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<document id="16D1553F93291410DC344910D13FA184" ID-CLB-Dataset="7951" ID-DOI="10.11646/zootaxa.4877.3.7" ID-GBIF-Dataset="2e697ef5-3591-4c49-a68c-e32c4d395d7d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4570503" ID-ZooBank="10BCCB7E-377B-4884-AE65-B7C2DD69E5B1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610032969486" checkinUser="plazi" docAuthor="Arun, A. &amp; Ramani, N." docDate="2020" docId="039587D0FFE2470559EB2C4CEB461A92" docLanguage="en" docName="zootaxa.4877.3.7.pdf" docOrigin="Zootaxa 4877 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Papillacarus (Vepracarus) acaciensis Arun &amp; Ramani 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="548" masterDocId="FFACFFA8FFE1470C597C2D4EEE601F20" masterDocTitle="Two remarkable new species of oribatid mites (Acari: Oribatida) from Acacia litter of Kerala, India" masterLastPageNumber="558" masterPageNumber="539" pageNumber="542" updateTime="1698923512612" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="699F427AF2FB0ED53C678E76FCBC2570">Two remarkable new species of oribatid mites (Acari: Oribatida) from Acacia litter of Kerala, India</mods:title>
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<mods:namePart id="564077E19B27182DB5178D862B8E8ED8">Arun, A.</mods:namePart>
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<mods:namePart id="D586F9407E2D45229BE1454630755600">Ramani, N.</mods:namePart>
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<mods:affiliation id="9BD7B17688CB3AF7ACEA9717EC5B12F9">Division of Acarology, Department of Zoology, University of Calicut, Kerala, PO 673635, India. &amp; drnramani @ gmail. com; https: // orcid. org / 0000 - 0002 - 7448 - 439 X</mods:affiliation>
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<treatment id="039587D0FFE2470559EB2C4CEB461A92" ID-DOI="http://doi.org/10.5281/zenodo.4570521" ID-GBIF-Taxon="180324272" ID-Zenodo-Dep="4570521" LSID="urn:lsid:plazi:treatment:039587D0FFE2470559EB2C4CEB461A92" httpUri="http://treatment.plazi.org/id/039587D0FFE2470559EB2C4CEB461A92" lastPageId="9" lastPageNumber="548" pageId="3" pageNumber="542">
<subSubSection id="C326654DFFE2470F59EB2C4CECDC1E3D" box="[151,700,258,285]" pageId="3" pageNumber="542" type="nomenclature">
<paragraph id="8B8336C6FFE2470F59EB2C4CECDC1E3D" blockId="3.[151,700,258,321]" box="[151,700,258,285]" pageId="3" pageNumber="542">
<heading id="D0CB81AAFFE2470F59EB2C4CECDC1E3D" bold="true" box="[151,700,258,285]" fontSize="11" level="1" pageId="3" pageNumber="542" reason="1">
<emphasis id="B948EAD4FFE2470F59EB2C4CECDC1E3D" bold="true" box="[151,700,258,285]" pageId="3" pageNumber="542">
<taxonomicName id="4C3C4D45FFE2470F59EB2C4CEC371E3D" ID-CoL="3778b284-a5a0-4b4d-a2df-6025eb67f2dd" authority="Arun &amp; Ramani, 2020" authorityName="Arun &amp; Ramani" authorityYear="2020" box="[151,599,258,285]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="acaciensis" status="sp. nov." subGenus="Vepracarus">
<emphasis id="B948EAD4FFE2470F59EB2C4CEF501E3D" bold="true" box="[151,304,258,285]" italics="true" pageId="3" pageNumber="542">Papillacarus</emphasis>
(
<emphasis id="B948EAD4FFE2470F583D2C4DEFAA1E3D" bold="true" box="[321,458,259,285]" italics="true" pageId="3" pageNumber="542">Vepracarus</emphasis>
)
<emphasis id="B948EAD4FFE2470F58A72C4DEC371E3D" bold="true" box="[475,599,259,285]" italics="true" pageId="3" pageNumber="542">acaciensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57AFFFE2470F5B232C4DECDC1E3D" box="[607,700,259,285]" pageId="3" pageNumber="542" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<subSubSection id="C326654DFFE2470F59EB2C69EF401E61" box="[151,288,295,321]" pageId="3" pageNumber="542" type="description">
<paragraph id="8B8336C6FFE2470F59EB2C69EF401E61" blockId="3.[151,700,258,321]" box="[151,288,295,321]" pageId="3" pageNumber="542">
(
<figureCitation id="13072A43FFE2470F59E32C69EF781E61" box="[159,280,295,321]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURES 58" captionStart-3="FIGURES 910" captionStart-4="FIGURES 1112" captionStartId-0="4.[152,255,1989,2013]" captionStartId-1="5.[152,255,1991,2015]" captionStartId-2="6.[152,270,1964,1988]" captionStartId-3="7.[152,270,1881,1905]" captionStartId-4="8.[152,270,1961,1985]" captionTargetBox-0="[297,1295,192,1958]" captionTargetBox-1="[388,1198,191,1958]" captionTargetBox-2="[184,1399,199,1926]" captionTargetBox-3="[171,1417,196,1842]" captionTargetBox-4="[267,1313,203,1920]" captionTargetId-0="figure-22@4.[279,1308,181,1965]" captionTargetId-1="figure-19@5.[373,1214,181,1967]" captionTargetId-2="figure-22@6.[172,1416,181,1940]" captionTargetId-3="figure-19@7.[151,1436,181,1857]" captionTargetId-4="figure-22@8.[248,1339,181,1937]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURE 3. Papillacarus (Vepracarus) acaciensis sp. nov., adult dorsal view. Scale bar 50 µm." captionText-1="FIGURE 4. Papillacarus (Vepracarus) acaciensis sp. nov., adult ventral view. Scale bar 50 µm." captionText-2="FIGURES 58. Papillacarus (Vepracarus) acaciensis sp. nov., adult; 5. notogastral seta; 6. subcapitulum, ventral view; 7. chelicera, left; 8. notogastral tubercles. Scale bar 30 µm." captionText-3="FIGURES 910. Papillacarus (Vepracarus) acaciensis sp. nov., adult legs showing segments femur to tarsus, antiaxial view. 9. leg I, left; 10. leg II, left. Scale bar 30 µm." captionText-4="FIGURES 1112. Papillacarus (Vepracarus) acaciensis sp. nov., adult; legs showing segments trochanter to tarsus, antiaxial view. 11. leg III, left; 12. leg IV, left. Scale bar 30 µm." figureDoi-0="http://doi.org/10.5281/zenodo.4570517" figureDoi-1="http://doi.org/10.5281/zenodo.4570519" figureDoi-2="http://doi.org/10.5281/zenodo.4570523" figureDoi-3="http://doi.org/10.5281/zenodo.4570525" figureDoi-4="http://doi.org/10.5281/zenodo.4570529" httpUri-0="https://zenodo.org/record/4570517/files/figure.png" httpUri-1="https://zenodo.org/record/4570519/files/figure.png" httpUri-2="https://zenodo.org/record/4570523/files/figure.png" httpUri-3="https://zenodo.org/record/4570525/files/figure.png" httpUri-4="https://zenodo.org/record/4570529/files/figure.png" pageId="3" pageNumber="542">Figs. 312</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C326654DFFE2470F59EB2C20EB391899" pageId="3" pageNumber="542" type="diagnosis">
<paragraph id="8B8336C6FFE2470F59EB2C20EC181ED5" blockId="3.[151,1437,366,1977]" pageId="3" pageNumber="542">
<emphasis id="B948EAD4FFE2470F59EB2C20EF721EA8" bold="true" box="[151,274,366,392]" pageId="3" pageNumber="542">Diagnosis.</emphasis>
Body size 376400 × 148171. Rostrum rounded. Entire dorsal surface covered with tubercles. Prodorsal, notogastral and neotrichal setae branched and tree-shaped. Sensillus pectinate, with 18 branches. Epimeral setal formula: 7-4-3-3. Genital setae
<emphasis id="B948EAD4FFE2470F58872CF9EC691EF0" box="[507,521,439,464]" italics="true" pageId="3" pageNumber="542">g</emphasis>
<subScript id="17B83483FFE2470F5B752C87EC711EF7" attach="left" box="[521,529,457,471]" fontSize="6" pageId="3" pageNumber="542">6</subScript>
,
<emphasis id="B948EAD4FFE2470F5B632CF9EC4D1EF0" box="[543,557,439,464]" italics="true" pageId="3" pageNumber="542">g</emphasis>
<subScript id="17B83483FFE2470F5B512C87EC551EF7" attach="left" box="[557,565,457,471]" fontSize="6" pageId="3" pageNumber="542">7</subScript>
,
<emphasis id="B948EAD4FFE2470F5B382CF9EC321EF0" box="[580,594,439,464]" italics="true" pageId="3" pageNumber="542">g</emphasis>
<subScript id="17B83483FFE2470F5B2E2C87EC3A1EF7" attach="left" box="[594,602,457,471]" fontSize="6" pageId="3" pageNumber="542">8</subScript>
,
<emphasis id="B948EAD4FFE2470F5B152CF9EC171EF0" box="[617,631,439,464]" italics="true" pageId="3" pageNumber="542">g</emphasis>
<subScript id="17B83483FFE2470F5B0B2C87ECE71EF7" attach="left" box="[631,647,457,471]" fontSize="6" pageId="3" pageNumber="542">10</subScript>
setiform, remaining pairs arboriform. Ano-adanal plates incompletely separated. Anal and adanal setae branched.
</paragraph>
<paragraph id="8B8336C6FFE2470F59BB2CB1EF731D1D" blockId="3.[151,1437,366,1977]" pageId="3" pageNumber="542">
<emphasis id="B948EAD4FFE2470F59BB2CB1EF111D38" box="[199,369,511,536]" italics="true" pageId="3" pageNumber="542">Measurements.</emphasis>
Body length 378 (
<typeStatus id="54878864FFE2470F5B3D2CB1ECC91D3A" box="[577,681,511,538]" pageId="3" pageNumber="542" type="holotype">holotype</typeStatus>
), 376400 (
<specimenCount id="9D3AFD4FFFE2470F5A512CB1EDA11D39" box="[813,961,511,537]" pageId="3" pageNumber="542" type="generic" typeStatus="paratypes">10 paratypes</specimenCount>
); body width 166 (
<typeStatus id="54878864FFE2470F5DEF2CB1EA9B1D3A" box="[1171,1275,511,538]" pageId="3" pageNumber="542" type="holotype">holotype</typeStatus>
), 148171 (
<specimenCount id="9D3AFD4FFFE2470F5C032CB1EF681D1D" pageId="3" pageNumber="542" type="generic" typeStatus="paratypes">10 paratypes</specimenCount>
).
</paragraph>
<paragraph id="8B8336C6FFE2470F59BB2F09EB2C1DA5" blockId="3.[151,1437,366,1977]" pageId="3" pageNumber="542">
<emphasis id="B948EAD4FFE2470F59BB2F09EF2D1D40" box="[199,333,583,608]" italics="true" pageId="3" pageNumber="542">Integument.</emphasis>
Body colour golden brown. Surface of prodorsum and notogaster with thickly punctate tubercles, subcapitulum and epimeral region ornamented with weakly developed tubercles showing thick punctations.
