]" class="Insecta" family="Heterobathmiidae" genus="Heterobathmia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
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<emphasis id="B941EA9C1C0F1C541D22FD089C9AB4A6" box="[1061,1230,715,739]" italics="true" pageId="2" pageNumber="63">Heterobathmia</emphasis>
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</taxonomicName>
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species; forewing length in male
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<quantity id="4CCD9B6B1C0F1C541839FD2C999CB542" box="[318,456,751,775]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" metricValueMax="5.5" metricValueMin="5.1" pageId="2" pageNumber="63" unit="mm" value="5.3" valueMax="5.5" valueMin="5.1">5.1–5.5 mm</quantity>
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||
; in single available female
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||
<quantity id="4CCD9B6B1C0F1C541BFEFD2C9B06B542" box="[761,850,751,775]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="63" unit="mm" value="4.4">4.4 mm</quantity>
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||
.
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||
</paragraph>
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||
<paragraph id="8B8A368E1C0F1C5419C0FCD79D29B246" blockId="2.[151,1437,714,1855]" pageId="2" pageNumber="63">
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||
<emphasis id="B941EA9C1C0F1C5419C0FCD7995EB56E" box="[199,266,788,811]" italics="true" pageId="2" pageNumber="63">Wings</emphasis>
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||
. Forewing ground coloration light brown with some purplish iridescence and in well-preserved specimens with light greyish appearance due to very extensive suffusion with silvery-white scales. This suffusion may be least developed or completely lacking on a large part of the anal area, on the dorsum just beyond the tornal spot, and in an area around the Rs-M1 crossvein and M1/M2 fork, but we are uncertain to which degree absence of silvery-white scales here and (in particular) in other areas is due to accidental scale loss. Tornal spot distinctive, but + pronouncedly confluent with adjacent white scaling. Hindwing grey with purplish and greenish iridescence. In marginal ‘fringes’ of both wing pairs silvery-white piliform scales form prominent spots around R-Rs1 and M2.
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</paragraph>
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||
<paragraph id="8B8A368E1C0F1C5419C0FBCC9CE9B006" blockId="2.[151,1437,714,1855]" pageId="2" pageNumber="63">
|
||
<emphasis id="B941EA9C1C0F1C5419C0FBCC9939B262" box="[199,365,1039,1063]" italics="true" pageId="2" pageNumber="63">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="130E2A0B1C0F1C541879FBCC9990B262" box="[382,452,1039,1063]" captionStart="FIGURE 2" captionStartId="3.[151,250,1513,1536]" captionTargetBox="[214,1370,195,1490]" captionTargetId="figure@3.[214,1372,193,1492]" captionTargetPageId="3" captionText="FIGURE 2. Heterobathmia megadecella n. sp., male genitalia, details in slide preparations as prepared for previous taxonomic descriptions of congeners (A & C – E), or stacked multifocus photographs of intact specimen (B). A, dorsum IX plus postgenital complex of holotype, dorsal view; ventral acanthae on the latter visible through cleared tergum X plate; B, postabdominal apex in dorsal view of paratype, notice contrast in pigmentation and texture between dorsum IX and extraordinarily large tergum X; C, venter IX with valvae of holotype; D, phallus of holotype; E, phallic apex of paratype, part of scaly-textured connecting zone between outer tube and syringe-like process from ejaculatory duct externally exposed. fr, sclerotized ‘ frame’ of postgenital complex; mp, medial teguminal process; lp, lateral teguminal process; te, ‘ tegumen’ (dorsum IX); tX, tergal plate of segment X; vi, ‘ vinculum’ (venter IX), vl, valve." httpUri="https://zenodo.org/record/215244/files/figure.png" pageId="2" pageNumber="63">Fig. 2</figureCitation>
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). Sternum VIII with large unmelanized median area. Segment IX with ‘tegumen’ and ‘vinculum’ completely separated by narrow membranous cleft. Anteromedian vinculum margin produced into prominent paired close-set processes on common base. Tegumen relatively longer than in other described species, anterior margin strongly convex, posterior margin more strongly concave. Tegumen posteriorly without strengthened ridge (present in the other described
