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<document id="CD1B52439F018FB8F59E9EB91415B7C7" ID-DOI="10.5281/zenodo.196425" ID-GBIF-Dataset="e18a85f3-43a2-4c80-8a1f-698148f15a9b" ID-ISSN="1175-5326" ID-Zenodo-Dep="196425" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460243284416" checkinUser="plazi" docAuthor="Mironov, Sergey V. &amp; Dabert, Jacek" docDate="2010" docId="039587A15333541EFF14DD11E2C6FDCF" docLanguage="en" docName="zt02526p036.pdf" docOrigin="Zootaxa 2526" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Protolichus gratus Favette et Trouessart 1904" docType="treatment" docVersion="10" lastPageNumber="33" masterDocId="FFACFFD9532C543EFF83DF16E753FFBD" masterDocTitle="Systematic revision of the feather mite genus Protolichus Trouessart, 1884 (Astigmata, Pterolichidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="32" updateTime="1698241288831" updateUser="plazi">
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<mods:title id="9C74276ED748F525516A3812475FF61E">Systematic revision of the feather mite genus Protolichus Trouessart, 1884 (Astigmata, Pterolichidae)</mods:title>
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<mods:namePart id="C2B3557D59466F996D6FBCC0D4046638">Mironov, Sergey V.</mods:namePart>
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<mods:namePart id="91580DF2C070F8C4EB505F62C604E2F7">Dabert, Jacek</mods:namePart>
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<mods:date id="47986F871001B9048A9666838CF9EC23">2010</mods:date>
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<treatment id="039587A15333541EFF14DD11E2C6FDCF" ID-DOI="http://doi.org/10.5281/zenodo.5686107" ID-GBIF-Taxon="119390082" ID-Zenodo-Dep="5686107" LSID="urn:lsid:plazi:treatment:039587A15333541EFF14DD11E2C6FDCF" httpUri="http://treatment.plazi.org/id/039587A15333541EFF14DD11E2C6FDCF" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<subSubSection id="C326653C53335421FF14DD11E63EFDF9" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8B8336B753335421FF14DD11E583FD9C" blockId="31.[151,720,519,580]" box="[151,720,519,545]" pageId="31" pageNumber="32">
<heading id="D0CB81DB53335421FF14DD11E583FD9C" bold="true" box="[151,720,519,545]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<taxonomicName id="4C3C4D3453335421FF14DD11E583FD9C" authority="Favette et Trouessart, 1904" authorityName="Favette et Trouessart" authorityYear="1904" box="[151,720,519,545]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="gratus">
<emphasis id="B948EAA553335421FF14DD11E583FD9C" bold="true" box="[151,720,519,545]" pageId="31" pageNumber="32">
<emphasis id="B948EAA553335421FF14DD11E625FD9C" bold="true" box="[151,374,519,545]" italics="true" pageId="31" pageNumber="32">Protolichus gratus</emphasis>
<bibRefCitation id="EFAD4B4653335421FEFDDD11E583FD9C" author="Favette" box="[382,720,519,545]" pageId="31" pageNumber="36" refString="Favette, J. &amp; Trouessart, E. L. (1904) Monographie du genre Protolichus (Trt) et revision des Sarcoptides plumicoles (Analgesinae) qui vivent sur les perroquets. Memoires de la Societe zoologique de France, 17, 120 - 166 + pls. V - XV." type="journal article" year="1904">Favette et Trouessart, 1904</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8336B753335421FF14DD3CE63EFDF9" blockId="31.[151,720,519,580]" box="[151,365,554,580]" pageId="31" pageNumber="32">
(
<figureCitation id="13072A3253335421FF1CDD3CE7A9FDF9" box="[159,250,554,580]" captionStart="FIGURE 19" captionStartId="32.[151,255,1953,1977]" captionTargetBox="[182,1430,690,1891]" captionTargetId="figure@32.[151,1436,682,1927]" captionTargetPageId="32" captionText="FIGURE 19. Protolichus gratus, heteromorph male. A—dorsal view, B—ventral view." httpUri="https://zenodo.org/record/196444/files/figure.png" pageId="31" pageNumber="32">Figs 19</figureCitation>
,
<figureCitation id="13072A3253335421FE85DD3CE675FDF9" box="[262,294,554,580]" captionStart="FIGURE 20" captionStartId="33.[151,255,1947,1971]" captionTargetBox="[163,1421,190,1946]" captionTargetId="figure@33.[151,1436,171,1961]" captionTargetPageId="33" captionText="FIGURE 20. Protolichus gratus, details of heteromorph male. A—leg I, dorsal view, B—leg I, ventral view, C—leg II, dorsal view, D—leg II, ventral view, E—tarsus IV, dorsal view, F—gnathosoma, ventral view, G—opisthosoma, dorsal view." httpUri="https://zenodo.org/record/196445/files/figure.png" pageId="31" pageNumber="32">20</figureCitation>
,
<figureCitation id="13072A3253335421FEB1DD3CE603FDF9" box="[306,336,554,580]" captionStart="FIGURE 21" captionStartId="34.[151,255,1299,1323]" captionTargetBox="[158,1426,219,1274]" captionTargetId="figure@34.[151,1436,194,1278]" captionTargetPageId="34" captionText="FIGURE 21. Protolichus females, dorsal view. A— Protolichus casuarinus, B— P. gratus." httpUri="https://zenodo.org/record/196446/files/figure.png" pageId="31" pageNumber="32">21</figureCitation>
B)
</paragraph>
</subSubSection>
<subSubSection id="C326653C53335421FF14DD63E5C7FD11" pageId="31" pageNumber="32" type="reference_group">
<paragraph id="8B8336B753335421FF14DD63E5C7FD11" blockId="31.[151,1384,629,684]" pageId="31" pageNumber="32">
<treatmentCitationGroup id="AB2C119953335421FF14DD63E5C7FD11" pageId="31" pageNumber="32">
<emphasis id="B948EAA553335421FF14DD63E575FD31" box="[151,550,629,652]" italics="true" pageId="31" pageNumber="32">
<taxonomicName id="4C3C4D3453335421FF14DD63E640FD31" box="[151,275,629,652]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">Protolichus</taxonomicName>
(s. str.)