</paragraph>
<paragraph id="8B8336C6FFE2470F59BB2FC1EFA61C85" blockId="3.[151,1437,366,1977]" pageId="3" pageNumber="542">
<emphasis id="B948EAD4FFE2470F59BB2FC1EF241D89" box="[199,324,655,681]" italics="true" pageId="3" pageNumber="542">Prodorsum</emphasis>
(
<figureCitation id="13072A43FFE2470F582F2FC1EFF81D89" box="[339,408,655,681]" captionStart="FIGURE 3" captionStartId="4.[152,255,1989,2013]" captionTargetBox="[297,1295,192,1958]" captionTargetId="figure-22@4.[279,1308,181,1965]" captionTargetPageId="4" captionText="FIGURE 3. Papillacarus (Vepracarus) acaciensis sp. nov., adult dorsal view. Scale bar 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4570517" httpUri="https://zenodo.org/record/4570517/files/figure.png" pageId="3" pageNumber="542">Fig. 3</figureCitation>
). Prodorsum broadly conical, rostrum rounded. All prodorsal setae branched and of varying length. Rostral seta the longest (
<emphasis id="B948EAD4FFE2470F58812FFDEC751DEC" box="[509,533,691,716]" italics="true" pageId="3" pageNumber="542">ro</emphasis>
, 52). Lamellar seta (
<emphasis id="B948EAD4FFE2470F5B802FFDED701DED" box="[764,784,691,717]" italics="true" pageId="3" pageNumber="542">le</emphasis>
, 41), interlamellar seta (
<emphasis id="B948EAD4FFE2470F5D5D2FFDEA571DEC" box="[1057,1079,691,716]" italics="true" pageId="3" pageNumber="542">in</emphasis>
, 45), anterior exobothridial seta (
<emphasis id="B948EAD4FFE2470F59DD2F99EEA81DD0" box="[161,200,727,752]" italics="true" pageId="3" pageNumber="542">exa</emphasis>
, 44), posterior exobothridial seta (
<emphasis id="B948EAD4FFE2470F5B362F99EC111DD0" box="[586,625,727,752]" italics="true" pageId="3" pageNumber="542">exp</emphasis>
, 42). Sensillus (
<emphasis id="B948EAD4FFE2470F5A5B2F99ED5D1DD0" box="[807,829,727,752]" italics="true" pageId="3" pageNumber="542">ss</emphasis>
, 55) pectinate, with 18 long branches on one side, and six small barbs on the opposite side, the branches more or less of equal length at middle, but progressively decreasing towards the anterior and posterior ends. Prodorsal surface ornamented with tubercles. A feebly developed postbothridial transverse band (
<emphasis id="B948EAD4FFE2470F58BB2E0DEFBD1C43" box="[455,477,835,867]" italics="true" pageId="3" pageNumber="542">
S
<subScript id="17B83483FFE2470F58A92E1BEFBD1C43" attach="left" box="[469,477,853,867]" fontSize="6" pageId="3" pageNumber="542">b</subScript>
</emphasis>
) present at the interlamellar area. Weakly developed polygonal sculptures bearing fine punctation present, posterior to
<emphasis id="B948EAD4FFE2470F58802E29EC721CA7" box="[508,530,871,903]" italics="true" pageId="3" pageNumber="542">
S
<subScript id="17B83483FFE2470F5B762E37EC721CA7" attach="left" box="[522,530,889,903]" fontSize="6" pageId="3" pageNumber="542">b</subScript>
</emphasis>
. Lateral prodorsal margin bears distinct tooth like projection, one on either side, above the level of seta
<emphasis id="B948EAD4FFE2470F58E52EC5EFA01C84" box="[409,448,907,932]" italics="true" pageId="3" pageNumber="542">exa</emphasis>
.
</paragraph>
<paragraph id="8B8336C6FFE2470F59BB2EE1EF931B5D" blockId="3.[151,1437,366,1977]" pageId="3" pageNumber="542">
<emphasis id="B948EAD4FFE2470F59BB2EE1EF231CE8" box="[199,323,943,968]" italics="true" pageId="3" pageNumber="542">Notogaster</emphasis>
(
<figureCitation id="13072A43FFE2470F582B2EE1EFA71CE9" box="[343,455,943,969]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="4.[152,255,1989,2013]" captionStartId-1="5.[152,255,1991,2015]" captionTargetBox-0="[297,1295,192,1958]" captionTargetBox-1="[388,1198,191,1958]" captionTargetId-0="figure-22@4.[279,1308,181,1965]" captionTargetId-1="figure-19@5.[373,1214,181,1967]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 3. Papillacarus (Vepracarus) acaciensis sp. nov., adult dorsal view. Scale bar 50 µm." captionText-1="FIGURE 4. Papillacarus (Vepracarus) acaciensis sp. nov., adult ventral view. Scale bar 50 µm." figureDoi-0="http://doi.org/10.5281/zenodo.4570517" figureDoi-1="http://doi.org/10.5281/zenodo.4570519" httpUri-0="https://zenodo.org/record/4570517/files/figure.png" httpUri-1="https://zenodo.org/record/4570519/files/figure.png" pageId="3" pageNumber="542">Figs. 34</figureCitation>
,
<figureCitation id="13072A43FFE2470F58AB2EE0EC651CE9" box="[471,517,942,969]" captionStart="FIGURES 58" captionStartId="6.[152,270,1964,1988]" captionTargetBox="[184,1399,199,1926]" captionTargetId="figure-22@6.[172,1416,181,1940]" captionTargetPageId="6" captionText="FIGURES 58. Papillacarus (Vepracarus) acaciensis sp. nov., adult; 5. notogastral seta; 6. subcapitulum, ventral view; 7. chelicera, left; 8. notogastral tubercles. Scale bar 30 µm." figureDoi="http://doi.org/10.5281/zenodo.4570523" httpUri="https://zenodo.org/record/4570523/files/figure.png" pageId="3" pageNumber="542">5, 8</figureCitation>
). Anterior notogastral margin straight. Entire notogastral surface covered with densely punctate tubercles, except the area lying just below the dorsosejugal suture where thick punctations present. Thick punctations are also seen scattered on the notogaster. Sixteen pairs of primary notogastral setae and more than 18 pairs of additional neotrichal setae present, all branched with 59 branches and show variation in length as:
<emphasis id="B948EAD4FFE2470F5CFD2955EBED1B14" box="[1409,1421,1051,1076]" italics="true" pageId="3" pageNumber="542">c</emphasis>
<subScript id="17B83483FFE2470F5CF12963EBF51B1B" attach="left" box="[1421,1429,1069,1083]" fontSize="6" pageId="3" pageNumber="542">1</subScript>
,
<emphasis id="B948EAD4FFE2470F59EB2971EEC51B79" box="[151,165,1087,1113]" italics="true" pageId="3" pageNumber="542">d</emphasis>
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,
<emphasis id="B948EAD4FFE2470F59C02971EEA81B78" box="[188,200,1087,1112]" italics="true" pageId="3" pageNumber="542">e</emphasis>
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,
<emphasis id="B948EAD4FFE2470F59A32971EE8D1B78" box="[223,237,1087,1112]" italics="true" pageId="3" pageNumber="542">p</emphasis>
<subScript id="17B83483FFE2470F5991291FEE951B7F" attach="left" box="[237,245,1105,1119]" fontSize="6" pageId="3" pageNumber="542">2</subScript>
,
<emphasis id="B948EAD4FFE2470F58782971EF721B79" box="[260,274,1087,1113]" italics="true" pageId="3" pageNumber="542">h</emphasis>
<subScript id="17B83483FFE2470F586E291FEF7A1B7F" attach="left" box="[274,282,1105,1119]" fontSize="6" pageId="3" pageNumber="542">2</subScript>
(2632);
<emphasis id="B948EAD4FFE2470F58F52971EFF51B78" box="[393,405,1087,1112]" italics="true" pageId="3" pageNumber="542">c</emphasis>
<subScript id="17B83483FFE2470F58EA291FEFFE1B7F" attach="right" box="[406,414,1105,1119]" fontSize="6" pageId="3" pageNumber="542">2</subScript>
,
<emphasis id="B948EAD4FFE2470F58D12971EFDB1B79" box="[429,443,1087,1113]" italics="true" pageId="3" pageNumber="542">d</emphasis>
<subScript id="17B83483FFE2470F58C7291FEFA31B7F" attach="left" box="[443,451,1105,1119]" fontSize="6" pageId="3" pageNumber="542">2</subScript>
,
<emphasis id="B948EAD4FFE2470F58AD2971EFB91B79" box="[465,473,1087,1113]" italics="true" pageId="3" pageNumber="542">f</emphasis>
<subScript id="17B83483FFE2470F58A5291FEF811B7F" attach="left" box="[473,481,1105,1119]" fontSize="6" pageId="3" pageNumber="542">1</subScript>
,
<emphasis id="B948EAD4FFE2470F588C2971EF9E1B79" box="[496,510,1087,1113]" italics="true" pageId="3" pageNumber="542">h</emphasis>
<subScript id="17B83483FFE2470F5882291FEC661B7F" attach="left" box="[510,518,1105,1119]" fontSize="6" pageId="3" pageNumber="542">1</subScript>
,
<emphasis id="B948EAD4FFE2470F5B692971EC431B78" box="[533,547,1087,1112]" italics="true" pageId="3" pageNumber="542">p</emphasis>
<subScript id="17B83483FFE2470F5B5F291FEC4B1B7F" attach="left" box="[547,555,1105,1119]" fontSize="6" pageId="3" pageNumber="542">1</subScript>
,
<emphasis id="B948EAD4FFE2470F5B452971EC251B78" box="[569,581,1087,1112]" italics="true" pageId="3" pageNumber="542">c</emphasis>
<subScript id="17B83483FFE2470F5B3A291FEC2E1B7F" attach="right" box="[582,590,1105,1119]" fontSize="6" pageId="3" pageNumber="542">3</subScript>
,
<emphasis id="B948EAD4FFE2470F5B202971EC0A1B79" box="[604,618,1087,1113]" italics="true" pageId="3" pageNumber="542">d</emphasis>
<subScript id="17B83483FFE2470F5B16291FEC121B7F" attach="left" box="[618,626,1105,1119]" fontSize="6" pageId="3" pageNumber="542">3</subScript>
,
<emphasis id="B948EAD4FFE2470F5BFD2971ECED1B78" box="[641,653,1087,1112]" italics="true" pageId="3" pageNumber="542">e</emphasis>
<subScript id="17B83483FFE2470F5BF2291FECF61B7F" attach="right" box="[654,662,1105,1119]" fontSize="6" pageId="3" pageNumber="542">2</subScript>
,
<emphasis id="B948EAD4FFE2470F5BD82971ECCC1B79" box="[676,684,1087,1113]" italics="true" pageId="3" pageNumber="542">f</emphasis>
<subScript id="17B83483FFE2470F5BD0291FECD41B7F" attach="left" box="[684,692,1105,1119]" fontSize="6" pageId="3" pageNumber="542">2</subScript>
,
<emphasis id="B948EAD4FFE2470F5BBF2971ECB11B79" box="[707,721,1087,1113]" italics="true" pageId="3" pageNumber="542">h</emphasis>
<subScript id="17B83483FFE2470F5BAD291FECB91B7F" attach="left" box="[721,729,1105,1119]" fontSize="6" pageId="3" pageNumber="542">3</subScript>
,
<emphasis id="B948EAD4FFE2470F5B942971EC961B78" box="[744,758,1087,1112]" italics="true" pageId="3" pageNumber="542">p</emphasis>
<subScript id="17B83483FFE2470F5B8A291FEC9E1B7F" attach="left" box="[758,766,1105,1119]" fontSize="6" pageId="3" pageNumber="542">3</subScript>
(3646). Neotrichal setae arise after
<emphasis id="B948EAD4FFE2470F5DE22971EAC61B79" box="[1182,1190,1087,1113]" italics="true" pageId="3" pageNumber="542">f</emphasis>
setae. Lyrifissures
<emphasis id="B948EAD4FFE2470F5C032971EBF51B78" box="[1407,1429,1087,1112]" italics="true" pageId="3" pageNumber="542">ia</emphasis>
,
<emphasis id="B948EAD4FFE2470F59EB292DEED31B5C" box="[151,179,1123,1148]" italics="true" pageId="3" pageNumber="542">im</emphasis>
,
<emphasis id="B948EAD4FFE2470F59BD292DEEB71B5D" box="[193,215,1123,1149]" italics="true" pageId="3" pageNumber="542">ih</emphasis>
and
<emphasis id="B948EAD4FFE2470F5871292DEF431B5C" box="[269,291,1123,1148]" italics="true" pageId="3" pageNumber="542">ip</emphasis>
visible as figured.