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<taxonomicName id="4C354D0D1C0F1C541BD4FB5C9B2FB2F2" box="[723,891,1183,1207]" class="Insecta" family="Heterobathmiidae" genus="Heterobathmia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
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<emphasis id="B941EA9C1C0F1C541BD4FB5C9B2FB2F2" box="[723,891,1183,1207]" italics="true" pageId="2" pageNumber="63">Heterobathmia</emphasis>
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</taxonomicName>
|
||
species). Lateral teguminal process longer than postgenital complex, tapering and with slight S-bend. Medial teguminal process relatively much smaller than in other named species, tapering, only slightly curved, apex pointed. Valve extremely broad, almost trapezoid; upper margin concave near base; lower hind corner slightly produced, more rounded than in other species, with densely clustered, stout sensilla; apex with particularly long and strong setae. Postgenital complex with tergum X exceptionally well developed, saddle-shaped, synscleritous with tegumen only in middle region, with smooth, glossy surface without setation. Anterior margin of tergum X almost straight, posterior margin convexly rounded. ‘Frame’ of postgenital complex clearly separate from tergum X, slender, without anterior bridge between lateral frame sclerites. Frame sclerites without lateral extensions, posteriorly strongly converging, apices medially fused. Fused apical regions of frame sclerites with numerous simple acanthae and tubular, apically pointed cuticular processes, much longer/more prominent than any counterparts in other described
|
||
<taxonomicName id="4C354D0D1C0F1C541D38F9C49CB3B05A" box="[1087,1255,1543,1567]" class="Insecta" family="Heterobathmiidae" genus="Heterobathmia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EA9C1C0F1C541D38F9C49CB3B05A" box="[1087,1255,1543,1567]" italics="true" pageId="2" pageNumber="63">Heterobathmia</emphasis>
|
||
</taxonomicName>
|
||
species. Lower surface of postgenital complex densely set with very long, delicate cuticular ‘hairs’ (acanthae).
|
||
</paragraph>
|
||
<paragraph id="8B8A368E1C0F1C5419C0F98C9986B17A" blockId="2.[151,1437,714,1855]" pageId="2" pageNumber="63">
|
||
Phallus overall similar to that of the named species, strongly sclerotized, with hyaline ‘punctures’ (the nature of which still remains to be ascertained) on base of outer tube. As in previously described congeners the intima of the ejaculatory duct is sclerotized; it is produced into a seemingly eversible and purely cuticular tube terminating in a syringe-like (dorsally open) apical spine. The membranous ‘endophallus’ connecting this tube to the apex of the outer phallic tube bears a patch of melanized and + pointed scale-like protuberances arranged in a tile-like fashion; depending on the degree of protrusion of the terminal tube none or some of these protuberances may become located externally (
|
||
<figureCitation id="130E2A0B1C0F1C541869F8E499E5B17A" box="[366,433,1831,1855]" captionStart="FIGURE 2" captionStartId="3.[151,250,1513,1536]" captionTargetBox="[214,1370,195,1490]" captionTargetId="figure@3.[214,1372,193,1492]" captionTargetPageId="3" captionText="FIGURE 2. Heterobathmia megadecella n. sp., male genitalia, details in slide preparations as prepared for previous taxonomic descriptions of congeners (A & C – E), or stacked multifocus photographs of intact specimen (B). A, dorsum IX plus postgenital complex of holotype, dorsal view; ventral acanthae on the latter visible through cleared tergum X plate; B, postabdominal apex in dorsal view of paratype, notice contrast in pigmentation and texture between dorsum IX and extraordinarily large tergum X; C, venter IX with valvae of holotype; D, phallus of holotype; E, phallic apex of paratype, part of scaly-textured connecting zone between outer tube and syringe-like process from ejaculatory duct externally exposed. fr, sclerotized ‘ frame’ of postgenital complex; mp, medial teguminal process; lp, lateral teguminal process; te, ‘ tegumen’ (dorsum IX); tX, tergal plate of segment X; vi, ‘ vinculum’ (venter IX), vl, valve." httpUri="https://zenodo.org/record/215244/files/figure.png" pageId="2" pageNumber="63">Fig. 2</figureCitation>
|
||
E).