<taxonomicName id="4C3C4D3453335421FEEADD63E688FD31" box="[361,475,629,652]" class="Arachnida" family="Pterolichidae" genus="Pterolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="brachiatus">brachiatus</taxonomicName>
<taxonomicName id="4C3C4D3453335421FE61DD63E575FD31" authority="Favette and Trouessart 1904: 136" authorityName="Favette and Trouessart" authorityPageNumber="136" authorityYear="1904" box="[482,550,629,652]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="gratus">gratus</taxonomicName>
</emphasis>
<treatmentCitation id="0A9D10A653335421FDAFDD63E4C1FD31" author="Favette" box="[556,914,629,652]" page="136" pageId="31" pageNumber="32" year="1904">
<bibRefCitation id="EFAD4B4653335421FDAFDD63E40DFD31" author="Favette" box="[556,862,629,652]" pageId="31" pageNumber="36" refString="Favette, J. &amp; Trouessart, E. L. (1904) Monographie du genre Protolichus (Trt) et revision des Sarcoptides plumicoles (Analgesinae) qui vivent sur les perroquets. Memoires de la Societe zoologique de France, 17, 120 - 166 + pls. V - XV." type="journal article" year="1904">Favette and Trouessart 1904</bibRefCitation>
: 136
</treatmentCitation>
, Pl. V, fig. 3;
<treatmentCitation id="0A9D10A653335421FBA7DD63E3A3FD31" author="Dubinin" box="[1060,1264,629,652]" page="294" pageId="31" pageNumber="32" year="1956">
<bibRefCitation id="EFAD4B4653335421FBA7DD63E3E8FD31" author="Dubinin" box="[1060,1211,629,652]" pageId="31" pageNumber="36" refString="Dubinin, V. B. (1956) Feather mites (Analgesoidea). Part III. Family Pterolichidae. Fauna SSSR, Paukoobraznye, 6 (7), 1 - 813. [In Russian]" type="journal article" year="1956">Dubinin 1956</bibRefCitation>
: 294
</treatmentCitation>
, fig. 140D.
<taxonomicName id="4C3C4D3453335421FF14DD83E5C3FD11" authority="Gaud and Atyeo 1996: 132" authorityName="Gaud and Atyeo" authorityPageNumber="132" authorityYear="1996" box="[151,656,661,684]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="gratus">
<emphasis id="B948EAA553335421FF14DD83E635FD11" box="[151,358,661,684]" italics="true" pageId="31" pageNumber="32">Protolichus gratus:</emphasis>
<treatmentCitation id="0A9D10A653335421FEEFDD83E5C3FD11" author="Gaud" box="[364,656,661,684]" page="132" pageId="31" pageNumber="32" year="1996">
<bibRefCitation id="EFAD4B4653335421FEEFDD83E508FD11" author="Gaud" box="[364,603,661,684]" pageId="31" pageNumber="36" refString="Gaud, J. &amp; Atyeo, W. T. (1996) Feather mites of the World (Acarina, Astigmata): the supraspecific taxa. Musee Royal de l'Afrique Centrale, Annales, Sciences Zoologiques, 277, 1 - 193 (Part 1, text), 1 - 436 (Part 2, illustrations)." type="journal article" year="1996">Gaud and Atyeo 1996</bibRefCitation>
: 132
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C326653C53335421FF14DDCCE3A2FCA7" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8B8336B753335421FF14DDCCE3A2FCA7" blockId="31.[151,1437,730,2026]" pageId="31" pageNumber="32">
<emphasis id="B948EAA553335421FF14DDCCE6C7FD49" bold="true" box="[151,404,730,756]" pageId="31" pageNumber="32">Material examined.</emphasis>
Heteromorph male
<typeStatus id="5487881553335421FD09DDCCE5A9FD49" box="[650,762,730,756]" pageId="31" pageNumber="32" type="lectotype">lectotype</typeStatus>
(
<collectionCode id="ED2DAE7253335421FC88DDCCE417FD49" LSID="urn:lsid:biocol.org:col:15277" box="[779,836,730,756]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15277" name="Royal Ontario Museum - Herbarium" pageId="31" pageNumber="32">TRT</collectionCode>
34A1) and female
<typeStatus id="5487881553335421FBB1DDCCE385FD49" box="[1074,1238,730,756]" pageId="31" pageNumber="32" type="paralectotype">paralectotype</typeStatus>
(
<collectionCode id="ED2DAE7253335421FB64DDCCE273FD49" LSID="urn:lsid:biocol.org:col:15277" box="[1255,1312,730,756]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15277" name="Royal Ontario Museum - Herbarium" pageId="31" pageNumber="32">TRT</collectionCode>
34A2) ex
<taxonomicName id="4C3C4D3453335421FF14DC16E535FCA7" authority="Bonaparte, 1850" authorityName="Bonaparte" authorityYear="1850" box="[151,614,768,794]" class="Aves" family="Psittacidae" genus="Trichoglossus" kingdom="Animalia" order="Psittaciformes" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="forsteni">