</paragraph>
<paragraph id="8B8336C6FFE2470F59BB29C9EB0A1A75" blockId="3.[151,1437,366,1977]" pageId="3" pageNumber="542">
<taxonomicName id="4C3C4D45FFE2470F59BB29C9EF361B81" authorityName="Krefft" authorityYear="1871" box="[199,342,1159,1185]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="3" pageNumber="542" phylum="Nematoda" rank="genus">
<emphasis id="B948EAD4FFE2470F59BB29C9EF361B81" box="[199,342,1159,1185]" italics="true" pageId="3" pageNumber="542">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="13072A43FFE2470F581929C9EFD31B81" box="[357,435,1159,1185]" captionStart="FIGURE 4" captionStartId="5.[152,255,1991,2015]" captionTargetBox="[388,1198,191,1958]" captionTargetId="figure-19@5.[373,1214,181,1967]" captionTargetPageId="5" captionText="FIGURE 4. Papillacarus (Vepracarus) acaciensis sp. nov., adult ventral view. Scale bar 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4570519" httpUri="https://zenodo.org/record/4570519/files/figure.png" pageId="3" pageNumber="542">Figs. 4</figureCitation>
,
<figureCitation id="13072A43FFE2470F58C229C9EF8B1B82" box="[446,491,1159,1186]" captionStart="FIGURES 58" captionStartId="6.[152,270,1964,1988]" captionTargetBox="[184,1399,199,1926]" captionTargetId="figure-22@6.[172,1416,181,1940]" captionTargetPageId="6" captionText="FIGURES 58. Papillacarus (Vepracarus) acaciensis sp. nov., adult; 5. notogastral seta; 6. subcapitulum, ventral view; 7. chelicera, left; 8. notogastral tubercles. Scale bar 30 µm." figureDoi="http://doi.org/10.5281/zenodo.4570523" httpUri="https://zenodo.org/record/4570523/files/figure.png" pageId="3" pageNumber="542">67</figureCitation>
). Subcapitulum longer than wide (100×67). Less prominent tubercles with fine punctations present on mentum. Five pairs of subcapitular setae detected,
<emphasis id="B948EAD4FFE2470F5ADC29E5EDCE1BE4" box="[928,942,1195,1220]" italics="true" pageId="3" pageNumber="542">a</emphasis>
(27) setiform, smooth and pointed apically;
<emphasis id="B948EAD4FFE2470F59EB2981EEC51BC9" box="[151,165,1231,1257]" italics="true" pageId="3" pageNumber="542">h</emphasis>
barbed (19),
<emphasis id="B948EAD4FFE2470F58422981EF321BC8" box="[318,338,1231,1256]" italics="true" pageId="3" pageNumber="542">m</emphasis>
<subScript id="17B83483FFE2470F582E29AFEF3A1BCF" attach="left" box="[338,346,1249,1263]" fontSize="6" pageId="3" pageNumber="542">1</subScript>
,
<emphasis id="B948EAD4FFE2470F58152981EF1D1BC8" box="[361,381,1231,1256]" italics="true" pageId="3" pageNumber="542">m</emphasis>
<subScript id="17B83483FFE2470F580129AFEFE51BCF" attach="left" box="[381,389,1249,1263]" fontSize="6" pageId="3" pageNumber="542">2</subScript>
,
<emphasis id="B948EAD4FFE2470F58E82981EFC81BC8" box="[404,424,1231,1256]" italics="true" pageId="3" pageNumber="542">m</emphasis>
<subScript id="17B83483FFE2470F58D529AFEFD11BCF" attach="left" box="[425,433,1249,1263]" fontSize="6" pageId="3" pageNumber="542">3</subScript>
(1820) setiform, and barbed. Three pairs of smooth adoral setae present:
<emphasis id="B948EAD4FFE2470F5D812981EB761BC8" box="[1277,1302,1231,1256]" italics="true" pageId="3" pageNumber="542">or</emphasis>
<subScript id="17B83483FFE2470F5C6A29AFEB7E1BCF" attach="left" box="[1302,1310,1249,1263]" fontSize="6" pageId="3" pageNumber="542">1</subScript>
(14) wider proximally, tapering apically and with a blunt tip;
<emphasis id="B948EAD4FFE2470F5BA429BDEC911A2C" box="[728,753,1267,1292]" italics="true" pageId="3" pageNumber="542">or</emphasis>
<subScript id="17B83483FFE2470F5B8D284BEC991A33" attach="left" box="[753,761,1285,1299]" fontSize="6" pageId="3" pageNumber="542">2</subScript>
long (15) and
<emphasis id="B948EAD4FFE2470F5ADE29BDEDDB1A2C" box="[930,955,1267,1292]" italics="true" pageId="3" pageNumber="542">or</emphasis>
<subScript id="17B83483FFE2470F5AC7284BEDA31A33" attach="left" box="[955,963,1285,1299]" fontSize="6" pageId="3" pageNumber="542">3</subScript>
small (11), both with blunt tip. Rutellum with 36 blunt dendrites. Palpal (39) surface punctated, palpal setal formula 010110 (+1
<emphasis id="B948EAD4FFE2470F5DC52859EAAD1A10" box="[1209,1229,1303,1328]" italics="true" pageId="3" pageNumber="542">ω</emphasis>
), solenidion blunt distally. Chelicera (117) with 35 teeth, seta
<emphasis id="B948EAD4FFE2470F5BF72875ECD31A75" box="[651,691,1339,1365]" italics="true" pageId="3" pageNumber="542">cha</emphasis>
(4) short, thorn-like, seta
<emphasis id="B948EAD4FFE2470F5AAB2875ED9F1A75" box="[983,1023,1339,1365]" italics="true" pageId="3" pageNumber="542">chb</emphasis>
(34) long, setiform and smooth.
</paragraph>
<paragraph id="8B8336C6FFE2470F59BB2811EF111AE1" blockId="3.[151,1437,366,1977]" pageId="3" pageNumber="542">
<emphasis id="B948EAD4FFE2470F59BB2811EF1F1A58" box="[199,383,1375,1401]" italics="true" pageId="3" pageNumber="542">Epimeral region</emphasis>
(
<figureCitation id="13072A43FFE2470F58F22811EFB21A59" box="[398,466,1375,1401]" captionStart="FIGURE 4" captionStartId="5.[152,255,1991,2015]" captionTargetBox="[388,1198,191,1958]" captionTargetId="figure-19@5.[373,1214,181,1967]" captionTargetPageId="5" captionText="FIGURE 4. Papillacarus (Vepracarus) acaciensis sp. nov., adult ventral view. Scale bar 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4570519" httpUri="https://zenodo.org/record/4570519/files/figure.png" pageId="3" pageNumber="542">Fig. 4</figureCitation>
). Epimeral plates distinct, surface with punctations. Epimeres I and II carry neotrichal setae, all branched, epimeral setal formula 7-4-3-3. Setae
<emphasis id="B948EAD4FFE2470F5A5628CDED941ABD" box="[810,1012,1411,1437]" italics="true" pageId="3" pageNumber="542">1a-e, 2a, 3a, 4a-c</emphasis>
(715) shorter and thinner than
<emphasis id="B948EAD4FFE2470F5C1428CDEF6A1AE1" italics="true" pageId="3" pageNumber="542">1f-g, 2b-d, 3b-c</emphasis>
(2134).
</paragraph>
<paragraph id="8B8336C6FFE2470F59BB2885EA871955" blockId="3.[151,1437,366,1977]" pageId="3" pageNumber="542">
<emphasis id="B948EAD4FFE2470F59BB2885EFF01AC4" box="[199,400,1483,1509]" italics="true" pageId="3" pageNumber="542">Anogenital region</emphasis>
(
<figureCitation id="13072A43FFE2470F58E22885EF801AC5" box="[414,480,1483,1509]" captionStart="FIGURE 4" captionStartId="5.[152,255,1991,2015]" captionTargetBox="[388,1198,191,1958]" captionTargetId="figure-19@5.[373,1214,181,1967]" captionTargetPageId="5" captionText="FIGURE 4. Papillacarus (Vepracarus) acaciensis sp. nov., adult ventral view. Scale bar 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4570519" httpUri="https://zenodo.org/record/4570519/files/figure.png" pageId="3" pageNumber="542">Fig. 4</figureCitation>
). Transverse suture divides each genital plate into smaller anterior and larger posterior sections. Genital setae
<emphasis id="B948EAD4FFE2470F58E028A1EFCA1928" box="[412,426,1519,1544]" italics="true" pageId="3" pageNumber="542">g</emphasis>
<subScript id="17B83483FFE2470F58D62B4FEFD2192F" attach="left" box="[426,434,1537,1551]" fontSize="6" pageId="3" pageNumber="542">6</subScript>
,
<emphasis id="B948EAD4FFE2470F58BE28A1EFB01928" box="[450,464,1519,1544]" italics="true" pageId="3" pageNumber="542">g</emphasis>
<subScript id="17B83483FFE2470F58AC2B4FEFB8192F" attach="left" box="[464,472,1537,1551]" fontSize="6" pageId="3" pageNumber="542">7</subScript>
,
<emphasis id="B948EAD4FFE2470F589B28A1EF951928" box="[487,501,1519,1544]" italics="true" pageId="3" pageNumber="542">g</emphasis>
<subScript id="17B83483FFE2470F58892B4FEF9D192F" attach="left" box="[501,509,1537,1551]" fontSize="6" pageId="3" pageNumber="542">8</subScript>
,
<emphasis id="B948EAD4FFE2470F5B7128A1EC7B1928" box="[525,539,1519,1544]" italics="true" pageId="3" pageNumber="542">g</emphasis>
<subScript id="17B83483FFE2470F5B672B4FEC4B192F" attach="left" box="[539,555,1537,1551]" fontSize="6" pageId="3" pageNumber="542">10</subScript>
setiform and smooth (1719), other setae branched (1114). Anal and adanal plates separated along the antero-median region while the postero-median regions fused. Two pairs of anal (
<emphasis id="B948EAD4FFE2470F5C3C2B5DEB3C190C" box="[1344,1372,1555,1580]" italics="true" pageId="3" pageNumber="542">an</emphasis>
<subScript id="17B83483FFE2470F5C202B6BEB041913" attach="left" box="[1372,1380,1573,1587]" fontSize="6" pageId="3" pageNumber="542">1</subScript>
,
<emphasis id="B948EAD4FFE2470F5C0D2B5DEBED190C" box="[1393,1421,1555,1580]" italics="true" pageId="3" pageNumber="542">an</emphasis>
<subScript id="17B83483FFE2470F5CF12B6BEBF51913" attach="left" box="[1421,1429,1573,1587]" fontSize="6" pageId="3" pageNumber="542">2</subScript>
, 2521) and four pairs of adanal setae (
<emphasis id="B948EAD4FFE2470F5B352B79EC051971" box="[585,613,1591,1617]" italics="true" pageId="3" pageNumber="542">ad</emphasis>
<subScript id="17B83483FFE2470F5B192B07EC1A1977" attach="left" box="[613,634,1609,1623]" fontSize="6" pageId="3" pageNumber="542">1-4</subScript>
, 3042) present, all possess several long branches. Adanal setae longer than the anal setae. Both the genital and ano-adanal plates bear punctation. Lyrifissure
<emphasis id="B948EAD4FFE2470F5D1E2B15EAE61955" box="[1122,1158,1627,1653]" italics="true" pageId="3" pageNumber="542">iad</emphasis>
distinct.