|
||
</paragraph>
|
||
<caption id="DF4A66061C0E1C551990FA2A9A7EB093" httpUri="https://zenodo.org/record/215244/files/figure.png" pageId="3" pageNumber="64" targetBox="[214,1370,195,1490]" targetPageId="3">
|
||
<paragraph id="8B8A368E1C0E1C551990FA2A9A7EB093" blockId="3.[151,1436,1513,1750]" pageId="3" pageNumber="64">
|
||
<emphasis id="B941EA9C1C0E1C551990FA2A9947B3BB" bold="true" box="[151,275,1513,1536]" pageId="3" pageNumber="64">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C354D0D1C0E1C55181EFA2A9A6CB3BA" box="[281,568,1513,1535]" class="Insecta" family="Heterobathmiidae" genus="Heterobathmia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="megadecella" status="sp. nov.">
|
||
<emphasis id="B941EA9C1C0E1C55181EFA2A9A6CB3BA" box="[281,568,1513,1535]" italics="true" pageId="3" pageNumber="64">Heterobathmia megadecella</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27257E71C0E1C551B39FA299A24B3BA" box="[574,624,1514,1535]" pageId="3" pageNumber="64" rank="species">
|
||
<emphasis id="B941EA9C1C0E1C551B39FA299A3EB3BA" bold="true" box="[574,618,1514,1535]" pageId="3" pageNumber="64">n.sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, male genitalia, details in slide preparations as prepared for previous taxonomic descriptions of congeners (A & C–E), or stacked multifocus photographs of intact specimen (B). A, dorsum IX plus postgenital complex of holotype, dorsal view; ventral acanthae on the latter visible through cleared tergum X plate; B, postabdominal apex in dorsal view of paratype, notice contrast in pigmentation and texture between dorsum IX and extraordinarily large tergum X; C, venter IX with valvae of holotype; D, phallus of holotype; E, phallic apex of paratype, part of scaly-textured connecting zone between outer tube and syringe-like process from ejaculatory duct externally exposed. fr, sclerotized ‘frame’ of postgenital complex; mp, medial teguminal process; lp, lateral teguminal process; te, ‘tegumen’ (dorsum IX); tX, tergal plate of segment X; vi, ‘vinculum’ (venter IX), vl, valve.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8A368E1C0E1C5519C0F8C19DC8B1B7" blockId="3.[151,1437,1794,2034]" pageId="3" pageNumber="64">
|
||
<emphasis id="B941EA9C1C0E1C5519C0F8C199D2B15F" box="[199,390,1794,1818]" italics="true" pageId="3" pageNumber="64">Female genitalia</emphasis>
|
||
(
|
||
<figureCitation id="130E2A0B1C0E1C551892F8C1998CB15F" box="[405,472,1794,1818]" captionStart="FIGURE 3" captionStartId="4.[151,250,1537,1560]" captionTargetBox="[224,1362,383,1515]" captionTargetId="figure@4.[224,1363,353,1517]" captionTargetPageId="4" captionText="FIGURE 3. Heterobathmia megadecella n. sp., postabdomen of unique female paratype. A, ventral view, stacked multifocus photographs of KOH-macerated specimen in ethanol, supported on ‘ Ballotili’ glass beads; B, lateral view, conventional slide mount in Euparal, rectum / anal tube partly filled with non-degradable material (including loosened wing scales), hence its position / course easily identifiable; C, magnified detail of B, showing optical section of anterior part of genital chamber. db, (non-spinose posterior part of) ductus bursae; ds, ductus spermathecae; gcs, sclerotization of invaginated thick-walled papilla in genital chamber; 1 - 2 - 3, sclerites on softwalled lower surface of terminal unit thus denoted in text." httpUri="https://zenodo.org/record/215245/files/figure.png" pageId="3" pageNumber="64">Fig. 3</figureCitation>
|
||