<emphasis id="B948EAA553335421FF14DC16E6C8FCA4" box="[151,411,768,793]" italics="true" pageId="31" pageNumber="32">Trichoglossus forsteni</emphasis>
Bonaparte, 1850
</taxonomicName>
, Timor, other data unknown;
<typeStatus id="5487881553335421FC43DC16E378FCA7" box="[960,1067,768,794]" pageId="31" pageNumber="32" type="lectotype">lectotype</typeStatus>
designated here.
</paragraph>
</subSubSection>
<subSubSection id="C326653C5333541EFF45DC33E653FED1" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="description">
<paragraph id="8B8336B753335421FF45DC33E39FFAD2" blockId="31.[151,1437,730,2026]" pageId="31" pageNumber="32">
<emphasis id="B948EAA553335421FF45DC33E60CFC82" bold="true" box="[198,351,805,831]" pageId="31" pageNumber="32">Description.</emphasis>
Heteromorph male (
<typeStatus id="5487881553335421FDDCDC33E582FC82" box="[607,721,805,831]" pageId="31" pageNumber="32" type="lectotype">lectotype</typeStatus>
). Idiosoma, length x width 640 x 360. Subcapitulum with posterior margin slightly convex, postero-lateral parts crossed by oblique folds (
<figureCitation id="13072A3253335421FBD9DC5CE3E0FCD9" box="[1114,1203,842,868]" captionStart="FIGURE 20" captionStartId="33.[151,255,1947,1971]" captionTargetBox="[163,1421,190,1946]" captionTargetId="figure@33.[151,1436,171,1961]" captionTargetPageId="33" captionText="FIGURE 20. Protolichus gratus, details of heteromorph male. A—leg I, dorsal view, B—leg I, ventral view, C—leg II, dorsal view, D—leg II, ventral view, E—tarsus IV, dorsal view, F—gnathosoma, ventral view, G—opisthosoma, dorsal view." httpUri="https://zenodo.org/record/196445/files/figure.png" pageId="31" pageNumber="32">Fig. 20</figureCitation>
F). Prodorsal shield with posterior margin with wide and short median extension, length along midline 185, surface monotonously punctate, transverse area around scapular setae and near bases of epimerites Ia poorly sclerotized. Setae
<emphasis id="B948EAA553335421FA0BDC83E2CFFC13" box="[1416,1436,917,942]" italics="true" pageId="31" pageNumber="32">si</emphasis>
spiculiform, 58 long. Distance between scapular setae:
<emphasis id="B948EAA553335421FC9FDCACE404FC6E" box="[796,855,954,979]" italics="true" pageId="31" pageNumber="32">se:se</emphasis>
70,
<emphasis id="B948EAA553335421FC09DCACE4E9FC6E" box="[906,954,954,979]" italics="true" pageId="31" pageNumber="32">si:si</emphasis>
40. Hysterosoma 440 long. Hysteronotal shield: length 430, width 275, anterior margin straight, surface monotonously punctate. Opisthosomal lobes with obliquely cut apices and transverse crest posterior to bases of setae
<emphasis id="B948EAA553335421FB81DB13E34CFBA3" box="[1026,1055,1029,1054]" italics="true" pageId="31" pageNumber="32">h3</emphasis>
(
<figureCitation id="13072A3253335421FBB2DB13E3D9FBA2" box="[1073,1162,1029,1055]" captionStart="FIGURE 20" captionStartId="33.[151,255,1947,1971]" captionTargetBox="[163,1421,190,1946]" captionTargetId="figure@33.[151,1436,171,1961]" captionTargetPageId="33" captionText="FIGURE 20. Protolichus gratus, details of heteromorph male. A—leg I, dorsal view, B—leg I, ventral view, C—leg II, dorsal view, D—leg II, ventral view, E—tarsus IV, dorsal view, F—gnathosoma, ventral view, G—opisthosoma, dorsal view." httpUri="https://zenodo.org/record/196445/files/figure.png" pageId="31" pageNumber="32">Fig. 20</figureCitation>
G). Supranal concavity ovate, poorly expressed. Terminal cleft as an inverted U with slightly divergent branches, length 72, width at level of setae
<geoCoordinate id="EE08507053335421FEBEDB46E6D1FBD7" box="[317,386,1104,1130]" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="925" value="1.5">
<emphasis id="B948EAA553335421FEBEDB46E609FBD4" box="[317,346,1104,1129]" italics="true" pageId="31" pageNumber="32">e1</emphasis>
30
</geoCoordinate>
, greatest width 40. Lateral setae
<emphasis id="B948EAA553335421FC8EDB46E479FBD4" box="[781,810,1104,1129]" italics="true" pageId="31" pageNumber="32">c2</emphasis>
spiculiform, 58 long; setae