</paragraph>
<paragraph id="8B8336C6FFE2470F59BB2B31EB391899" blockId="3.[151,1437,366,1977]" pageId="3" pageNumber="542">
<emphasis id="B948EAD4FFE2470F59BB2B31EE9C19B8" box="[199,252,1663,1688]" italics="true" pageId="3" pageNumber="542">Legs</emphasis>
(
<figureCitation id="13072A43FFE2470F58762B31EFE519B9" box="[266,389,1663,1689]" captionStart-0="FIGURES 910" captionStart-1="FIGURES 1112" captionStartId-0="7.[152,270,1881,1905]" captionStartId-1="8.[152,270,1961,1985]" captionTargetBox-0="[171,1417,196,1842]" captionTargetBox-1="[267,1313,203,1920]" captionTargetId-0="figure-19@7.[151,1436,181,1857]" captionTargetId-1="figure-22@8.[248,1339,181,1937]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 910. Papillacarus (Vepracarus) acaciensis sp. nov., adult legs showing segments femur to tarsus, antiaxial view. 9. leg I, left; 10. leg II, left. Scale bar 30 µm." captionText-1="FIGURES 1112. Papillacarus (Vepracarus) acaciensis sp. nov., adult; legs showing segments trochanter to tarsus, antiaxial view. 11. leg III, left; 12. leg IV, left. Scale bar 30 µm." figureDoi-0="http://doi.org/10.5281/zenodo.4570525" figureDoi-1="http://doi.org/10.5281/zenodo.4570529" httpUri-0="https://zenodo.org/record/4570525/files/figure.png" httpUri-1="https://zenodo.org/record/4570529/files/figure.png" pageId="3" pageNumber="542">Figs. 912</figureCitation>
). All legs bearing a single claw each, and claws with a small basoventral tooth. Segments of all legs finely punctate. Trochanters I &amp; II devoid of setae while III &amp; IV with two setae each. Femur I with a posteroventral tooth and an antero-ventral flange like expansion, femora III and IV each with antero-ventral tooth. On genu I, two solenidia (
<emphasis id="B948EAD4FFE2470F58242BA5EFF3182B" box="[344,403,1771,1803]" italics="true" pageId="3" pageNumber="542">
σ
<subScript id="17B83483FFE2470F581A2BB3EF0E182B" attach="left" box="[358,366,1789,1803]" fontSize="6" pageId="3" pageNumber="542">1</subScript>
, σ
<subScript id="17B83483FFE2470F58F72BB3EFF3182B" attach="left" box="[395,403,1789,1803]" fontSize="6" pageId="3" pageNumber="542">2</subScript>
</emphasis>
) present, the latter longer than the former and tapering distally. Genua II, III and IV with a single solenidion each. A single solenidion (
<emphasis id="B948EAD4FFE2470F5BF42A41ECF71808" box="[648,663,1807,1832]" italics="true" pageId="3" pageNumber="542">φ</emphasis>
) present on tibia of all legs, except that of leg IV. Tarsus I bears two solenidia (
<emphasis id="B948EAD4FFE2470F58702A7DEF311873" box="[268,337,1843,1875]" italics="true" pageId="3" pageNumber="542">
ω
<subScript id="17B83483FFE2470F585C2A0BEF481873" attach="left" box="[288,296,1861,1875]" fontSize="6" pageId="3" pageNumber="542">1</subScript>
, ω
<subScript id="17B83483FFE2470F58352A0BEF311873" attach="left" box="[329,337,1861,1875]" fontSize="6" pageId="3" pageNumber="542">2</subScript>
</emphasis>
) while tarsus II possesses a single solenidion. The remaining tarsi devoid of solenidia. Famulus (
<emphasis id="B948EAD4FFE2470F5CF42A7DEBF3186C" box="[1416,1427,1843,1868]" italics="true" pageId="3" pageNumber="542">ɛ</emphasis>
) of tarsus I short and blunt and placed distal to the stout solenidion
<emphasis id="B948EAD4FFE2470F5AF42A19EDAE1857" box="[904,974,1879,1911]" italics="true" pageId="3" pageNumber="542">
ω
<subScript id="17B83483FFE2470F5AE02A27EDC41857" attach="left" box="[924,932,1897,1911]" fontSize="6" pageId="3" pageNumber="542">1</subScript>
, ω
<subScript id="17B83483FFE2470F5ABA2A27EDAE1857" attach="left" box="[966,974,1897,1911]" fontSize="6" pageId="3" pageNumber="542">2</subScript>
</emphasis>
of tarsus I long and tapering. Solenidion on tibial segment (
<emphasis id="B948EAD4FFE2470F58162A35EF1918B4" box="[362,377,1915,1940]" italics="true" pageId="3" pageNumber="542">φ</emphasis>
) of legs III and IV thick with blunt tip while those of other legs thin and tapering. Chaetotaxy of legs IIV: 0-5-2(2)-3(1)-17(2); 0-5-2(1)-2(1)-12(1); 2-4-2(1)-1(1)-11(0); 2-3-2(1)-3(0)-11(0) (see
<tableCitation id="C6BE037DFFE2470F5D812AD1EB2E1899" box="[1277,1358,1951,1977]" captionStart="TABLE 1" captionStartId="9.[151,239,151,175]" captionTargetPageId="9" captionText="TABLE 1. Setation and solenidia on legs in adult specimens of Papillacarus (Vepracarus) acaciensis sp. nov." httpUri="http://table.plazi.org/id/DF43664EFFE8470559EB2DD9EB4E1F8F" pageId="3" pageNumber="542" tableUuid="DF43664EFFE8470559EB2DD9EB4E1F8F">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<caption id="DF43664EFFE5470859E42A8BEA0618FE" ID-DOI="http://doi.org/10.5281/zenodo.4570517" ID-Zenodo-Dep="4570517" box="[152,1126,1989,2014]" httpUri="https://zenodo.org/record/4570517/files/figure.png" pageId="4" pageNumber="543" startId="4.[152,255,1989,2013]" targetBox="[297,1295,192,1958]" targetPageId="4">
<paragraph id="8B8336C6FFE5470859E42A8BEA0618FE" blockId="4.[152,1126,1989,2014]" box="[152,1126,1989,2014]" pageId="4" pageNumber="543">
<emphasis id="B948EAD4FFE5470859E42A8BEF7818FD" bold="true" box="[152,280,1989,2013]" pageId="4" pageNumber="543">FIGURE 3.</emphasis>
<taxonomicName id="4C3C4D45FFE5470858632A8BECFD18FD" authority="Arun &amp; Ramani, 2020" authorityName="Arun &amp; Ramani" authorityYear="2020" box="[287,669,1989,2013]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="543" phylum="Arthropoda" rank="species" species="acaciensis" status="sp. nov." subGenus="Vepracarus">
<emphasis id="B948EAD4FFE5470858632A8BEFC218FD" box="[287,418,1989,2013]" italics="true" pageId="4" pageNumber="543">Papillacarus</emphasis>
(
<emphasis id="B948EAD4FFE5470858CC2A88EC4318FD" box="[432,547,1990,2013]" italics="true" pageId="4" pageNumber="543">Vepracarus</emphasis>
)
<emphasis id="B948EAD4FFE547085B4F2A88ECFD18FD" box="[563,669,1990,2013]" italics="true" pageId="4" pageNumber="543">acaciensis</emphasis>
</taxonomicName>
<emphasis id="B948EAD4FFE547085BDE2A8BEC9318FD" bold="true" box="[674,755,1989,2013]" pageId="4" pageNumber="543">
<taxonomicNameLabel id="A27B57AFFFE547085BDE2A8BEC9318FD" box="[674,755,1989,2013]" pageId="4" pageNumber="543" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult dorsal view. Scale bar 50 µm.
</paragraph>
</caption>
<caption id="DF43664EFFE4470959E42A89EA0F18C0" ID-DOI="http://doi.org/10.5281/zenodo.4570519" ID-Zenodo-Dep="4570519" box="[152,1135,1991,2016]" httpUri="https://zenodo.org/record/4570519/files/figure.png" pageId="5" pageNumber="544" startId="5.[152,255,1991,2015]" targetBox="[388,1198,191,1958]" targetPageId="5">
<paragraph id="8B8336C6FFE4470959E42A89EA0F18C0" blockId="5.[152,1135,1991,2016]" box="[152,1135,1991,2016]" pageId="5" pageNumber="544">
<emphasis id="B948EAD4FFE4470959E42A89EF7818FF" bold="true" box="[152,280,1991,2015]" pageId="5" pageNumber="544">FIGURE 4.</emphasis>
<taxonomicName id="4C3C4D45FFE4470958632A86ECFD18FF" authority="Arun &amp; Ramani, 2020" authorityName="Arun &amp; Ramani" authorityYear="2020" box="[287,669,1992,2016]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="544" phylum="Arthropoda" rank="species" species="acaciensis" status="sp. nov." subGenus="Vepracarus">
<emphasis id="B948EAD4FFE4470958632A86EFC218C0" box="[287,418,1992,2016]" italics="true" pageId="5" pageNumber="544">Papillacarus</emphasis>
(
<emphasis id="B948EAD4FFE4470958CC2A86EC4318FF" box="[432,547,1992,2015]" italics="true" pageId="5" pageNumber="544">Vepracarus</emphasis>
)
<emphasis id="B948EAD4FFE447095B4F2A86ECFD18FF" box="[563,669,1992,2015]" italics="true" pageId="5" pageNumber="544">acaciensis</emphasis>
</taxonomicName>
<emphasis id="B948EAD4FFE447095BDE2A86EC9318C0" bold="true" box="[674,755,1992,2016]" pageId="5" pageNumber="544">
<taxonomicNameLabel id="A27B57AFFFE447095BDE2A86EC9318C0" box="[674,755,1992,2016]" pageId="5" pageNumber="544" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult ventral view. Scale bar 50 µm.
</paragraph>
</caption>
<caption id="DF43664EFFE7470A59E42AE2ECB418C8" ID-DOI="http://doi.org/10.5281/zenodo.4570523" ID-Zenodo-Dep="4570523" httpUri="https://zenodo.org/record/4570523/files/figure.png" pageId="6" pageNumber="545" startId="6.[152,270,1964,1988]" targetBox="[184,1399,199,1926]" targetPageId="6">
<paragraph id="8B8336C6FFE7470A59E42AE2ECB418C8" blockId="6.[151,1436,1964,2024]" pageId="6" pageNumber="545">
<emphasis id="B948EAD4FFE7470A59E42AE2EF2318E4" bold="true" box="[152,323,1964,1988]" pageId="6" pageNumber="545">FIGURES 58.</emphasis>
<taxonomicName id="4C3C4D45FFE7470A58302AE2ECAE18E4" authority="Arun &amp; Ramani, 2020" authorityName="Arun &amp; Ramani" authorityYear="2020" box="[332,718,1964,1988]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="545" phylum="Arthropoda" rank="species" species="acaciensis" status="sp. nov." subGenus="Vepracarus">
<emphasis id="B948EAD4FFE7470A58302AE2EFAF18E4" box="[332,463,1964,1988]" italics="true" pageId="6" pageNumber="545">Papillacarus</emphasis>
(
<emphasis id="B948EAD4FFE7470A58A32AE3EC3218E4" box="[479,594,1965,1988]" italics="true" pageId="6" pageNumber="545">Vepracarus</emphasis>
)
<emphasis id="B948EAD4FFE7470A5B182AE3ECAE18E4" box="[612,718,1965,1988]" italics="true" pageId="6" pageNumber="545">acaciensis</emphasis>
</taxonomicName>
<emphasis id="B948EAD4FFE7470A5BAA2AE2ED4918E4" bold="true" box="[726,809,1964,1988]" pageId="6" pageNumber="545">
<taxonomicNameLabel id="A27B57AFFFE7470A5BAA2AE2ED4918E4" box="[726,809,1964,1988]" pageId="6" pageNumber="545" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult; 5. notogastral seta; 6. subcapitulum, ventral view; 7. chelicera, left; 8. notogastral tubercles. Scale bar 30 µm.
</paragraph>
</caption>
<caption id="DF43664EFFE6470B59E42A17EC3E18B6" ID-DOI="http://doi.org/10.5281/zenodo.4570525" ID-Zenodo-Dep="4570525" httpUri="https://zenodo.org/record/4570525/files/figure.png" pageId="7" pageNumber="546" startId="7.[152,270,1881,1905]" targetBox="[171,1417,196,1842]" targetPageId="7">
<paragraph id="8B8336C6FFE6470B59E42A17EC3E18B6" blockId="7.[151,1437,1881,1942]" pageId="7" pageNumber="546">
<emphasis id="B948EAD4FFE6470B59E42A17EF2E1851" bold="true" box="[152,334,1881,1905]" pageId="7" pageNumber="546">FIGURES 910.</emphasis>
<taxonomicName id="4C3C4D45FFE6470B58292A17ECB31851" authority="Arun &amp; Ramani, 2020" authorityName="Arun &amp; Ramani" authorityYear="2020" box="[341,723,1881,1906]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="546" phylum="Arthropoda" rank="species" species="acaciensis" status="sp. nov." subGenus="Vepracarus">
<emphasis id="B948EAD4FFE6470B58292A17EFB81851" box="[341,472,1881,1905]" italics="true" pageId="7" pageNumber="546">Papillacarus</emphasis>
(
<emphasis id="B948EAD4FFE6470B589B2A14EC3A1851" box="[487,602,1882,1905]" italics="true" pageId="7" pageNumber="546">Vepracarus</emphasis>
)
<emphasis id="B948EAD4FFE6470B5B152A14ECB31851" box="[617,723,1882,1905]" italics="true" pageId="7" pageNumber="546">acaciensis</emphasis>
</taxonomicName>
<emphasis id="B948EAD4FFE6470B5BA52A14ED4A1852" bold="true" box="[729,810,1882,1906]" pageId="7" pageNumber="546">
<taxonomicNameLabel id="A27B57AFFFE6470B5BA52A14ED4A1852" box="[729,810,1882,1906]" pageId="7" pageNumber="546" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult legs showing segments femur to tarsus, antiaxial view. 9. leg I, left; 10. leg II, left. Scale bar 30 µm.
</paragraph>
</caption>
<caption id="DF43664EFFE9470459E42AE7ECDE18C6" ID-DOI="http://doi.org/10.5281/zenodo.4570529" ID-Zenodo-Dep="4570529" httpUri="https://zenodo.org/record/4570529/files/figure.png" pageId="8" pageNumber="547" startId="8.[152,270,1961,1985]" targetBox="[267,1313,203,1920]" targetPageId="8">
<paragraph id="8B8336C6FFE9470459E42AE7ECDE18C6" blockId="8.[151,1436,1961,2022]" pageId="8" pageNumber="547">
<emphasis id="B948EAD4FFE9470459E42AE7EF3918E1" bold="true" box="[152,345,1961,1985]" pageId="8" pageNumber="547">FIGURES 1112.</emphasis>
<taxonomicName id="4C3C4D45FFE94704581C2AE7ECBE18E1" authority="Arun &amp; Ramani, 2020" authorityName="Arun &amp; Ramani" authorityYear="2020" box="[352,734,1961,1986]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="547" phylum="Arthropoda" rank="species" species="acaciensis" status="sp. nov." subGenus="Vepracarus">
<emphasis id="B948EAD4FFE94704581C2AE7EF8318E1" box="[352,483,1961,1985]" italics="true" pageId="8" pageNumber="547">Papillacarus</emphasis>
(
<emphasis id="B948EAD4FFE94704588E2AE4EC0518E1" box="[498,613,1962,1985]" italics="true" pageId="8" pageNumber="547">Vepracarus</emphasis>
)
<emphasis id="B948EAD4FFE947045B082AE4ECBE18E1" box="[628,734,1962,1985]" italics="true" pageId="8" pageNumber="547">acaciensis</emphasis>
</taxonomicName>
<emphasis id="B948EAD4FFE947045B992AE4ED5618E2" bold="true" box="[741,822,1962,1986]" pageId="8" pageNumber="547">
<taxonomicNameLabel id="A27B57AFFFE947045B992AE4ED5618E2" box="[741,822,1962,1986]" pageId="8" pageNumber="547" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult; legs showing segments trochanter to tarsus, antiaxial view. 11. leg III, left; 12. leg IV, left. Scale bar 30 µm.