). Subgenital plate (sclerotized venter VIII plate) ≈ 1.8x length of tergum VIII. Dorsal sclerotization of terminal unit (i.e., dorsum IX+X) with area of unmelanized cuticle along mid-line. Apophyses from anterolateral corners ≈ 0.2x dorsolateral length of terminal unit. Posterior part of dorsal sclerotization appearing set off from the larger anterior part by distinctive unmelanized indentation in lower margin of dorsal sclerite. Membranous lower surface of terminal unit with paired longitudinal elevations, each of which bearing 1) a moderately melanized, posteriorly tapering plate extending beyond mid-length of elevation 2) an adjacent smaller and more anterior, strongly melanized plate, and 3) a long ribbon-like plate on anterior part of inner surface (
|
||
<figureCitation id="130E2A0B1C0E1C551C69F8199DC8B1B7" box="[1390,1436,2010,2034]" captionStart="FIGURE 3" captionStartId="4.[151,250,1537,1560]" captionTargetBox="[224,1362,383,1515]" captionTargetId="figure@4.[224,1363,353,1517]" captionTargetPageId="4" captionText="FIGURE 3. Heterobathmia megadecella n. sp., postabdomen of unique female paratype. A, ventral view, stacked multifocus photographs of KOH-macerated specimen in ethanol, supported on ‘ Ballotili’ glass beads; B, lateral view, conventional slide mount in Euparal, rectum / anal tube partly filled with non-degradable material (including loosened wing scales), hence its position / course easily identifiable; C, magnified detail of B, showing optical section of anterior part of genital chamber. db, (non-spinose posterior part of) ductus bursae; ds, ductus spermathecae; gcs, sclerotization of invaginated thick-walled papilla in genital chamber; 1 - 2 - 3, sclerites on softwalled lower surface of terminal unit thus denoted in text." httpUri="https://zenodo.org/record/215245/files/figure.png" pageId="3" pageNumber="64">Fig.</figureCitation>
|
||
</paragraph>
|
||
<paragraph id="8B8A368E1C091C521990FF549986B77A" blockId="4.[151,1436,151,319]" pageId="4" pageNumber="65">
|
||
3A). Last-mentioned plate as strongly melanized as the second, its blunt anterior apex extending particularly far anteriorly. Genital chamber in front of origin of ductus spermathecae remarkable for having a conspicuous annular sclerotization (gcs) encircling the invaginated thick-walled papilla on the (posteriorly produced) apex of which the ductus bursae opens into the chamber; this sclerotization is most strongly melanized around a transverse groove on its ventral surface (
|
||
<figureCitation id="130E2A0B1C091C52186AFEE499E4B77A" box="[365,432,295,319]" captionStart="FIGURE 3" captionStartId="4.[151,250,1537,1560]" captionTargetBox="[224,1362,383,1515]" captionTargetId="figure@4.[224,1363,353,1517]" captionTargetPageId="4" captionText="FIGURE 3. Heterobathmia megadecella n. sp., postabdomen of unique female paratype. A, ventral view, stacked multifocus photographs of KOH-macerated specimen in ethanol, supported on ‘ Ballotili’ glass beads; B, lateral view, conventional slide mount in Euparal, rectum / anal tube partly filled with non-degradable material (including loosened wing scales), hence its position / course easily identifiable; C, magnified detail of B, showing optical section of anterior part of genital chamber. db, (non-spinose posterior part of) ductus bursae; ds, ductus spermathecae; gcs, sclerotization of invaginated thick-walled papilla in genital chamber; 1 - 2 - 3, sclerites on softwalled lower surface of terminal unit thus denoted in text." httpUri="https://zenodo.org/record/215245/files/figure.png" pageId="4" pageNumber="65">Fig. 3</figureCitation>
|
||
C).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4A66061C091C521990F9C29C20B0F4" httpUri="https://zenodo.org/record/215245/files/figure.png" pageId="4" pageNumber="65" targetBox="[224,1362,383,1515]" targetPageId="4">