<emphasis id="B948EAA553335421FBF9DB46E3C4FBD4" box="[1146,1175,1104,1129]" italics="true" pageId="31" pageNumber="32">d2</emphasis>
missing in specimen; setae
<emphasis id="B948EAA553335421FF5BDB63E7A7FB33" box="[216,244,1141,1166]" italics="true" pageId="31" pageNumber="32">e2</emphasis>
thickened, extending to level of lobar apices, 100102 long; setae
<emphasis id="B948EAA553335421FC7DDB63E34AFB33" box="[1022,1049,1141,1166]" italics="true" pageId="31" pageNumber="32">e1</emphasis>
setiform, situated approximately at level of macrosetae
<emphasis id="B948EAA553335421FE33DB8DE69CFB09" box="[432,463,1179,1204]" italics="true" pageId="31" pageNumber="32">h2</emphasis>
, setae
<emphasis id="B948EAA553335421FDA5DB8DE56CFB09" box="[550,575,1179,1204]" italics="true" pageId="31" pageNumber="32">f2</emphasis>
leaf-like, with 1 or 2 teeth on distal part, setae
<emphasis id="B948EAA553335421FB13DB8DE3FEFB09" box="[1168,1197,1179,1204]" italics="true" pageId="31" pageNumber="32">e1</emphasis>
situated at level of macrosetae
<emphasis id="B948EAA553335421FEA2DBD6E66DFB64" box="[289,318,1216,1241]" italics="true" pageId="31" pageNumber="32">h2</emphasis>
. Distance between dorsal setae:
<emphasis id="B948EAA553335421FD39DBD6E5AEFB64" box="[698,765,1216,1241]" italics="true" pageId="31" pageNumber="32">c2:d2</emphasis>
140,
<emphasis id="B948EAA553335421FCBDDBD6E4D3FB64" box="[830,896,1216,1241]" italics="true" pageId="31" pageNumber="32">d2:e2</emphasis>
165,
<emphasis id="B948EAA553335421FC42DBD6E350FB64" box="[961,1027,1216,1241]" italics="true" pageId="31" pageNumber="32">e2:h3</emphasis>
80,
<emphasis id="B948EAA553335421FBB5DBD6E324FB64" box="[1078,1143,1216,1241]" italics="true" pageId="31" pageNumber="32">
e2:
<geoCoordinate id="EE08507053335421FBDFDBD6E324FB64" box="[1116,1143,1216,1241]" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="925" value="1.8666667">e1</geoCoordinate>
</emphasis>
52,
<emphasis id="B948EAA553335421FB29DBD6E3B8FB64" box="[1194,1259,1216,1241]" italics="true" pageId="31" pageNumber="32">e2:e2</emphasis>
135,
<emphasis id="B948EAA553335421FAAFDBD6E222FB64" box="[1324,1393,1216,1241]" italics="true" pageId="31" pageNumber="32">h2:h2</emphasis>
95,
<emphasis id="B948EAA553335421FF14DBF3E788FB43" box="[151,219,1253,1278]" italics="true" pageId="31" pageNumber="32">h3:h3</emphasis>
88,
<emphasis id="B948EAA553335421FE8CDBF3E639FB43" box="[271,362,1253,1278]" italics="true" pageId="31" pageNumber="32">ps1:ps1</emphasis>
56,
<emphasis id="B948EAA553335421FE1DDBF3E6B2FB43" box="[414,481,1253,1278]" italics="true" pageId="31" pageNumber="32">e2:h2</emphasis>
58,
<emphasis id="B948EAA553335421FD97DBF3E50BFB43" box="[532,600,1253,1278]" italics="true" pageId="31" pageNumber="32">h2:h3</emphasis>
25,
<emphasis id="B948EAA553335421FD0EDBF3E58EFB43" box="[653,733,1253,1278]" italics="true" pageId="31" pageNumber="32">ps1:h3</emphasis>
12. Genital apparatus situated at level of anterior margin of trochanters IV,
<date id="FF82107753335421FECEDA1CE6FCFA99" box="[333,431,1290,1316]" pageId="31" pageNumber="32" value="1930-10-22">30 x 22.</date>
Paragenital apodemes with anterior end extended to midlevel of coxal fields III, not connected to epimerites IIIa, apodemes not connected to each other by transverse bridge (
<figureCitation id="13072A3253335421FB36DA26E258FAF7" box="[1205,1291,1328,1354]" captionStart="FIGURE 19" captionStartId="32.[151,255,1953,1977]" captionTargetBox="[182,1430,690,1891]" captionTargetId="figure@32.[151,1436,682,1927]" captionTargetPageId="32" captionText="FIGURE 19. Protolichus gratus, heteromorph male. A—dorsal view, B—ventral view." httpUri="https://zenodo.org/record/196444/files/figure.png" pageId="31" pageNumber="32">Fig. 19</figureCitation>
B). Distance between ventral setae:
<emphasis id="B948EAA553335421FE20DA43E6B5FAD3" box="[419,486,1365,1390]" italics="true" pageId="31" pageNumber="32">3b:3a</emphasis>
10,
<emphasis id="B948EAA553335421FD99DA43E503FAD3" box="[538,592,1365,1390]" italics="true" pageId="31" pageNumber="32">3a:g</emphasis>
28,
<emphasis id="B948EAA553335421FD00DA43E5EBFAD3" box="[643,696,1365,1390]" italics="true" pageId="31" pageNumber="32">g:4a</emphasis>