</paragraph>
</caption>
<caption id="DF43664EFFE8470559EB2DD9EB4E1F8F" ID-Table-UUID="DF43664EFFE8470559EB2DD9EB4E1F8F" box="[151,1326,151,175]" httpUri="http://table.plazi.org/id/DF43664EFFE8470559EB2DD9EB4E1F8F" pageId="9" pageNumber="548" targetBox="[166,1359,191,377]" targetIsTable="true" targetPageId="9">
<paragraph id="8B8336C6FFE8470559EB2DD9EB4E1F8F" blockId="9.[151,1326,151,175]" box="[151,1326,151,175]" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE8470559EB2DD9EF6A1F8F" bold="true" box="[151,266,151,175]" pageId="9" pageNumber="548">TABLE 1.</emphasis>
Setation and solenidia on legs in adult specimens of
<taxonomicName id="4C3C4D45FFE847055A3C2DD9EAB11F8F" authority="Arun &amp; Ramani, 2020" authorityName="Arun &amp; Ramani" authorityYear="2020" box="[832,1233,151,175]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="acaciensis" status="sp. nov." subGenus="Vepracarus">
<emphasis id="B948EAD4FFE847055A3C2DD9EDAA1F8F" box="[832,970,151,175]" italics="true" pageId="9" pageNumber="548">Papillacarus</emphasis>
(
<emphasis id="B948EAD4FFE847055AA52DD6EA321F8F" box="[985,1106,152,175]" italics="true" pageId="9" pageNumber="548">Vepracarus</emphasis>
)
<emphasis id="B948EAD4FFE847055D1E2DD6EAB11F8F" box="[1122,1233,152,175]" italics="true" pageId="9" pageNumber="548">acaciensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57AFFFE847055DA42DD9EB4E1F8F" box="[1240,1326,151,175]" pageId="9" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B8336C6FFE8470559DA2D8EEA0F1E59" pageId="9" pageNumber="548">
<table id="F93CC466FFE8B8F359DA2DF1EB2F1E59" box="[166,1359,191,377]" gridcols="7" gridrows="5" pageId="9" pageNumber="548">
<tr id="350C3484FFE8B8F359DA2DF1EB2F1FF8" box="[166,1359,191,216]" gridrow="0" pageId="9" pageNumber="548" rowspan-6="1">
<th id="76DD5DF8FFE8B8F359DA2DF1EEAD1FF8" box="[166,205,191,216]" gridcol="0" gridrow="0" pageId="9" pageNumber="548">Leg</th>
<th id="76DD5DF8FFE8B8F3598D2DF1EF3F1FF8" box="[241,351,191,216]" gridcol="1" gridrow="0" pageId="9" pageNumber="548">Trochanter</th>
<th id="76DD5DF8FFE8B8F358F22DF1EC6D1FF8" box="[398,525,191,216]" gridcol="2" gridrow="0" pageId="9" pageNumber="548">Femur</th>
<th id="76DD5DF8FFE8B8F35B3E2DF1ECD11FF8" box="[578,689,191,216]" gridcol="3" gridrow="0" pageId="9" pageNumber="548">Genu</th>
<th id="76DD5DF8FFE8B8F35B9D2DF1ED381FF8" box="[737,856,191,216]" gridcol="4" gridrow="0" pageId="9" pageNumber="548">Tibia</th>
<th id="76DD5DF8FFE8B8F35AFC2DF1EA931FF8" box="[896,1267,191,216]" gridcol="5" gridrow="0" pageId="9" pageNumber="548">Tarsus</th>
</tr>
<tr id="350C3484FFE8B8F359DA2DA6EB2F1E25" box="[166,1359,232,261]" gridrow="1" pageId="9" pageNumber="548">
<th id="76DD5DF8FFE8B8F359DA2DA6EEAD1E25" box="[166,205,232,261]" gridcol="0" gridrow="1" pageId="9" pageNumber="548">I</th>
<td id="76DD5DF8FFE8B8F3598D2DA6EF3F1E25" box="[241,351,232,261]" gridcol="1" gridrow="1" pageId="9" pageNumber="548">-</td>
<td id="76DD5DF8FFE8B8F358F22DA6EC6D1E25" box="[398,525,232,261]" gridcol="2" gridrow="1" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE8470558F22DA6EFFB1E20" box="[398,411,232,256]" italics="true" pageId="9" pageNumber="548">d</emphasis>
, (
<emphasis id="B948EAD4FFE8470558CC2DA6EFD71E20" box="[432,439,232,256]" italics="true" pageId="9" pageNumber="548">l</emphasis>
),
<emphasis id="B948EAD4FFE8470558B02DA6EF841E20" box="[460,484,232,256]" italics="true" pageId="9" pageNumber="548">bv</emphasis>
,
<emphasis id="B948EAD4FFE84705588D2DA6EF9C1FDF" box="[497,508,232,255]" italics="true" pageId="9" pageNumber="548">v</emphasis>
</td>
<td id="76DD5DF8FFE8B8F35B3E2DA6ECD11E25" box="[578,689,232,261]" gridcol="3" gridrow="1" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE847055B3E2DA6ECE91FDF" box="[578,649,232,256]" italics="true" pageId="9" pageNumber="548">L”, d, σ</emphasis>
<emphasis id="B948EAD4FFE847055BF52DB9ECF01E25" box="[649,656,247,261]" italics="true" pageId="9" pageNumber="548">1</emphasis>
<emphasis id="B948EAD4FFE847055BED2DA6ECC91FDF" box="[657,681,232,255]" italics="true" pageId="9" pageNumber="548">, σ</emphasis>
<emphasis id="B948EAD4FFE847055BD62DB9ECD11E25" box="[682,689,247,261]" italics="true" pageId="9" pageNumber="548">2</emphasis>
</td>
<td id="76DD5DF8FFE8B8F35B9D2DA6ED381E25" box="[737,856,232,261]" gridcol="4" gridrow="1" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE847055B9D2DA6EC931FDF" box="[737,755,232,255]" italics="true" pageId="9" pageNumber="548">xt</emphasis>
<emphasis id="B948EAD4FFE847055B8F2DB9EC9A1E25" box="[755,762,247,261]" italics="true" pageId="9" pageNumber="548">2</emphasis>
<emphasis id="B948EAD4FFE847055B872DA6ED791FDF" box="[763,793,232,255]" italics="true" pageId="9" pageNumber="548">, xt</emphasis>
<emphasis id="B948EAD4FFE847055A662DB9ED411E25" box="[794,801,247,261]" italics="true" pageId="9" pageNumber="548">1</emphasis>
<emphasis id="B948EAD4FFE847055A5D2DA6ED381FDF" box="[801,856,232,255]" italics="true" pageId="9" pageNumber="548">, L, φ</emphasis>
</td>
<td id="76DD5DF8FFE8B8F35AFC2DA6EA931E25" box="[896,1267,232,261]" gridcol="5" gridrow="1" pageId="9" pageNumber="548">
(
<emphasis id="B948EAD4FFE847055AF52DA6EDF71E20" box="[905,919,232,256]" italics="true" pageId="9" pageNumber="548">ft</emphasis>
), (
<emphasis id="B948EAD4FFE847055AC82DA6EDA61FDF" box="[948,966,232,255]" italics="true" pageId="9" pageNumber="548">tc</emphasis>
),
<emphasis id="B948EAD4FFE847055AA02DA6ED911FDF" box="[988,1009,232,255]" italics="true" pageId="9" pageNumber="548">it,</emphasis>
(
<emphasis id="B948EAD4FFE847055A832DA6EA6C1FDF" box="[1023,1036,232,255]" italics="true" pageId="9" pageNumber="548">p</emphasis>
), (
<emphasis id="B948EAD4FFE847055D552DA6EA561FDF" box="[1065,1078,232,255]" italics="true" pageId="9" pageNumber="548">u</emphasis>
), (
<emphasis id="B948EAD4FFE847055D2F2DA6EA001FDF" box="[1107,1120,232,255]" italics="true" pageId="9" pageNumber="548">a</emphasis>
),
<emphasis id="B948EAD4FFE847055D092DA6EADD1FDF" box="[1141,1213,232,255]" italics="true" pageId="9" pageNumber="548">s, m, n,</emphasis>
(
<emphasis id="B948EAD4FFE847055DB12DA6EA851FDF" box="[1229,1253,232,255]" italics="true" pageId="9" pageNumber="548">pv</emphasis>
),
</td>
<td id="76DD5DF8FFE8B8F35D862DA6EB2F1E25" box="[1274,1359,232,261]" gridcol="6" gridrow="1" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE847055D862DA6EB421FDF" box="[1274,1314,232,255]" italics="true" pageId="9" pageNumber="548">ɛ, ω</emphasis>
<emphasis id="B948EAD4FFE847055C5E2DB9EB491E25" box="[1314,1321,247,261]" italics="true" pageId="9" pageNumber="548">1</emphasis>
<emphasis id="B948EAD4FFE847055C552DA6EB281FDF" box="[1321,1352,232,255]" italics="true" pageId="9" pageNumber="548">, ω</emphasis>
<emphasis id="B948EAD4FFE847055C342DB9EB2F1E25" box="[1352,1359,247,261]" italics="true" pageId="9" pageNumber="548">2</emphasis>
</td>
</tr>
<tr id="350C3484FFE8B8F359DA2C5EEB2F1E08" box="[166,1359,272,296]" gridrow="2" pageId="9" pageNumber="548" rowspan-6="1">
<th id="76DD5DF8FFE8B8F359DA2C5EEEAD1E08" box="[166,205,272,296]" gridcol="0" gridrow="2" pageId="9" pageNumber="548">II</th>
<td id="76DD5DF8FFE8B8F3598D2C5EEF3F1E08" box="[241,351,272,296]" gridcol="1" gridrow="2" pageId="9" pageNumber="548">-</td>
<td id="76DD5DF8FFE8B8F358F22C5EEC6D1E08" box="[398,525,272,296]" gridcol="2" gridrow="2" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE8470558F22C5EEFFB1E08" box="[398,411,272,296]" italics="true" pageId="9" pageNumber="548">d</emphasis>
, (
<emphasis id="B948EAD4FFE8470558CC2C5EEFD71E08" box="[432,439,272,296]" italics="true" pageId="9" pageNumber="548">l</emphasis>
),
<emphasis id="B948EAD4FFE8470558B02C5EEF841E08" box="[460,484,272,296]" italics="true" pageId="9" pageNumber="548">bv</emphasis>
,
<emphasis id="B948EAD4FFE84705588D2C5FEF9C1E08" box="[497,508,273,296]" italics="true" pageId="9" pageNumber="548">v</emphasis>
</td>
<td id="76DD5DF8FFE8B8F35B3E2C5EECD11E08" box="[578,689,272,296]" gridcol="3" gridrow="2" pageId="9" pageNumber="548">
(
<emphasis id="B948EAD4FFE847055B362C5EEC311E08" box="[586,593,272,296]" italics="true" pageId="9" pageNumber="548">l</emphasis>
),
<emphasis id="B948EAD4FFE847055B1A2C5FEC121E08" box="[614,626,273,296]" italics="true" pageId="9" pageNumber="548">σ</emphasis>
</td>
<td id="76DD5DF8FFE8B8F35B9D2C5EED381E08" box="[737,856,272,296]" gridcol="4" gridrow="2" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE847055B9D2C5FEC931E08" box="[737,755,273,296]" italics="true" pageId="9" pageNumber="548">xt</emphasis>
,
<emphasis id="B948EAD4FFE847055A7C2C5EED671E08" box="[768,775,272,296]" italics="true" pageId="9" pageNumber="548">l</emphasis>
,
<emphasis id="B948EAD4FFE847055A602C5FED4A1E08" box="[796,810,273,296]" italics="true" pageId="9" pageNumber="548">φ</emphasis>
</td>
<td id="76DD5DF8FFE8B8F35AFC2C5EEA931E08" box="[896,1267,272,296]" gridcol="5" gridrow="2" pageId="9" pageNumber="548">
(
<emphasis id="B948EAD4FFE847055AF52C5EEDF71E08" box="[905,919,272,296]" italics="true" pageId="9" pageNumber="548">ft</emphasis>
), (
<emphasis id="B948EAD4FFE847055AC82C5FEDA61E08" box="[948,966,273,296]" italics="true" pageId="9" pageNumber="548">tc</emphasis>
), (
<emphasis id="B948EAD4FFE847055A982C5FED911E08" box="[996,1009,273,296]" italics="true" pageId="9" pageNumber="548">p</emphasis>
), (
<emphasis id="B948EAD4FFE847055D732C5FEA7C1E08" box="[1039,1052,273,296]" italics="true" pageId="9" pageNumber="548">u</emphasis>
),
<emphasis id="B948EAD4FFE847055D4C2C5FEA5A1E08" box="[1072,1082,273,296]" italics="true" pageId="9" pageNumber="548">s</emphasis>
,
<emphasis id="B948EAD4FFE847055D3B2C5FEA351E08" box="[1095,1109,273,296]" italics="true" pageId="9" pageNumber="548">it</emphasis>
, (
<emphasis id="B948EAD4FFE847055D162C5FEAE21E08" box="[1130,1154,273,296]" italics="true" pageId="9" pageNumber="548">pv</emphasis>
),
<emphasis id="B948EAD4FFE847055DEB2C5FEAC91E08" box="[1175,1193,273,296]" italics="true" pageId="9" pageNumber="548">ω</emphasis>
</td>
</tr>
<tr id="350C3484FFE8B8F359DA2C76EB2F1E76" box="[166,1359,312,342]" gridrow="3" pageId="9" pageNumber="548" rowspan-6="1">
<th id="76DD5DF8FFE8B8F359DA2C76EEAD1E76" box="[166,205,312,342]" gridcol="0" gridrow="3" pageId="9" pageNumber="548">III</th>
<td id="76DD5DF8FFE8B8F3598D2C76EF3F1E76" box="[241,351,312,342]" gridcol="1" gridrow="3" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE84705598D2C76EE981E70" box="[241,248,312,336]" italics="true" pageId="9" pageNumber="548">l</emphasis>
,
<emphasis id="B948EAD4FFE8470558712C77EF781E70" box="[269,280,313,336]" italics="true" pageId="9" pageNumber="548">v</emphasis>
</td>
<td id="76DD5DF8FFE8B8F358F22C76EC6D1E76" box="[398,525,312,342]" gridcol="2" gridrow="3" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE8470558F22C76EFFB1E70" box="[398,411,312,336]" italics="true" pageId="9" pageNumber="548">d</emphasis>
,
<emphasis id="B948EAD4FFE8470558D42C76EFCF1E70" box="[424,431,312,336]" italics="true" pageId="9" pageNumber="548">l</emphasis>
1,
<emphasis id="B948EAD4FFE8470558B72C76EFB21E70" box="[459,466,312,336]" italics="true" pageId="9" pageNumber="548">l</emphasis>
2,
<emphasis id="B948EAD4FFE8470558932C77EC651E70" box="[495,517,313,336]" italics="true" pageId="9" pageNumber="548">ev</emphasis>
</td>
<td id="76DD5DF8FFE8B8F35B3E2C76ECD11E76" box="[578,689,312,342]" gridcol="3" gridrow="3" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE847055B3E2C76EC2F1E70" box="[578,591,312,336]" italics="true" pageId="9" pageNumber="548">d</emphasis>
,
<emphasis id="B948EAD4FFE847055B272C76EC021E70" box="[603,610,312,336]" italics="true" pageId="9" pageNumber="548">l</emphasis>
,
<emphasis id="B948EAD4FFE847055B0B2C77ECE31E70" box="[631,643,313,336]" italics="true" pageId="9" pageNumber="548">σ</emphasis>
</td>
<td id="76DD5DF8FFE8B8F35B9D2C76ED381E76" box="[737,856,312,342]" gridcol="4" gridrow="3" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE847055B9D2C76EC8E1E70" box="[737,750,312,336]" italics="true" pageId="9" pageNumber="548">d</emphasis>
,
<emphasis id="B948EAD4FFE847055B862C77ED681E70" box="[762,776,313,336]" italics="true" pageId="9" pageNumber="548">φ</emphasis>
</td>
<td id="76DD5DF8FFE8B8F35AFC2C76EA931E76" box="[896,1267,312,342]" gridcol="5" gridrow="3" pageId="9" pageNumber="548">
(
<emphasis id="B948EAD4FFE847055AF52C76EDF71E70" box="[905,919,312,336]" italics="true" pageId="9" pageNumber="548">ft</emphasis>
), (
<emphasis id="B948EAD4FFE847055AC82C77EDA61E70" box="[948,966,313,336]" italics="true" pageId="9" pageNumber="548">tc</emphasis>
), (
<emphasis id="B948EAD4FFE847055A982C77ED911E70" box="[996,1009,313,336]" italics="true" pageId="9" pageNumber="548">p</emphasis>
), (
<emphasis id="B948EAD4FFE847055D732C77EA7C1E70" box="[1039,1052,313,336]" italics="true" pageId="9" pageNumber="548">u</emphasis>
),
<emphasis id="B948EAD4FFE847055D4C2C77EA5A1E70" box="[1072,1082,313,336]" italics="true" pageId="9" pageNumber="548">s</emphasis>
, (
<emphasis id="B948EAD4FFE847055D332C77EA071E70" box="[1103,1127,313,336]" italics="true" pageId="9" pageNumber="548">pv</emphasis>
)
</td>
</tr>
<tr id="350C3484FFE8B8F359DA2C2EEB2F1E59" box="[166,1359,352,377]" gridrow="4" pageId="9" pageNumber="548" rowspan-6="1">
<th id="76DD5DF8FFE8B8F359DA2C2EEEAD1E59" box="[166,205,352,377]" gridcol="0" gridrow="4" pageId="9" pageNumber="548">IV</th>
<td id="76DD5DF8FFE8B8F3598D2C2EEF3F1E59" box="[241,351,352,377]" gridcol="1" gridrow="4" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE84705598D2C2EEE981E58" box="[241,248,352,376]" italics="true" pageId="9" pageNumber="548">l</emphasis>
,
<emphasis id="B948EAD4FFE8470558712C2FEF781E58" box="[269,280,353,376]" italics="true" pageId="9" pageNumber="548">v</emphasis>
</td>
<td id="76DD5DF8FFE8B8F358F22C2EEC6D1E59" box="[398,525,352,377]" gridcol="2" gridrow="4" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE8470558F22C2EEFFB1E58" box="[398,411,352,376]" italics="true" pageId="9" pageNumber="548">d</emphasis>
,
<emphasis id="B948EAD4FFE8470558D42C2EEFCF1E58" box="[424,431,352,376]" italics="true" pageId="9" pageNumber="548">l</emphasis>
,
<emphasis id="B948EAD4FFE8470558B82C2FEFBA1E58" box="[452,474,353,376]" italics="true" pageId="9" pageNumber="548">ev</emphasis>
</td>
<td id="76DD5DF8FFE8B8F35B3E2C2EECD11E59" box="[578,689,352,377]" gridcol="3" gridrow="4" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE847055B3E2C2EEC2F1E58" box="[578,591,352,376]" italics="true" pageId="9" pageNumber="548">d</emphasis>
,
<emphasis id="B948EAD4FFE847055B272C2EEC021E58" box="[603,610,352,376]" italics="true" pageId="9" pageNumber="548">l</emphasis>
,
<emphasis id="B948EAD4FFE847055B0B2C2FECE31E58" box="[631,643,353,376]" italics="true" pageId="9" pageNumber="548">σ</emphasis>
</td>
<td id="76DD5DF8FFE8B8F35B9D2C2EED381E59" box="[737,856,352,377]" gridcol="4" gridrow="4" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE847055B9D2C2EED611E58" box="[737,769,352,376]" italics="true" pageId="9" pageNumber="548">d, l</emphasis>
,
<emphasis id="B948EAD4FFE847055A6B2C2FED421E58" box="[791,802,353,376]" italics="true" pageId="9" pageNumber="548">v</emphasis>
</td>
<td id="76DD5DF8FFE8B8F35AFC2C2EEA931E59" box="[896,1267,352,377]" gridcol="5" gridrow="4" pageId="9" pageNumber="548">
(
<emphasis id="B948EAD4FFE847055AF52C2EEDF71E58" box="[905,919,352,376]" italics="true" pageId="9" pageNumber="548">ft</emphasis>
), (
<emphasis id="B948EAD4FFE847055AC82C2FEDA61E58" box="[948,966,353,376]" italics="true" pageId="9" pageNumber="548">tc</emphasis>
), (
<emphasis id="B948EAD4FFE847055A982C2FED911E58" box="[996,1009,353,376]" italics="true" pageId="9" pageNumber="548">p</emphasis>
), (
<emphasis id="B948EAD4FFE847055D732C2FEA7C1E58" box="[1039,1052,353,376]" italics="true" pageId="9" pageNumber="548">u</emphasis>
),
<emphasis id="B948EAD4FFE847055D4C2C2FEA5A1E58" box="[1072,1082,353,376]" italics="true" pageId="9" pageNumber="548">s</emphasis>
, (
<emphasis id="B948EAD4FFE847055D332C2FEA071E58" box="[1103,1127,353,376]" italics="true" pageId="9" pageNumber="548">pv</emphasis>
)
</td>
</tr>
</table>
</paragraph>
<subSubSection id="C326654DFFE8470559BB2CE6EA3F1D56" pageId="9" pageNumber="548" type="materials_examined">
<paragraph id="8B8336C6FFE8470559BB2CE6EFBC1D2A" blockId="9.[151,1437,424,1458]" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE8470559BB2CE6EFDF1EE2" bold="true" box="[199,447,424,450]" pageId="9" pageNumber="548">Materials examined.</emphasis>
<materialsCitation id="3B543C9BFFE8470558B52CE6EFBC1D2A" ID-GBIF-Occurrence="3044016301" collectingDate="2019-04-13" collectorName="Coll. Arun, A." country="India" latitude="11.13513" location="University Campus" longLatPrecision="1" longitude="75.88847" municipality="Malappuram" pageId="9" pageNumber="548" specimenCount="10" stateProvince="Kerala" typeStatus="holotype">
<typeStatus id="54878864FFE8470558B52CE6EC511EE2" box="[457,561,424,450]" pageId="9" pageNumber="548" type="holotype">Holotype</typeStatus>
and
<specimenCount id="9D3AFD4FFFE847055B122CE6ED641EE2" box="[622,772,424,450]" pageId="9" pageNumber="548" type="generic" typeStatus="paratypes">10 paratypes</specimenCount>
: Collected from densely accumulated litter samples of
<emphasis id="B948EAD4FFE847055CF82CE6EF201EC6" italics="true" pageId="9" pageNumber="548">A. auriculiformis¸</emphasis>
Calicut
<location id="8EE3601DFFE8470558E32C82EC181EC6" LSID="urn:lsid:plazi:treatment:039587D0FFE2470559EB2C4CEB461A92:8EE3601DFFE8470558E32C82EC181EC6" box="[415,632,460,486]" country="India" latitude="11.13513" longLatPrecision="1" longitude="75.88847" municipality="Malappuram" name="University Campus" pageId="9" pageNumber="548" stateProvince="Kerala">University Campus</location>
(
<geoCoordinate id="EE085001FFE847055BF42C82ED5B1EC7" box="[648,827,460,487]" degrees="11" direction="north" minutes="8" orientation="latitude" pageId="9" pageNumber="548" precision="1" seconds="6.468" value="11.13513">11° 8 6.468 N</geoCoordinate>
<geoCoordinate id="EE085001FFE847055A472C82EA761EC6" box="[827,1046,460,487]" degrees="75" direction="east" minutes="53" orientation="longitude" pageId="9" pageNumber="548" precision="1" seconds="18.5028" value="75.88847">75° 53 18.5028 E</geoCoordinate>
),
<collectingMunicipality id="6BE7ACBCFFE847055D562C82EAD91EC6" box="[1066,1209,460,486]" pageId="9" pageNumber="548">Malappuram</collectingMunicipality>
(Dt.),
<collectingRegion id="49F8F824FFE847055C7F2C82EB301EC6" box="[1283,1360,460,486]" country="India" name="Kerala" pageId="9" pageNumber="548">Kerala</collectingRegion>
,
<collectingCountry id="F32B7656FFE847055C202C82EBF81EC6" box="[1372,1432,460,486]" name="India" pageId="9" pageNumber="548">India</collectingCountry>
,
<collectorName id="26C95310FFE8470559EB2CBEEF551D2A" box="[151,309,496,522]" pageId="9" pageNumber="548">Coll. Arun, A.</collectorName>
on
<date id="FF821006FFE8470558232CBEEFB81D2A" box="[351,472,496,522]" pageId="9" pageNumber="548" value="2019-04-13">
<collectingDate id="EFC6E9EEFFE8470558232CBEEFB81D2A" box="[351,472,496,522]" pageId="9" pageNumber="548" value="2019-04-13">13.iv.2019</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8B8336C6FFE8470559BB2F5AEA3F1D56" blockId="9.[151,1437,424,1458]" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE8470559BB2F5AEFEE1D0E" bold="true" box="[199,398,532,558]" pageId="9" pageNumber="548">Type deposition.</emphasis>
<materialsCitation id="3B543C9BFFE8470558EA2F5AEB541D0E" ID-GBIF-Occurrence="3044016309" box="[406,1332,532,558]" country="India" location="Zoological" pageId="9" pageNumber="548" specimenCount="2" typeStatus="holotype">
The
<typeStatus id="54878864FFE8470558B02F5AEC4E1D0E" box="[460,558,532,558]" pageId="9" pageNumber="548" type="holotype">holotype</typeStatus>
and five
<typeStatus id="54878864FFE847055BE22F5AED691D0E" box="[670,777,532,558]" pageId="9" pageNumber="548" type="paratype">paratypes</typeStatus>
are deposited in the
<location id="8EE3601DFFE847055A832F5AEA181D0E" LSID="urn:lsid:plazi:treatment:039587D0FFE2470559EB2C4CEB461A92:8EE3601DFFE847055A832F5AEA181D0E" box="[1023,1144,532,558]" country="India" name="Zoological" pageId="9" pageNumber="548">Zoological</location>
survey of
<collectingCountry id="F32B7656FFE847055D842F5AEB541D0E" box="[1272,1332,532,558]" name="India" pageId="9" pageNumber="548">India</collectingCountry>
</materialsCitation>
,
<materialsCitation id="3B543C9BFFE847055C3D2F5AEC791D72" ID-GBIF-Occurrence="3044016312" country="India" location="Western Ghat Regional Centre" pageId="9" pageNumber="548" specimenCount="1" typeStatus="holotype">
<location id="8EE3601DFFE847055C3D2F5AEFF01D72" LSID="urn:lsid:plazi:treatment:039587D0FFE2470559EB2C4CEB461A92:8EE3601DFFE847055C3D2F5AEFF01D72" country="India" name="Western Ghat Regional Centre" pageId="9" pageNumber="548">Western Ghat Regional Centre</location>
,
<collectingCountry id="F32B7656FFE8470558E72F76EC791D72" box="[411,537,568,594]" name="India" pageId="9" pageNumber="548">Kozhikode</collectingCountry>
</materialsCitation>
,
<materialsCitation id="3B543C9BFFE847055B592F76ECDB1D72" ID-GBIF-Occurrence="3044016308" box="[549,699,568,594]" country="India" location="Kerala" pageId="9" pageNumber="548" specimenCount="1" stateProvince="Kerala" typeStatus="holotype">
<collectingRegion id="49F8F824FFE847055B592F76EC121D72" box="[549,626,568,594]" country="India" name="Kerala" pageId="9" pageNumber="548">Kerala</collectingRegion>
,
<collectingCountry id="F32B7656FFE847055B022F76ECDB1D72" box="[638,699,568,594]" name="India" pageId="9" pageNumber="548">India</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B543C9BFFE847055BBB2F76ED7E1D56" ID-GBIF-Occurrence="3044016306" country="India" county="University" location="Acarology Collections" municipality="Department of Zoology" pageId="9" pageNumber="548" specimenCount="1" typeStatus="paratype">
and four
<typeStatus id="54878864FFE847055A522F76EDF91D72" box="[814,921,568,594]" pageId="9" pageNumber="548" type="paratype">paratypes</typeStatus>
are in the
<location id="8EE3601DFFE847055D6D2F76EB701D72" LSID="urn:lsid:plazi:treatment:039587D0FFE2470559EB2C4CEB461A92:8EE3601DFFE847055D6D2F76EB701D72" box="[1041,1296,568,594]" country="India" county="University" municipality="Department of Zoology" name="Acarology Collections" pageId="9" pageNumber="548">Acarology Collections</location>
,
<location id="8EE3601DFFE847055C612F76EF6E1D56" LSID="urn:lsid:plazi:treatment:039587D0FFE2470559EB2C4CEB461A92:8EE3601DFFE847055C612F76EF6E1D56" country="India" county="University" municipality="Department of Zoology" name="Division of Acarology" pageId="9" pageNumber="548">Division of Acarology</location>
,
<collectingMunicipality id="6BE7ACBCFFE8470558662F12EC431D56" box="[282,547,604,630]" pageId="9" pageNumber="548">Department of Zoology</collectingMunicipality>
,
<collectingCounty id="62E24E4AFFE847055B522F12ECC41D56" box="[558,676,604,630]" pageId="9" pageNumber="548">University</collectingCounty>
of
<collectingCountry id="F32B7656FFE847055BB62F12ED7E1D56" box="[714,798,604,630]" name="India" pageId="9" pageNumber="548">Calicut</collectingCountry>
</materialsCitation>
,
<materialsCitation id="3B543C9BFFE847055A552F12EA3B1D56" ID-GBIF-Occurrence="3044016304" box="[809,1115,604,630]" country="India" location="Kerala" pageId="9" pageNumber="548" specimenCount="1" stateProvince="Kerala" typeStatus="paratype">
Malappuram,
<collectingRegion id="49F8F824FFE847055ABA2F12EA731D56" box="[966,1043,604,630]" country="India" name="Kerala" pageId="9" pageNumber="548">Kerala</collectingRegion>
,
<collectingCountry id="F32B7656FFE847055D622F12EA3B1D56" box="[1054,1115,604,630]" name="India" pageId="9" pageNumber="548">India</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C326654DFFE8470559BB2FCEEFBF1D9E" pageId="9" pageNumber="548" type="etymology">
<paragraph id="8B8336C6FFE8470559BB2FCEEFBF1D9E" blockId="9.[151,1437,424,1458]" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE8470559BB2FCEEF2F1DBA" bold="true" box="[199,335,640,666]" pageId="9" pageNumber="548">Etymology.</emphasis>
The species name “
<taxonomicName id="4C3C4D45FFE847055B442FCFECCD1DBA" authorityName="Arun &amp; Ramani" authorityYear="2020" box="[568,685,641,666]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="acaciensis">
<emphasis id="B948EAD4FFE847055B442FCFECCD1DBA" box="[568,685,641,666]" italics="true" pageId="9" pageNumber="548">acaciensis</emphasis>
</taxonomicName>
” is derived from the litter of
<emphasis id="B948EAD4FFE847055D7E2FCEEAAA1DBA" box="[1026,1226,640,666]" italics="true" pageId="9" pageNumber="548">
<taxonomicName id="4C3C4D45FFE847055D7E2FCEEAA61DBA" box="[1026,1222,640,666]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="9" pageNumber="548" phylum="Tracheophyta" rank="species" species="auriculiformis">A. auriculiformis</taxonomicName>
,
</emphasis>
which formed the natural habitat of the species.
</paragraph>
</subSubSection>
<subSubSection id="C326654DFFE8470559BB2F86EFFA1C53" pageId="9" pageNumber="548" type="biology_ecology">
<paragraph id="8B8336C6FFE8470559BB2F86EFFA1C53" blockId="9.[151,1437,424,1458]" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE8470559BB2F86EF4D1DC2" bold="true" box="[199,301,712,738]" pageId="9" pageNumber="548">Ecology.</emphasis>
All the life stages of
<taxonomicName id="4C3C4D45FFE847055B4E2F86ECA31DC2" authorityName="Kunst" authorityYear="1959" box="[562,707,712,738]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="548" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAD4FFE847055B4E2F86ECA31DC2" box="[562,707,712,738]" italics="true" pageId="9" pageNumber="548">Papillacarus</emphasis>
</taxonomicName>
<taxonomicName id="4C3C4D45FFE847055BB12F86ED171DC2" authority="Arun &amp; Ramani, 2020" authorityName="Arun &amp; Ramani" authorityYear="2020" box="[717,887,712,738]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="acaciensis" status="sp. nov." subGenus="Vepracarus">
(
<emphasis id="B948EAD4FFE847055BAA2F87EC871DC2" box="[726,743,713,738]" italics="true" pageId="9" pageNumber="548">V</emphasis>
.)
<emphasis id="B948EAD4FFE847055A7E2F87ED171DC2" box="[770,887,713,738]" italics="true" pageId="9" pageNumber="548">acaciensis</emphasis>
</taxonomicName>
<emphasis id="B948EAD4FFE847055AFD2F86EDBE1DC2" bold="true" box="[897,990,712,738]" pageId="9" pageNumber="548">
<taxonomicNameLabel id="A27B57AFFFE847055AFD2F86EDBE1DC2" box="[897,990,712,738]" pageId="9" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
such as the larvae, the three nymphal stages and adults were recovered from the decomposing litter of
<emphasis id="B948EAD4FFE847055A0C2FA2EA581C26" box="[880,1080,748,774]" italics="true" pageId="9" pageNumber="548">
<taxonomicName id="4C3C4D45FFE847055A0C2FA2EA541C26" box="[880,1076,748,774]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="9" pageNumber="548" phylum="Tracheophyta" rank="species" species="auriculiformis">A. auriculiformis</taxonomicName>
.
</emphasis>
<taxonomicName id="4C3C4D45FFE847055D422FA2EAEC1C26" box="[1086,1164,748,774]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="9" pageNumber="548" phylum="Tracheophyta" rank="genus">Acacia</taxonomicName>
litter was recognized as the naturally preferred diet of the species, supporting its reproduction and subsequent completion of life history as evidenced through the detection of varied number of food boli in the gut of field collected life stages of the species (personal observation).
</paragraph>
</subSubSection>
<subSubSection id="C326654DFFE8470559BB2E32EB461A92" pageId="9" pageNumber="548" type="discussion">
<paragraph id="8B8336C6FFE8470559BB2E32EF531BDF" blockId="9.[151,1437,424,1458]" pageId="9" pageNumber="548">
<emphasis id="B948EAD4FFE8470559BB2E32EF541CB6" bold="true" box="[199,308,892,918]" pageId="9" pageNumber="548">Remarks</emphasis>
. The specimens of
<taxonomicName id="4C3C4D45FFE847055B762E32ECFB1CB6" authorityName="Kunst" authorityYear="1959" box="[522,667,892,918]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="548" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAD4FFE847055B762E32ECFB1CB6" box="[522,667,892,918]" italics="true" pageId="9" pageNumber="548">Papillacarus</emphasis>
</taxonomicName>
<taxonomicName id="4C3C4D45FFE847055BDC2E32ED251CB6" authority="Arun &amp; Ramani, 2020" authorityName="Arun &amp; Ramani" authorityYear="2020" box="[672,837,892,918]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="acaciensis" status="sp. nov." subGenus="Vepracarus">
(
<emphasis id="B948EAD4FFE847055BD52E33ECDA1CB6" box="[681,698,893,918]" italics="true" pageId="9" pageNumber="548">V</emphasis>
.)
<emphasis id="B948EAD4FFE847055BAC2E33ED251CB6" box="[720,837,893,918]" italics="true" pageId="9" pageNumber="548">acaciensis</emphasis>
</taxonomicName>
<emphasis id="B948EAD4FFE847055A372E32EDC31CB6" bold="true" box="[843,931,892,918]" pageId="9" pageNumber="548">
<taxonomicNameLabel id="A27B57AFFFE847055A372E32EDC31CB6" box="[843,931,892,918]" pageId="9" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
when compared with the other known congeners, were found to show most resemblance to the North Indian species,
<taxonomicName id="4C3C4D45FFE847055AC52EEFEB1C1C9A" authority="(Sarkar &amp; Subias, 1984)" baseAuthorityName="Sarkar &amp; Subias" baseAuthorityYear="1984" box="[953,1404,928,954]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="cornutus" subGenus="Vepracarus">
<emphasis id="B948EAD4FFE847055AC52EEFEDAE1C9A" box="[953,974,929,954]" italics="true" pageId="9" pageNumber="548">P.</emphasis>
(
<emphasis id="B948EAD4FFE847055AA12EEFED8E1C9A" box="[989,1006,929,954]" italics="true" pageId="9" pageNumber="548">V</emphasis>
.)
<emphasis id="B948EAD4FFE847055D782EEFEA061C9A" box="[1028,1126,929,954]" italics="true" pageId="9" pageNumber="548">cornutus</emphasis>
(
<bibRefCitation id="EFAD4B37FFE847055D082EEEEB141C9A" author="Sarkar, S. &amp; Subias, L. S." box="[1140,1396,928,954]" pageId="9" pageNumber="548" pagination="25 - 30" refId="ref13547" refString="Sarkar, S. &amp; Subias, L. S. (1984) New lohmanniids (Acarida: Oribatida) from India. Oriental Insects, 18, 25 - 30. https: // doi. org / 10.1080 / 00305316.1984.10432192" type="journal article" year="1984">Sarkar &amp; Subías, 1984</bibRefCitation>
)
</taxonomicName>
, in the nature of prodorsal and notogastral setae, presence of a tooth on the lateral prodorsal margin, above the insertion of seta
<emphasis id="B948EAD4FFE8470559992EA7EF6C1B22" box="[229,268,1001,1026]" italics="true" pageId="9" pageNumber="548">exa</emphasis>
and the possession of tubercles on prodorsum and notogaster. However, the new species keeps its identity separate from the above species in the following respects (1) incompletely separated nature of anal and adanal plates (in
<taxonomicName id="4C3C4D45FFE8470559C3297FEF141B6A" baseAuthorityName="Sarkar &amp; Subias" baseAuthorityYear="1984" box="[191,372,1072,1098]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="cornutus" subGenus="Vepracarus">
<emphasis id="B948EAD4FFE8470559C3297FEEB41B6A" box="[191,212,1073,1098]" italics="true" pageId="9" pageNumber="548">P.</emphasis>
(
<emphasis id="B948EAD4FFE847055999297FEE961B6A" box="[229,246,1073,1098]" italics="true" pageId="9" pageNumber="548">V</emphasis>
.)
<emphasis id="B948EAD4FFE847055873297FEF141B6A" box="[271,372,1073,1098]" italics="true" pageId="9" pageNumber="548">cornutus</emphasis>
</taxonomicName>
<emphasis id="B948EAD4FFE847055808297FEF181B6A" box="[372,376,1073,1098]" italics="true" pageId="9" pageNumber="548">,</emphasis>
anal and adanal plates are completely separated); (2) elongated nature of posteriormost pairs of the notogastral setae (short in
<taxonomicName id="4C3C4D45FFE847055B71291BECDF1B4E" baseAuthorityName="Sarkar &amp; Subias" baseAuthorityYear="1984" box="[525,703,1108,1134]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="cornutus" subGenus="Vepracarus">
<emphasis id="B948EAD4FFE847055B71291BEC421B4E" box="[525,546,1109,1134]" italics="true" pageId="9" pageNumber="548">P.</emphasis>
(
<emphasis id="B948EAD4FFE847055B4F291BEC241B4E" box="[563,580,1109,1134]" italics="true" pageId="9" pageNumber="548">V</emphasis>
.)
<emphasis id="B948EAD4FFE847055B21291BECDF1B4E" box="[605,703,1109,1134]" italics="true" pageId="9" pageNumber="548">cornutus</emphasis>
</taxonomicName>
); (3) presence of feebly developed polygonal sculptures beneath prodorsal band (absent in
<taxonomicName id="4C3C4D45FFE8470558BD2937EC141BB2" baseAuthorityName="Sarkar &amp; Subias" baseAuthorityYear="1984" box="[449,628,1144,1170]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="cornutus" subGenus="Vepracarus">
<emphasis id="B948EAD4FFE8470558BD2937EFB61BB2" box="[449,470,1145,1170]" italics="true" pageId="9" pageNumber="548">P.</emphasis>
(
<emphasis id="B948EAD4FFE8470558942937EF991BB2" box="[488,505,1145,1170]" italics="true" pageId="9" pageNumber="548">V</emphasis>
.)
<emphasis id="B948EAD4FFE847055B6E2937EC141BB2" box="[530,628,1145,1170]" italics="true" pageId="9" pageNumber="548">cornutus</emphasis>
</taxonomicName>
); (4) differences in the length of barbs on
<emphasis id="B948EAD4FFE847055D232937EA151BB2" box="[1119,1141,1145,1170]" italics="true" pageId="9" pageNumber="548">ss</emphasis>
(distal and basal barbs of
<emphasis id="B948EAD4FFE8470559EB29D3EECD1B96" box="[151,173,1181,1206]" italics="true" pageId="9" pageNumber="548">ss</emphasis>
shorter than the medial barbs in the new species while in
<taxonomicName id="4C3C4D45FFE847055A4329D3ED931B96" baseAuthorityName="Sarkar &amp; Subias" baseAuthorityYear="1984" box="[831,1011,1180,1206]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="cornutus" subGenus="Vepracarus">
<emphasis id="B948EAD4FFE847055A4329D3ED341B96" box="[831,852,1181,1206]" italics="true" pageId="9" pageNumber="548">P.</emphasis>
(
<emphasis id="B948EAD4FFE847055A1929D3ED161B96" box="[869,886,1181,1206]" italics="true" pageId="9" pageNumber="548">V</emphasis>
.)
<emphasis id="B948EAD4FFE847055AF229D3ED931B96" box="[910,1011,1181,1206]" italics="true" pageId="9" pageNumber="548">cornutus</emphasis>
</taxonomicName>
<emphasis id="B948EAD4FFE847055A8E29D3ED961B96" box="[1010,1014,1181,1206]" italics="true" pageId="9" pageNumber="548">,</emphasis>
distal barbs are longer than the basal ones); (5) genital setae
<emphasis id="B948EAD4FFE8470558DC298FEFCE1BFA" box="[416,430,1217,1242]" italics="true" pageId="9" pageNumber="548">g</emphasis>
<subScript id="17B83483FFE8470558D2299CEFD61BC0" attach="left" box="[430,438,1234,1248]" fontSize="6" pageId="9" pageNumber="548">6</subScript>
,
<emphasis id="B948EAD4FFE8470558B9298FEFB31BFA" box="[453,467,1217,1242]" italics="true" pageId="9" pageNumber="548">g</emphasis>
<subScript id="17B83483FFE8470558AF299DEFBB1BC1" attach="left" box="[467,475,1235,1249]" fontSize="6" pageId="9" pageNumber="548">7</subScript>
,
<emphasis id="B948EAD4FFE847055896298FEF981BFA" box="[490,504,1217,1242]" italics="true" pageId="9" pageNumber="548">g</emphasis>
<subScript id="17B83483FFE847055884299CEC601BC0" attach="left" box="[504,512,1234,1248]" fontSize="6" pageId="9" pageNumber="548">8</subScript>
,
<emphasis id="B948EAD4FFE847055B73298FEC7D1BFA" box="[527,541,1217,1242]" italics="true" pageId="9" pageNumber="548">g</emphasis>
<subScript id="17B83483FFE847055B61299CEC4D1BC0" attach="left" box="[541,557,1234,1248]" fontSize="6" pageId="9" pageNumber="548">10</subScript>
setiform and smooth, and other setae branched (in
<taxonomicName id="4C3C4D45FFE847055D0F298FEB441BFA" baseAuthorityName="Sarkar &amp; Subias" baseAuthorityYear="1984" box="[1139,1316,1216,1242]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="cornutus" subGenus="Vepracarus">
<emphasis id="B948EAD4FFE847055D0F298FEAE91BFA" box="[1139,1161,1217,1242]" italics="true" pageId="9" pageNumber="548">P.</emphasis>
(
<emphasis id="B948EAD4FFE847055DE5298FEACA1BFA" box="[1177,1194,1217,1242]" italics="true" pageId="9" pageNumber="548">V</emphasis>
.)
<emphasis id="B948EAD4FFE847055DBE298FEB441BFA" box="[1218,1316,1217,1242]" italics="true" pageId="9" pageNumber="548">cornutus</emphasis>
</taxonomicName>
all genital setae barbed).
</paragraph>
<paragraph id="8B8336C6FFE8470559BB2846EB461A92" blockId="9.[151,1437,424,1458]" pageId="9" pageNumber="548">
The new species
<emphasis id="B948EAD4FFE8470558EB2847EFCC1A02" box="[407,428,1289,1314]" italics="true" pageId="9" pageNumber="548">P.</emphasis>
<taxonomicName id="4C3C4D45FFE8470558CA2846EC041A02" authorityName="Arun &amp; Ramani" authorityYear="2020" box="[438,612,1288,1314]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="acaciensis" subGenus="Vepracarus">
(
<emphasis id="B948EAD4FFE8470558BC2847EFB11A02" box="[448,465,1289,1314]" italics="true" pageId="9" pageNumber="548">V</emphasis>
.)
<emphasis id="B948EAD4FFE8470558912847EC041A02" box="[493,612,1289,1314]" italics="true" pageId="9" pageNumber="548">acaciensis</emphasis>
</taxonomicName>
shows resemblance to another described species,
<taxonomicName id="4C3C4D45FFE847055DC72847EF3F1A67" authority="(Mahunka, 1985)" baseAuthorityName="Mahunka" baseAuthorityYear="1985" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="549" phylum="Arthropoda" rank="species" species="incompletus" subGenus="Vepracarus">
<emphasis id="B948EAD4FFE847055DC72847EAB11A02" box="[1211,1233,1289,1314]" italics="true" pageId="9" pageNumber="548">P.</emphasis>
(
<emphasis id="B948EAD4FFE847055D992847EA961A02" box="[1253,1270,1289,1314]" italics="true" pageId="9" pageNumber="548">V</emphasis>
.)
<emphasis id="B948EAD4FFE847055C6E2846EBFC1A02" box="[1298,1436,1288,1314]" italics="true" pageId="9" pageNumber="548">incompletus</emphasis>
(
<bibRefCitation id="EFAD4B37FFE8470559E32862EF371A67" author="Mahunka, S." box="[159,343,1324,1351]" pageId="9" pageNumber="548" pagination="119 - 178" refId="ref13121" refString="Mahunka, S. (1985) Mites (Acari) from St. Lucia (Antilles). 2. Oribatida. Acta Zoologica Hungarica, 31 (1 - 3), 119 - 178." type="journal article" year="1985">Mahunka, 1985</bibRefCitation>
)
</taxonomicName>
described from
<collectingCountry id="F32B7656FFE847055B672862EC351A66" box="[539,597,1324,1350]" name="India" pageId="9" pageNumber="548">India</collectingCountry>
in the branched nature of prodorsal and notogastral setae and incomplete separation of ano-adanal plates. However, the new species is distinct from the above species in having: (1) an ornamentation of dorsal surface with tubercles (papilli in
<taxonomicName id="4C3C4D45FFE847055A7E283BEDB61AAE" baseAuthorityName="Mahunka" baseAuthorityYear="1985" box="[770,982,1396,1422]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="549" phylum="Arthropoda" rank="species" species="incompletus" subGenus="Vepracarus">
<emphasis id="B948EAD4FFE847055A7E283BED771AAE" box="[770,791,1397,1422]" italics="true" pageId="9" pageNumber="548">P.</emphasis>
(
<emphasis id="B948EAD4FFE847055A5B283BED581AAE" box="[807,824,1397,1422]" italics="true" pageId="9" pageNumber="548">V</emphasis>
.)
<emphasis id="B948EAD4FFE847055A33283AEDB61AAE" box="[847,982,1396,1422]" italics="true" pageId="9" pageNumber="548">incompletus</emphasis>
</taxonomicName>
); (2) seta
<emphasis id="B948EAD4FFE847055D34283BEA0F1AAE" box="[1096,1135,1397,1422]" italics="true" pageId="9" pageNumber="548">exp</emphasis>
shorter than
<emphasis id="B948EAD4FFE847055C7D283BEB771AAE" box="[1281,1303,1397,1422]" italics="true" pageId="9" pageNumber="548">ss</emphasis>
(
<emphasis id="B948EAD4FFE847055C5B283BEB2E1AAE" box="[1319,1358,1397,1422]" italics="true" pageId="9" pageNumber="548">exp</emphasis>
longer than
<emphasis id="B948EAD4FFE8470559B228D7EE841A92" box="[206,228,1433,1458]" italics="true" pageId="9" pageNumber="548">ss</emphasis>
in
<taxonomicName id="4C3C4D45FFE84705587428D7EFBC1A92" baseAuthorityName="Mahunka" baseAuthorityYear="1985" box="[264,476,1432,1458]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="549" phylum="Arthropoda" rank="species" species="incompletus" subGenus="Vepracarus">
<emphasis id="B948EAD4FFE84705587428D7EF7D1A92" box="[264,285,1433,1458]" italics="true" pageId="9" pageNumber="548">P.</emphasis>
(
<emphasis id="B948EAD4FFE84705585128D7EF5E1A92" box="[301,318,1433,1458]" italics="true" pageId="9" pageNumber="548">V</emphasis>
.)
<emphasis id="B948EAD4FFE84705582928D6EFBC1A92" box="[341,476,1432,1458]" italics="true" pageId="9" pageNumber="548">incompletus</emphasis>
</taxonomicName>
) and (3) epimeral setal formula of 7-4-3-3 (9-4-
<quantity id="4CC49B23FFE847055A8428D6EA5A1A92" box="[1016,1082,1432,1458]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="9" pageNumber="548" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3-4 in</quantity>
<taxonomicName id="4C3C4D45FFE847055D3D28D7EB751A92" baseAuthorityName="Mahunka" baseAuthorityYear="1985" box="[1089,1301,1432,1458]" class="Arachnida" family="Lohmanniidae" genus="Papillacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="549" phylum="Arthropoda" rank="species" species="incompletus" subGenus="Vepracarus">
<emphasis id="B948EAD4FFE847055D3D28D7EA371A92" box="[1089,1111,1433,1458]" italics="true" pageId="9" pageNumber="548">P.</emphasis>
(
<emphasis id="B948EAD4FFE847055D1A28D7EA171A92" box="[1126,1143,1433,1458]" italics="true" pageId="9" pageNumber="548">V</emphasis>
.)
<emphasis id="B948EAD4FFE847055DF228D6EB751A92" box="[1166,1301,1432,1458]" italics="true" pageId="9" pageNumber="548">incompletus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>