|
||
<paragraph id="8B8A368E1C091C521990F9C29C20B0F4" blockId="4.[151,1437,1537,1713]" pageId="4" pageNumber="65">
|
||
<emphasis id="B941EA9C1C091C521990F9C29940B052" bold="true" box="[151,276,1537,1560]" pageId="4" pageNumber="65">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C354D0D1C091C52181BF9C19A68B05D" box="[284,572,1538,1560]" class="Insecta" family="Heterobathmiidae" genus="Heterobathmia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="megadecella" status="sp. nov.">
|
||
<emphasis id="B941EA9C1C091C52181BF9C19A68B05D" box="[284,572,1538,1560]" italics="true" pageId="4" pageNumber="65">Heterobathmia megadecella</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27257E71C091C521B43F9C19A22B05D" box="[580,630,1538,1560]" pageId="4" pageNumber="65" rank="species">
|
||
<emphasis id="B941EA9C1C091C521B43F9C19A24B052" bold="true" box="[580,624,1538,1559]" pageId="4" pageNumber="65">n.sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, postabdomen of unique female paratype. A, ventral view, stacked multifocus photographs of KOH-macerated specimen in ethanol, supported on ‘Ballotili’ glass beads; B, lateral view, conventional slide mount in Euparal, rectum/anal tube partly filled with non-degradable material (including loosened wing scales), hence its position/course easily identifiable; C, magnified detail of B, showing optical section of anterior part of genital chamber. db, (non-spinose posterior part of) ductus bursae; ds, ductus spermathecae; gcs, sclerotization of invaginated thick-walled papilla in genital chamber; 1-2-3, sclerites on softwalled lower surface of terminal unit thus denoted in text.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32F65051C091C5319C0F91E9B8BB691" lastPageId="5" lastPageNumber="66" pageId="4" pageNumber="65" type="distribution">
|
||
<paragraph id="8B8A368E1C091C5219C0F91E9970B15F" blockId="4.[151,1436,1757,2034]" pageId="4" pageNumber="65">
|
||
<emphasis id="B941EA9C1C091C5219C0F91E990DB0B3" bold="true" box="[199,345,1757,1782]" pageId="4" pageNumber="65">Distribution</emphasis>
|
||
: Known from just two localities in Neuquen Province (
|
||
<collectingCountry id="F322761E1C091C521ACAF91D9C14B0B3" box="[973,1088,1758,1782]" name="Argentina" pageId="4" pageNumber="65">Argentina</collectingCountry>
|
||
) and Osorno Province (
|
||
<collectingCountry id="F322761E1C091C521C4AF91D9DDBB0B3" box="[1357,1423,1758,1782]" name="Chile" pageId="4" pageNumber="65">Chile</collectingCountry>
|
||
), respectively.
|
||
</paragraph>
|
||
<paragraph id="8B8A368E1C091C5319C0F8E59B8BB691" blockId="4.[151,1436,1757,2034]" lastBlockId="5.[151,1436,151,356]" lastPageId="5" lastPageNumber="66" pageId="4" pageNumber="65">
|
||
<emphasis id="B941EA9C1C091C5219C0F8E599ACB17A" bold="true" box="[199,504,1830,1855]" pageId="4" pageNumber="65">Phenology and bionomics</emphasis>
|
||
: Adults were collected from
|
||
<date id="FF8B104E1C091C521A44F8E59C2BB17B" box="[835,1151,1830,1854]" pageId="4" pageNumber="65">15 October to 14 November</date>
|
||
at altitudes from
|
||
<date id="FF8B104E1C091C521C40F8E5989AB127" pageId="4" pageNumber="65" value="1000" valueMax="1100">1000 to 1100</date>
|
||
(–1300) meters. Early stages, host plant affinities and life cycle are unknown to us. Judging from the account in
|
||
<bibRefCitation id="EFA44B7F1C091C5219B4F8AD99EAB1C3" box="[179,446,1902,1926]" pageId="4" pageNumber="65" refString="Humphries, C. J., Cox, J. M. & Nielsen, E. S. (1986) Nothofagus and its parasites: A cladistic approach to coevolution. In Stone, A. R. & Hawksworth, D. L. (Eds) Coevolution and Systematics. The Systematics Association Special Volume No. 32. Clarendon Press, Oxford, pp. 55 - 76." type="book chapter">