75,
<emphasis id="B948EAA553335421FD6FDA43E468FAD3" box="[748,827,1365,1390]" italics="true" pageId="31" pageNumber="32">4a:ps3</emphasis>
112. Diameter of anal suckers 23.
</paragraph>
<paragraph id="8B8336B753335421FF47DA6CE460F959" blockId="31.[151,1437,730,2026]" pageId="31" pageNumber="32">
Legs II approximately 1.41.5 times longer and thicker than legs I. Tarsus I with crest-like ventral extension stretching along entire segment, proximal end of this extension widely rounded (
<figureCitation id="13072A3253335421FB70DAB6E202FA07" box="[1267,1361,1440,1466]" captionStart="FIGURE 20" captionStartId="33.[151,255,1947,1971]" captionTargetBox="[163,1421,190,1946]" captionTargetId="figure@33.[151,1436,171,1961]" captionTargetPageId="33" captionText="FIGURE 20. Protolichus gratus, details of heteromorph male. A—leg I, dorsal view, B—leg I, ventral view, C—leg II, dorsal view, D—leg II, ventral view, E—tarsus IV, dorsal view, F—gnathosoma, ventral view, G—opisthosoma, dorsal view." httpUri="https://zenodo.org/record/196445/files/figure.png" pageId="31" pageNumber="32">Figs 20</figureCitation>
A, B). Tibia, genu I without ventral apophysis; genual seta
<emphasis id="B948EAA553335421FD7ADAD3E448FA63" box="[761,795,1477,1502]" italics="true" pageId="31" pageNumber="32">cG</emphasis>
I setiform, strongly thickened basally, smooth. Femur I with longitudinal ventral crest. Tarsus II with large tubercle-like ventral extension covered with verrucae and bearing setae
<emphasis id="B948EAA553335421FEBFD906E60CF994" box="[316,351,1552,1577]" italics="true" pageId="31" pageNumber="32">wa</emphasis>
and
<emphasis id="B948EAA553335421FE19D906E6E7F994" box="[410,436,1552,1577]" italics="true" pageId="31" pageNumber="32">ra</emphasis>
on apex; dorsal surface between bases of solenidion ω
<emphasis id="B948EAA553335421FBCBD906E304F994" box="[1096,1111,1552,1577]" italics="true" pageId="31" pageNumber="32">1</emphasis>
and seta
<emphasis id="B948EAA553335421FB4BD906E384F994" box="[1224,1239,1552,1577]" italics="true" pageId="31" pageNumber="32">d</emphasis>
slightly inflated and verrucous (
<figureCitation id="13072A3253335421FED1D923E6FCF9F2" box="[338,431,1589,1615]" captionStart="FIGURE 20" captionStartId="33.[151,255,1947,1971]" captionTargetBox="[163,1421,190,1946]" captionTargetId="figure@33.[151,1436,171,1961]" captionTargetPageId="33" captionText="FIGURE 20. Protolichus gratus, details of heteromorph male. A—leg I, dorsal view, B—leg I, ventral view, C—leg II, dorsal view, D—leg II, ventral view, E—tarsus IV, dorsal view, F—gnathosoma, ventral view, G—opisthosoma, dorsal view." httpUri="https://zenodo.org/record/196445/files/figure.png" pageId="31" pageNumber="32">Figs 20</figureCitation>
C, D). Tibiae II with tubercle-like antero-ventral apophysis with verrucous surface. Genu II with ventral surface verrucous, genual seta
<emphasis id="B948EAA553335421FD77D94DE445F9C9" box="[756,790,1627,1652]" italics="true" pageId="31" pageNumber="32">cG</emphasis>
II strongly thickened, with verrucous distal part, 1.82 times longer than segment. Femur II with two spine-like dorso-apical apophyses and with verrucous ventral surface. Solenidia φ of tibiae I, II verrucous in basal part. Tarsus IV slightly longer than corresponding tibia; setae
<emphasis id="B948EAA553335421FF5AD9DDE7BCF959" box="[217,239,1739,1764]" italics="true" pageId="31" pageNumber="32">d,</emphasis>
e spiculiform, with distinct basal ring (
<figureCitation id="13072A3253335421FD3FD9DCE441F959" box="[700,786,1738,1764]" captionStart="FIGURE 20" captionStartId="33.[151,255,1947,1971]" captionTargetBox="[163,1421,190,1946]" captionTargetId="figure@33.[151,1436,171,1961]" captionTargetPageId="33" captionText="FIGURE 20. Protolichus gratus, details of heteromorph male. A—leg I, dorsal view, B—leg I, ventral view, C—leg II, dorsal view, D—leg II, ventral view, E—tarsus IV, dorsal view, F—gnathosoma, ventral view, G—opisthosoma, dorsal view." httpUri="https://zenodo.org/record/196445/files/figure.png" pageId="31" pageNumber="32">Fig. 20</figureCitation>
E).
</paragraph>
<paragraph id="8B8336B753335421FF45D9E6E560F8B7" blockId="31.[151,1437,730,2026]" box="[198,563,1776,1802]" pageId="31" pageNumber="32">Homeomorph male. Unknown.</paragraph>
<paragraph id="8B8336B75333541EFF45D803E653FED1" blockId="31.[151,1437,730,2026]" lastBlockId="32.[151,1436,152,626]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
Female (
<typeStatus id="5487881553335421FEB3D803E686F892" box="[304,469,1813,1839]" pageId="31" pageNumber="32" type="paralectotype">paralectotype</typeStatus>
). Subcapitulum with transverse striae in median part, posterior margin strongly convex. Idiosoma, length x width, 470 x 275. Prodorsal shield with posterior margin slightly convex, length along midline 130, surface monotonously punctate, transverse bar across scapular setae weakly sclerotized. Setae
<emphasis id="B948EAA553335421FF5DD893E7A1F823" box="[222,242,1925,1950]" italics="true" pageId="31" pageNumber="32">si</emphasis>
spiculiform, 3032 long. Distance between scapular setae:
<emphasis id="B948EAA553335421FC37D893E4BCF823" box="[948,1007,1925,1950]" italics="true" pageId="31" pageNumber="32">se:se</emphasis>
90,
<emphasis id="B948EAA553335421FBA0D893E307F823" box="[1059,1108,1925,1950]" italics="true" pageId="31" pageNumber="32">si:si</emphasis>
60. Hysterosoma 340 long. Hysteronotal shield: length 320, width 230, anterior margin slightly concave, surface monotonously punctate; subtegumental sclerotized bands posterior to setae
<emphasis id="B948EAA553335421FD6AD8C6E456F854" box="[745,773,2000,2025]" italics="true" pageId="31" pageNumber="32">e2</emphasis>
slightly enlarged (
<figureCitation id="13072A3253335421FC5CD8C6E366F857" box="[991,1077,2000,2026]" captionStart="FIGURE 21" captionStartId="34.[151,255,1299,1323]" captionTargetBox="[158,1426,219,1274]" captionTargetId="figure@34.[151,1436,194,1278]" captionTargetPageId="34" captionText="FIGURE 21. Protolichus females, dorsal view. A— Protolichus casuarinus, B— P. gratus." httpUri="https://zenodo.org/record/196446/files/figure.png" pageId="31" pageNumber="32">Fig. 21</figureCitation>
B). Setae
<emphasis id="B948EAA553335421FB26D8C6E393F854" box="[1189,1216,2000,2025]" italics="true" pageId="31" pageNumber="32">c2</emphasis>
missing (probably spiculiform), setae
<emphasis id="B948EAA5530C541EFEFDDF8EE6CFFF0C" box="[382,412,152,177]" italics="true" pageId="32" pageNumber="33">d2</emphasis>
setiform, short; setae
<emphasis id="B948EAA5530C541EFD2BDF8EE597FF0C" box="[680,708,152,177]" italics="true" pageId="32" pageNumber="33">e2</emphasis>
thick setiform, setae
<emphasis id="B948EAA5530C541EFC4BDF8EE48CFF0C" box="[968,991,152,177]" italics="true" pageId="32" pageNumber="33">f2</emphasis>
with narrow membranous expansion and acute spine in medial part; setae
<emphasis id="B948EAA5530C541EFDEBDFABE5C1FF6B" box="[616,658,189,214]" italics="true" pageId="32" pageNumber="33">ps2</emphasis>
missing in specimen; setae
<emphasis id="B948EAA5530C541EFC70DFABE34EFF6B" box="[1011,1053,189,214]" italics="true" pageId="32" pageNumber="33">ps1</emphasis>
partly broken in specimen but certainly long, setae
<emphasis id="B948EAA5530C541EFE0FDFF4E6FBFF46" box="[396,424,226,251]" italics="true" pageId="32" pageNumber="33">e1</emphasis>
situated slightly posterior to level of cupules
<emphasis id="B948EAA5530C541EFC45DFF4E4B0FF46" box="[966,995,226,251]" italics="true" pageId="32" pageNumber="33">im</emphasis>
. Length of lateral hysteronotal setae:
<emphasis id="B948EAA5530C541EFF14DE1EE7E6FE9C" box="[151,181,264,289]" italics="true" pageId="32" pageNumber="33">d2</emphasis>
10,
<emphasis id="B948EAA5530C541EFF65DE1EE651FE9C" box="[230,258,264,289]" italics="true" pageId="32" pageNumber="33">e2</emphasis>
over 65 (apical parts broken off),
<emphasis id="B948EAA5530C541EFD13DE1EE5F4FE9C" box="[656,679,264,289]" italics="true" pageId="32" pageNumber="33">f2</emphasis>
48. Distance between hysterosomal setae:
<emphasis id="B948EAA5530C541EFB1FDE1EE38DFE9C" box="[1180,1246,264,289]" italics="true" pageId="32" pageNumber="33">c2:d2</emphasis>
120,
<emphasis id="B948EAA5530C541EFA9CDE1EE231FE9C" box="[1311,1378,264,289]" italics="true" pageId="32" pageNumber="33">d2:e2</emphasis>
142,
<emphasis id="B948EAA5530C541EFF14DE3BE789FEFB" box="[151,218,301,326]" italics="true" pageId="32" pageNumber="33">e2:h3</emphasis>
62,
<emphasis id="B948EAA5530C541EFE8DDE3BE603FEFB" box="[270,336,301,326]" italics="true" pageId="32" pageNumber="33">e2:e2</emphasis>
170,
<emphasis id="B948EAA5530C541EFE08DE3BE69FFEFB" box="[395,460,301,326]" italics="true" pageId="32" pageNumber="33">
e1:
<geoCoordinate id="EE085070530C541EFE2DDE3BE69FFEFB" box="[430,460,301,326]" direction="east" orientation="longitude" pageId="32" pageNumber="33" precision="925" value="3.0">e2</geoCoordinate>
</emphasis>
60,
<emphasis id="B948EAA5530C541EFD82DE3BE515FEFB" box="[513,582,301,326]" italics="true" pageId="32" pageNumber="33">h2:h2</emphasis>
85,
<emphasis id="B948EAA5530C541EFDF8DE3BE585FEFB" box="[635,726,301,326]" italics="true" pageId="32" pageNumber="33">ps1:ps1</emphasis>
28. Epigynum bow-shaped,
<date id="FF821077530C541EFBA9DE3BE3D4FEFA" box="[1066,1159,301,327]" pageId="32" pageNumber="33" value="1950-10-18">18 x 50</date>
. Setae
<emphasis id="B948EAA5530C541EFB58DE3BE255FEFB" box="[1243,1286,301,326]" italics="true" pageId="32" pageNumber="33">mG</emphasis>
of genu I, II setiform.
</paragraph>
</subSubSection>
<subSubSection id="C326653C530C541EFF45DE61E415FE2F" box="[198,838,375,402]" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8B8336B7530C541EFF45DE61E415FE2F" blockId="32.[151,1436,152,626]" box="[198,838,375,402]" pageId="32" pageNumber="33">
<emphasis id="B948EAA5530C541EFF45DE61E68DFE2C" bold="true" box="[198,478,375,401]" pageId="32" pageNumber="33">
<typeStatus id="54878815530C541EFF45DE61E650FE2C" box="[198,259,375,401]" pageId="32" pageNumber="33">Type</typeStatus>
host and locality.
</emphasis>
<taxonomicName id="4C3C4D34530C541EFE66DE6EE415FE2F" authority="Timor." authorityName="Timor." box="[485,838,376,402]" class="Aves" family="Psittacidae" genus="Trichoglossus" kingdom="Animalia" order="Psittaciformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="forsteni">
<emphasis id="B948EAA5530C541EFE66DE6EE5BAFE2C" box="[485,745,376,401]" italics="true" pageId="32" pageNumber="33">Trichoglossus forsteni</emphasis>
, Timor.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C326653C530C541EFF45DE8BE2C6FDCF" pageId="32" pageNumber="33" type="discussion">
<paragraph id="8B8336B7530C541EFF45DE8BE2C6FDCF" blockId="32.[151,1436,152,626]" pageId="32" pageNumber="33">
<emphasis id="B948EAA5530C541EFF45DE8BE613FE0A" bold="true" box="[198,320,413,439]" pageId="32" pageNumber="33">Remarks.</emphasis>
This species was originally described from
<taxonomicName id="4C3C4D34530C541EFCCADE8BE347FE0B" box="[841,1044,413,438]" class="Aves" family="Psittacidae" genus="Pseudeos" kingdom="Animalia" order="Psittaciformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="fuscata">
<emphasis id="B948EAA5530C541EFCCADE8BE347FE0B" box="[841,1044,413,438]" italics="true" pageId="32" pageNumber="33">Pseudeos fuscata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3C4D34530C541EFBCCDE8BE207FE0B" box="[1103,1364,413,438]" class="Aves" family="Psittacidae" genus="Trichoglossus" kingdom="Animalia" order="Psittaciformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="forsteni">
<emphasis id="B948EAA5530C541EFBCCDE8BE207FE0B" box="[1103,1364,413,438]" italics="true" pageId="32" pageNumber="33">Trichoglossus forsteni</emphasis>
</taxonomicName>
, from New
<collectingCountry id="F32B7627530C541EFF55DED4E678FE61" box="[214,299,450,476]" name="Guinea-Bissau" pageId="32" pageNumber="33">Guinea</collectingCountry>
and Timor, respectively (
<bibRefCitation id="EFAD4B46530C541EFDDADED4E4CAFE61" author="Favette" box="[601,921,450,476]" pageId="32" pageNumber="36" refString="Favette, J. &amp; Trouessart, E. L. (1904) Monographie du genre Protolichus (Trt) et revision des Sarcoptides plumicoles (Analgesinae) qui vivent sur les perroquets. Memoires de la Societe zoologique de France, 17, 120 - 166 + pls. V - XV." type="journal article" year="1904">Favette &amp; Trouessart 1904</bibRefCitation>
). One male and one female from
<taxonomicName id="4C3C4D34530C541EFAA6DED4E2C9FE66" box="[1317,1434,450,475]" class="Aves" family="Psittacidae" genus="Trichoglossus" kingdom="Animalia" order="Psittaciformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="forsteni">
<emphasis id="B948EAA5530C541EFAA6DED4E2C9FE66" box="[1317,1434,450,475]" italics="true" pageId="32" pageNumber="33">T. forsteni</emphasis>
</taxonomicName>
are present in the Trouessart collection; these slides are marked by W.T. Atyeo as
<typeStatus id="54878815530C541EFBD0DEFEE3EDFDBF" box="[1107,1214,488,514]" pageId="32" pageNumber="33" type="lectotype">lectotype</typeStatus>
and
<typeStatus id="54878815530C541EFB75DEFEE2CBFDBF" box="[1270,1432,488,514]" pageId="32" pageNumber="33" type="paralectotype">paralectotype</typeStatus>
. No specimens form
<taxonomicName id="4C3C4D34530C541EFE01DD1BE550FD9B" box="[386,515,525,550]" class="Aves" family="Psittacidae" genus="Pseudeos" kingdom="Animalia" order="Psittaciformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="fuscata">
<emphasis id="B948EAA5530C541EFE01DD1BE550FD9B" box="[386,515,525,550]" italics="true" pageId="32" pageNumber="33">Ps. fuscata</emphasis>
</taxonomicName>
labeled as
<taxonomicName id="4C3C4D34530C541EFD07DD1BE40DFD9B" box="[644,862,525,550]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="gratus">
<emphasis id="B948EAA5530C541EFD07DD1BE40DFD9B" box="[644,862,525,550]" italics="true" pageId="32" pageNumber="33">Protolichus gratus</emphasis>
</taxonomicName>
were found; apparently they were not retained in the Trouessart collection. The male from
<taxonomicName id="4C3C4D34530C541EFDFCDD24E5A6FDF6" box="[639,757,562,587]" class="Aves" family="Psittacidae" genus="Trichoglossus" kingdom="Animalia" order="Psittaciformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="forsteni">
<emphasis id="B948EAA5530C541EFDFCDD24E5A6FDF6" box="[639,757,562,587]" italics="true" pageId="32" pageNumber="33">T. forsteni</emphasis>
</taxonomicName>
corresponds quite well to the photo of
<taxonomicName id="4C3C4D34530C541EFB46DD24E27DFDF6" box="[1221,1326,562,587]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="gratus">
<emphasis id="B948EAA5530C541EFB46DD24E27DFDF6" box="[1221,1326,562,587]" italics="true" pageId="32" pageNumber="33">P. gratus</emphasis>
</taxonomicName>
given by
<bibRefCitation id="EFAD4B46530C541EFF14DD4EE6ABFDCF" author="Favette" box="[151,504,600,626]" pageId="32" pageNumber="36" refString="Favette, J. &amp; Trouessart, E. L. (1904) Monographie du genre Protolichus (Trt) et revision des Sarcoptides plumicoles (Analgesinae) qui vivent sur les perroquets. Memoires de la Societe zoologique de France, 17, 120 - 166 + pls. V - XV." type="journal article" year="1904">Favette and Trouessart (1904)</bibRefCitation>
, particularly in the structure of genual seta
<emphasis id="B948EAA5530C541EFC6DDD4EE343FDCC" box="[1006,1040,600,625]" italics="true" pageId="32" pageNumber="33">cG</emphasis>
II and ventral tubercle of tibia II.
</paragraph>
</subSubSection>
</treatment>
</document>