|
||
Humphries
|
||
<emphasis id="B941EA9C1C091C521831F8AC9932B1C3" box="[310,358,1902,1926]" italics="true" pageId="4" pageNumber="65">et al</emphasis>
|
||
. (1986)
|
||
</bibRefCitation>
|
||
the
|
||
<taxonomicName id="4C354D0D1C091C5218EAF8AD9A3AB1C3" box="[493,622,1902,1926]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="65" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B941EA9C1C091C5218EAF8AD9A3AB1C3" box="[493,622,1902,1926]" italics="true" pageId="4" pageNumber="65">Nothofagus</emphasis>
|
||
</taxonomicName>
|
||
host associations of all
|
||
<taxonomicName id="4C354D0D1C091C521A70F8AD9C4BB1C3" box="[887,1055,1902,1926]" class="Insecta" family="Heterobathmiidae" genus="Heterobathmia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EA9C1C091C521A70F8AD9C4BB1C3" box="[887,1055,1902,1926]" italics="true" pageId="4" pageNumber="65">Heterobathmia</emphasis>
|
||
</taxonomicName>
|
||
species (as then recognized) were known to the late Ebbe S. Nielsen, at least as inferred from observations on the occurrence of the adult moths. We are unable to ascertain how Nielsen’s designations of the undescribed species as ‘species A–H’ in that publication relate to the entities into which he eventually sorted the bulk of the available
|
||
<taxonomicName id="4C354D0D1C091C521D02F8199CF9B1B7" box="[1029,1197,2010,2034]" class="Insecta" family="Heterobathmiidae" genus="Heterobathmia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EA9C1C091C521D02F8199CF9B1B7" box="[1029,1197,2010,2034]" italics="true" pageId="4" pageNumber="65">Heterobathmia</emphasis>
|
||
</taxonomicName>
|
||
material (now in the Natural History Museum of
|
||
<collectingCountry id="F322761E1C081C5318E6FF549A1AB6EA" box="[481,590,151,175]" name="Denmark" pageId="5" pageNumber="66">Denmark</collectingCountry>
|
||
), except that during the description of
|
||
<emphasis id="B941EA9C1C081C531D13FF5A9CDAB6F5" box="[1044,1166,152,176]" italics="true" pageId="5" pageNumber="66">H. valvifer</emphasis>
|
||
a few years before his untimely passing (in 2001) he informed NPK that this one was ‘species B’.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F65051C081C5319C0FF23996BB721" pageId="5" pageNumber="66" type="etymology">
|
||
<paragraph id="8B8A368E1C081C5319C0FF23996BB721" blockId="5.[151,1436,151,356]" pageId="5" pageNumber="66">
|
||
<emphasis id="B941EA9C1C081C5319C0FF23991CB6BD" bold="true" box="[199,328,224,248]" pageId="5" pageNumber="66">Etymology</emphasis>
|
||
: The name
|
||
<taxonomicName id="4C354D0D1C081C5318CCFF239A0DB6BD" box="[459,601,224,248]" class="Insecta" family="Heterobathmiidae" genus="Heterobathmia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="megadecella">
|
||
<emphasis id="B941EA9C1C081C5318CCFF239A0DB6BD" box="[459,601,224,248]" italics="true" pageId="5" pageNumber="66">megadecella</emphasis>
|
||
</taxonomicName>
|
||
refers to the extraordinarily strongly developed dorsum X sclerotization of the male. Ebbe S. Nielsen had given the preliminary name “megadeca” to specimens sorted to this taxon, and we just emended it by using the “-ella” suffix, which is in such widespread use for moths that would have belonged in Linnean
|
||
<taxonomicName id="4C354D0D1C081C5319FDFE8E9963B721" box="[250,311,333,356]" class="Insecta" family="Tineidae" genus="Tinea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941EA9C1C081C5319FDFE8E9963B721" box="[250,311,333,356]" italics="true" pageId="5" pageNumber="66">Tinea</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |