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<document id="C8460E4E2410D45DC3B42F4808CEB23F" ID-CLB-Dataset="306634" ID-DOI="10.1093/zoolinnean/zlae137" ID-GBIF-Dataset="b9ccfb40-9194-4228-abd2-acaf275f5749" ID-ISSN="0024-4082" ID-Zenodo-Dep="14516114" ID-ZooBank="9059AA2-5086-46AD-85C6-DBDA56CA72E0" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1734033959621" checkinUser="plazi" docAuthor="Simon-Pražák, Jan, Yamamoto, Shûhei, Lackner, Tomáš, Fikáček, Martin, Prokop, Jakub &amp; Caterino, Michael S" docDate="2024" docId="038C87BE1701817A4C9FF9C6FF707CBF" docLanguage="en" docName="zlae137.pdf" docOrigin="Zoological Journal of the Linnean Society (zlae 137) 202 (3)" docSource="https://doi.org/10.1093/zoolinnean/zlae137" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Palaeabraeus glabrus Simon-Pražák, Yamamoto, Lackner, Fikáček, Prokop &amp; Caterino, 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="13" masterDocId="FFB5FFC6170881764F58FFA0FFA07B0E" masterDocTitle="A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="10" updateTime="1734537818702" updateUser="ExternalLinkService">
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<mods:title id="8A366EAC042B44B92EBA498157ADB72A">A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)</mods:title>
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<mods:namePart id="3EB45693D7B1735121A31B7FC0F5F07A">Simon-Pražák, Jan</mods:namePart>
<mods:affiliation id="C2C7FC90443C4ED35A6C505C2FA62643">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic &amp; Museum of Eastern Bohemia in Hradec Králové, Eliščino nábřeží 465, 500 03 Hradec Králové 3, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="F6B63D6B6A82D8295531B0D3C5F8D61A" type="email">j.prazak@muzeumhk.cz</mods:nameIdentifier>
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<mods:namePart id="909FD34F2FDA7B66A830EAFE97B82FE0">Yamamoto, Shûhei</mods:namePart>
<mods:affiliation id="14347B832E46C6CAA1A5F0BA62AEC15E">The Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo 060 - 0810, Japan</mods:affiliation>
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<mods:namePart id="64DAACF3E9432938D49F498C51BFAC40">Lackner, Tomáš</mods:namePart>
<mods:affiliation id="2DEB93D10A972FE900CC8380F4F82C15">Department of Environmental Systems Science, ETH Zürich, Weinbergstrasse 56, 8092 Zürich, Switzerland</mods:affiliation>
</mods:name>
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<mods:namePart id="73E22F7EA05A58F4844DE82372CC4155">Fikáček, Martin</mods:namePart>
<mods:affiliation id="554532724BF765A5A8AC2CBB0695DEDB">Department of Biological Sciences, National Sun Yat-sen University, No. 70, Lienhai Rd., Kaohsiung 80424, Taiwan &amp; Department of Entomology, National Museum, Cirkusová 1740 CZ- 19300 Praha 9, Czech Republic</mods:affiliation>
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<mods:namePart id="35530A59361AB2A7E50A2C9F900B6A7B">Prokop, Jakub</mods:namePart>
<mods:affiliation id="5FEC6BA24CB8855F8AAC5D16408E1333">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic &amp; Department of Plant and Environmental Sciences, Clemson University, Clemson, SC 29634, United States</mods:affiliation>
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<mods:namePart id="92EBD8DB7D20B31350F3383FB095C8C2">Caterino, Michael S</mods:namePart>
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<mods:title id="A9B426972CCF3A1F58B5C34FC926EE6B">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="6AD689693856F4750D7C136D289C14E0">2024</mods:date>
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<mods:title id="0831F89B9A067CAAD1F5E31E21774B14">zlae 137</mods:title>
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<mods:number id="D023728BF33196C150C892858325FE5D">2024-11-20</mods:number>
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<mods:number id="9F64B63F3FD151AB3E6DF1D475C350C0">202</mods:number>
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<treatment id="038C87BE1701817A4C9FF9C6FF707CBF" ID-DOI="http://doi.org/10.5281/zenodo.14424556" ID-Zenodo-Dep="14424556" LSID="urn:lsid:plazi:treatment:038C87BE1701817A4C9FF9C6FF707CBF" httpUri="http://treatment.plazi.org/id/038C87BE1701817A4C9FF9C6FF707CBF" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10" scope_family="Histeridae" scope_order="Coleoptera" scope_subFamily="Abraeinae" scope_tribe="Abraeini">
<subSubSection id="C33F65231701817F4C9FF9C6FAB67D8F" box="[967,1302,1638,1665]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8B9A36A81701817F4C9FF9C6FAB67D8F" blockId="9.[967,1302,1638,1665]" box="[967,1302,1638,1665]" pageId="9" pageNumber="10">
<heading id="D0D281C41701817F4C9FF9C6FAB67D8F" bold="true" box="[967,1302,1638,1665]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
<emphasis id="B951EABA1701817F4C9FF9C6FAB67D8F" bold="true" box="[967,1302,1638,1665]" pageId="9" pageNumber="10">
<taxonomicName id="4C254D2B1701817F4C9FF9C6FB1A7D8F" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[967,1210,1638,1665]" class="Insecta" family="Histeridae" genus="Palaeabraeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="glabrus" status="sp. nov.">
<emphasis id="B951EABA1701817F4C9FF9C6FB1A7D8F" bold="true" box="[967,1210,1638,1665]" italics="true" pageId="9" pageNumber="10">Palaeabraeus glabrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C11701817F4BE7F9C7FAB67D8F" box="[1215,1302,1639,1665]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C33F65231701817F4B1FF905FB367DB1" box="[1095,1174,1701,1727]" pageId="9" pageNumber="10" type="description">
<paragraph id="8B9A36A81701817F4B1FF905FB367DB1" blockId="9.[1095,1174,1701,1727]" box="[1095,1174,1701,1727]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4B1FF905FB367DB1" bold="true" box="[1095,1174,1701,1727]" pageId="9" pageNumber="10">
(
<figureCitation id="131E2A2D1701817F4B0AF905FB2B7DB1" box="[1106,1163,1701,1727]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." figureDoi="http://doi.org/10.5281/zenodo.14516132" httpUri="https://zenodo.org/record/14516132/files/figure.png" pageId="9" pageNumber="10">Fig. 8</figureCitation>
)
</emphasis>
</paragraph>
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<subSubSection id="C33F65231701817F4C72F946FA9A7C33" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8B9A36A81701817F4C72F946FA9A7C33" blockId="9.[810,1460,1766,1853]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4C72F946FC267DF0" box="[810,902,1766,1790]" italics="true" pageId="9" pageNumber="10">Material:</emphasis>
<materialsCitation id="3B4D3CF51701817F4CD5F946FA957C33" collectionCode="HUM" location="Kachin" pageId="9" pageNumber="10" specimenCode="SEHU-0000121213" specimenCount="1" stateProvince="Kachin">
<collectingRegion id="49E1F84A1701817F4CD5F946FC777DF0" box="[909,983,1766,1790]" country="Myanmar" name="Kachin" pageId="9" pageNumber="10">Kachin</collectingRegion>
amber (
<emphasis id="B951EABA1701817F4B69F947FB9A7DF0" box="[1073,1082,1767,1790]" italics="true" pageId="9" pageNumber="10">c</emphasis>
. 99 Mya). Amber piece dimensions: 5 × 2 ×
<quantity id="4CDD9B4D1701817F4C25F8A5FC187C13" box="[893,952,1797,1821]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="10" unit="mm" value="2.0">2 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at
<collectionCode id="ED34AE6D1701817F4B54F885FBEB7C32" box="[1036,1099,1829,1852]" pageId="9" pageNumber="10">HUM</collectionCode>
(
<specimenCode id="DB839ED31701817F4B0FF885FA8E7C33" box="[1111,1326,1829,1853]" pageId="9" pageNumber="10">SEHU-0000121213</specimenCode>
)
</materialsCitation>
.
</paragraph>
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<subSubSection id="C33F65231701817A4C72F8FFFDEE7931" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10" type="description">
<paragraph id="8B9A36A81701817A4C72F8FFFDEE7931" blockId="9.[810,1459,1886,1973]" lastBlockId="12.[129,778,144,575]" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4C72F8FFFC1C7C78" box="[810,956,1887,1910]" italics="true" pageId="9" pageNumber="10">Measurements:</emphasis>
Head width:
<quantity id="4CDD9B4D1701817F4B0AF8FEFB0D7C7B" box="[1106,1197,1886,1910]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="10" unit="mm" value="0.23">0.23 mm</quantity>
; width between anterior pronotal angles:
<quantity id="4CDD9B4D1701817F4C86F8DDFB997C9B" box="[990,1081,1917,1941]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="10" unit="mm" value="0.25">0.25 mm</quantity>
; width between posterior pronotal angles:
<quantity id="4CDD9B4D1701817F4CDAF83CFC6F7CBA" box="[898,975,1948,1972]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="10" unit="mm" value="0.5">0.5 mm</quantity>
; pronotal length:
<quantity id="4CDD9B4D1701817F4BF1F83DFAA47CBA" box="[1193,1284,1949,1973]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="10" unit="mm" value="0.23">0.23 mm</quantity>
; sutural elytral length:
<quantity id="4CDD9B4D1702817C4F8CF990FE827D49" box="[212,290,1584,1608]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="11" unit="mm" value="0.6">0.6 mm</quantity>
; elytral width (in pair, across widest point):
<quantity id="4CDD9B4D1702817C4FD9F9EFFF7D7D69" box="[129,221,1615,1639]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="10" pageNumber="11" unit="mm" value="0.56">0.56 mm</quantity>
.—
<emphasis id="B951EABA1702817C4FA4F9EFFE9C7D69" bold="true" box="[252,316,1615,1639]" italics="true" pageId="10" pageNumber="11">Body:</emphasis>
Oval, tapered anteriorly and posteriorly, strongly convex, impunctate, glabrous.—
<emphasis id="B951EABA1702817C4D75F9CEFDCF7D88" bold="true" box="[557,623,1646,1670]" italics="true" pageId="10" pageNumber="11">Head:</emphasis>
Frons convex, weakly and sparsely punctate; frontal stria absent; narrow epistoma with fine, sparse setae; eye convex, coarsely faceted, anterior portion exposed laterally; labrum short, convex, bearing two dominant setae curving ventrad and several secondary setae; mandibles largely concealed, with a few setae on lateral margins; galeae exposed, densely setose; maxillary palps long, penultimate palpomere about third length of terminal; terminal palpomere swollen at base, tapered apically, shallowly concave on inner surface; labial palpi with apical palpomere long, slender, penultimate palpomere about half its length; antennal insertions slightly open anteriorly, under weak frontolateral bulge; scape short, weakly widening apically, curved posteriad below eye; pedicel cylindrical, almost as long as antennomeres 3 and 4 combined; antennomeres 38 widening slightly toward club, each with peripheral setae, 36 moniliform, 7 and 8 with sharper basal margins, each narrowed to apex; antennal club round, slightly tapered to apex, tomentose, without annuli.—
<emphasis id="B951EABA1702817C4A35F8ABFA627C2D" bold="true" box="[1389,1474,1803,1827]" italics="true" pageId="10" pageNumber="11">Thorax:</emphasis>
Pronotal disk convex, almost twice as wide as long, sides weakly curved, evenly tapered from base to apex; anterior emargination deep; anterior corners strongly downturned, acute; marginal pronotal stria present, distant from anterior margin, closer to lateral margins. Elytra strongly convex, apically truncate, glabrous, and impunctate, without distinct dorsal striae; epipleuron with single weak submarginal carina. Wings fully developed (protruding beyond elytra). Prosternum short, lacking distinct lobe; keel shallowly emarginate at base, with lateral striae narrowed at middle, widened to base and apex; prosternum incised on either side for reception of the antennal funicle; antennal cavities located anterolaterally, well defined by marginal ridge posteriorly and laterally. Mesoventrite rather long, more than half as long as intercoxal width, strongly developed anteriorly, with broadly acute median process inserting into base of prosternal keel; marginal mesoventral stria complete, broadly rounded; mesometaventral suture well developed; metaventrite slightly impressed along midline; postmesocoxal stria complete, strongly recurved behind coxa to mesepimeron; lateral metaventral stria not evident; metepisternum broadly exposed, tapered beneath anteriorly produced metepimeron.—
<emphasis id="B951EABA1703817D4BEDF893FB4D7C44" bold="true" box="[1205,1261,1843,1866]" italics="true" pageId="11" pageNumber="12">Legs:</emphasis>
Profemur slightly convex along inner margin, both inner and outer margins setose; protibia weakly curved, widened slightly to apex, setose and with one subapical denticle along outer margin; protarsal groove well developed; protibial spurs absent; mesofemur similar to profemur; mesotibia slender, widened slightly to apex, with fine setae along inner and outer margins, few stronger spines present near inner apical corners; metafemur and metatibia similar to those of mesothorax; tarsi 5-5-5, tarsomeres 14 similar in length, ultimate tarsomeres as long as tarsomeres 24 combined; protarsomeres 14 with expanded and flattened ventral setae (probable sexual dimorphism); all tarsomeres with numerous ventral setae (not just one apicoventral pair); tarsal claws simple.—
<emphasis id="B951EABA1704817A4FBDFE09FEF17ACF" bold="true" box="[229,337,425,449]" italics="true" pageId="12" pageNumber="13">Abdomen:</emphasis>
Abdominal ventrite 1 flat between coxae, with slightly curved parallel striae from inner corner of metacoxa to ventrite 2; ventrites 25 short; propygidium shorter than pygidium, glabrous; pygidium slightly longer than basal width, weakly convex, apically rounded, impunctate.
</paragraph>
</subSubSection>
<caption id="DF5A66201702817C4FD9FA6CFAB07D0E" ID-DOI="http://doi.org/10.5281/zenodo.14516130" ID-Zenodo-Dep="14516130" httpUri="https://zenodo.org/record/14516130/files/figure.png" pageId="10" pageNumber="11" startId="10.[129,194,1484,1508]" targetBox="[129,1473,144,1456]" targetPageId="10" targetType="figure">
<paragraph id="8B9A36A81702817C4FD9FA6CFAB07D0E" blockId="10.[129,1382,1484,1536]" pageId="10" pageNumber="11">
<emphasis id="B951EABA1702817C4FD9FA6CFF787EEA" bold="true" box="[129,216,1484,1508]" pageId="10" pageNumber="11">Figure 7.</emphasis>
<taxonomicName id="4C254D2B1702817C4F85FA6DFE347EEA" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[221,404,1484,1508]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
<emphasis id="B951EABA1702817C4F85FA6DFE347EEA" box="[221,404,1484,1508]" italics="true" pageId="10" pageNumber="11">Pantostictus hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C11702817C4EC2FA6CFE7B7EED" box="[410,475,1484,1507]" pageId="10" pageNumber="11" rank="species">sp. nov</taxonomicNameLabel>
.. A, dorsal view. B, ventral view. C, lateral view. D, elytral setation, E, dorsal view. F, metaventrite, abdomen. Abbreviations: ac, antennal club; as, antennal scape; ls, labral setae; pl, prosternal lobe; pmss, postmesocoxal stria.
</paragraph>
</caption>
<caption id="DF5A66201703817D4F29FA7AFA3D7D48" ID-DOI="http://doi.org/10.5281/zenodo.14516132" ID-Zenodo-Dep="14516132" httpUri="https://zenodo.org/record/14516132/files/figure.png" pageId="11" pageNumber="12" startId="11.[113,178,1498,1522]" targetBox="[114,1458,144,1471]" targetPageId="11" targetType="figure">
<paragraph id="8B9A36A81703817D4F29FA7AFA3D7D48" blockId="11.[113,1437,1498,1607]" pageId="11" pageNumber="12">
<emphasis id="B951EABA1703817D4F29FA7AFF687EFD" bold="true" box="[113,200,1498,1523]" pageId="11" pageNumber="12">Figure 8.</emphasis>
<taxonomicName id="4C254D2B1703817D4F95FA7BFE2B7EFD" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[205,395,1499,1523]" class="Insecta" family="Histeridae" genus="Palaeabraeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="glabrus" status="gen. et sp. nov">
<emphasis id="B951EABA1703817D4F95FA7BFE2B7EFD" box="[205,395,1499,1523]" italics="true" pageId="11" pageNumber="12">Palaeabraeus glabrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C11703817D4EC8FA7BFDB37EFC" box="[400,531,1499,1523]" pageId="11" pageNumber="12" rank="species">gen. et sp. nov</taxonomicNameLabel>
.. A, dorsal view. B, ventral view. C, mouthparts lateral view. D, head, prosternal process and mesoventrite ventral view. E, frontal view. F, lateral view. Abbreviations: ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process.
</paragraph>
</caption>
<subSubSection id="C33F65231704817A4FD9FDC1FDBB7EE8" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8B9A36A81704817A4FD9FDC1FDBB7EE8" blockId="12.[128,779,609,1510]" pageId="12" pageNumber="13">
<emphasis id="B951EABA1704817A4FD9FDC1FECA7976" box="[129,362,609,632]" italics="true" pageId="12" pageNumber="13">Taxonomic assignment:</emphasis>
The species can be assigned to the subfamily
<taxonomicName id="4C254D2B1704817A4F9DFD20FE8F7996" authorityName="W. S. MacLeay" authorityYear="1819" box="[197,303,640,664]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
based on the lack of distinctive prosternal lobe (
<figureCitation id="131E2A2D1704817A4FD3FD3FFF5579B9" box="[139,245,671,695]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." figureDoi="http://doi.org/10.5281/zenodo.14516132" httpUri="https://zenodo.org/record/14516132/files/figure.png" pageId="12" pageNumber="13">Fig. 8B, D</figureCitation>
), convex eyes (
<figureCitation id="131E2A2D1704817A4EC1FD3FFE5E79B9" box="[409,510,671,695]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." figureDoi="http://doi.org/10.5281/zenodo.14516132" httpUri="https://zenodo.org/record/14516132/files/figure.png" pageId="12" pageNumber="13">Fig. 8E, F</figureCitation>
) and fused antennal club (
<figureCitation id="131E2A2D1704817A4FD3FD1EFF5379D8" box="[139,243,702,727]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." figureDoi="http://doi.org/10.5281/zenodo.14516132" httpUri="https://zenodo.org/record/14516132/files/figure.png" pageId="12" pageNumber="13">Fig. 8A, B</figureCitation>
).
<taxonomicName id="4C254D2B1704817A4E52FD1EFED479D8" authorityName="W. S. MacLeay" authorityYear="1819" box="[266,372,702,726]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
contain the following tribes:
<taxonomicName id="4C254D2B1704817A4DF1FD1EFCA579D8" authorityName="MacLeay" authorityYear="1819" box="[681,773,702,726]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Abraeini">Abraeini</taxonomicName>
,
<taxonomicName id="4C254D2B1704817A4FD8FD7EFF7379F8" authorityName="Wenzel" authorityYear="1944" box="[128,211,734,758]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Acritini">Acritini</taxonomicName>
,
<taxonomicName id="4C254D2B1704817A4F86FD7EFE2579F8" authorityName="Wenzel" authorityYear="1944" box="[222,389,734,758]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Acritomorphini">Acritomorphini</taxonomicName>
,
<taxonomicName id="4C254D2B1704817A4EC9FD7EFDAA79F8" authorityName="Portevin" authorityYear="1929" box="[401,522,734,758]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Plegaderini">Plegaderini</taxonomicName>
,
<taxonomicName id="4C254D2B1704817A4D4DFD7EFDDA79F8" authorityName="Bickhardt" authorityYear="1914" box="[533,634,734,758]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Teretriini">Teretriini</taxonomicName>
,
<taxonomicName id="4C254D2B1704817A4DDDFD7EFCA579F8" authorityName="Marseul" authorityYear="1857" box="[645,773,734,758]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Trypanaeini">Trypanaeini</taxonomicName>
,
<taxonomicName id="4C254D2B1704817A4FD8FD5DFF57781B" authorityName="Bickhardt" authorityYear="1913" box="[128,247,765,789]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Trypeticini">Trypeticini</taxonomicName>
, and the extinct
<taxonomicName id="4C254D2B1704817A4EA2FD5DFD22781B" box="[506,642,765,789]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Pantostictini">Pantostictini</taxonomicName>
.
<taxonomicName id="4C254D2B1704817A4DF9FD5DFCA6781B" authorityName="Bickhardt" authorityYear="1914" box="[673,774,765,789]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Teretriini">Teretriini</taxonomicName>
,
<taxonomicName id="4C254D2B1704817A4FD8FCBCFEA1783A" authorityName="Marseul" authorityYear="1857" box="[128,257,796,820]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Trypanaeini">Trypanaeini</taxonomicName>
, and
<taxonomicName id="4C254D2B1704817A4E63FCBCFE10783A" authorityName="Bickhardt" authorityYear="1913" box="[315,432,796,820]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Trypeticini">Trypeticini</taxonomicName>
can be ruled out as they all represent groups with elongate cylindrical body shapes (
<bibRefCitation id="EFB44B591704817A4DE5FC9CFF15787D" author="Wenzel RL" pageId="12" pageNumber="13" pagination="51 - 151" refId="ref19054" refString="Wenzel RL. On the classification of the histerid beetles. Fieldiana Zoology 1944; 28: 51 - 151. hups: // doi. org / 10.1093 / aesa / 37.3.375" type="journal article" year="1944">Wenzel 1944</bibRefCitation>
).
<taxonomicName id="4C254D2B1704817A4F8BFCFBFED8787D" authorityName="Wenzel" authorityYear="1944" box="[211,376,859,883]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Acritomorphini">Acritomorphini</taxonomicName>
have an elongate body shape, long prosternum, anteriorly positioned antennal cavities, and annulate antennal club (
<bibRefCitation id="EFB44B591704817A4E3FFC3AFE4F78BC" author="Wenzel RL" box="[359,495,922,946]" pageId="12" pageNumber="13" pagination="51 - 151" refId="ref19054" refString="Wenzel RL. On the classification of the histerid beetles. Fieldiana Zoology 1944; 28: 51 - 151. hups: // doi. org / 10.1093 / aesa / 37.3.375" type="journal article" year="1944">Wenzel 1944</bibRefCitation>
,
<bibRefCitation id="EFB44B591704817A4EA4FC3AFDDB78BC" author="Mazur" box="[508,635,922,946]" pageId="12" pageNumber="13" pagination="27 - 32" refId="ref18506" refString="Mazur º. New species of histerid beetles (Coleoptera, Histeridae). Annales Zoologici 1997; 38: 27 - 32. hup: // rcin. org. pl" type="journal article" year="1997">Mazur 1997</bibRefCitation>
).
<taxonomicName id="4C254D2B1704817A4DCBFC3AFCA978BC" authorityName="Portevin" authorityYear="1929" box="[659,777,922,946]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Plegaderini">Plegaderini</taxonomicName>
have a flattened body shape and pronotum with a pair of longitudinal grooves laterally, often with a transverse pronotal stria connecting the grooves (
<bibRefCitation id="EFB44B591704817A4ED0FC58FDB17F1E" author="Wenzel RL" box="[392,529,1016,1040]" pageId="12" pageNumber="13" pagination="51 - 151" refId="ref19054" refString="Wenzel RL. On the classification of the histerid beetles. Fieldiana Zoology 1944; 28: 51 - 151. hups: // doi. org / 10.1093 / aesa / 37.3.375" type="journal article" year="1944">Wenzel 1944</bibRefCitation>
).
<taxonomicName id="4C254D2B1704817A4D71FC58FD0E7F1E" box="[553,686,1016,1040]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Pantostictini">Pantostictini</taxonomicName>
likewise have a slightly flattened and rugosely punctuate body surface (
<bibRefCitation id="EFB44B591704817A4FD3FB96FE2E7F41" author="Poinar G &amp; Brown AE" box="[139,398,1078,1103]" pageId="12" pageNumber="13" pagination="38 - 46" refId="ref18750" refString="Poinar G, Brown AE. Pantostictus burmanicus, a new genus and species of Cretaceous beetles (Coleoptera: Hydrophiloidea: Histeridae) in Burmese amber. Proceedings of the Entomological Society of Washington 2009; 111: 38 - 46. hups: // doi. org / 10.4289 / 0013 - 8797 - 111.1.38" type="journal article" year="2009">Poinar and Brown 2009</bibRefCitation>
).
<taxonomicName id="4C254D2B1704817A4EF0FB96FE587F40" authorityName="Wenzel" authorityYear="1944" box="[424,504,1078,1102]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Acritini">Acritini</taxonomicName>
have their tarsal formula 5-5-4 (
<bibRefCitation id="EFB44B591704817A4F94FBF6FDE07F60" author="Kryzhanovskij OL &amp; Reichardt AN" box="[204,576,1110,1134]" pageId="12" pageNumber="13" refId="ref18111" refString="Kryzhanovskij OL, Reichardt AN. Zhuki Nadsemejstva Histeroidea (semejstva Sphaeritidae, Histeridae, Synteliidae) [in Russian]. Nauka, ed., Fauna ºººR, 1976." type="book" year="1976">Kryzhanovskij and Reichardt 1976</bibRefCitation>
). Therefore, based on the round and convex body shape, fused antennal clubs and the 5-5-5 tarsal formula (
<figureCitation id="131E2A2D1704817A4ED0FB34FE4B7FA2" box="[392,491,1172,1197]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." figureDoi="http://doi.org/10.5281/zenodo.14516132" httpUri="https://zenodo.org/record/14516132/files/figure.png" pageId="12" pageNumber="13">Fig. 8B, F</figureCitation>
) the species may be placed in the
<taxonomicName id="4C254D2B1704817A4F9AFB14FEBC7FC2" authorityName="MacLeay" authorityYear="1819" box="[194,284,1204,1228]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Abraeini">Abraeini</taxonomicName>
tribe. Other characters supporting the position of the species within
<taxonomicName id="4C254D2B1704817A4E05FB73FE177FE5" authorityName="MacLeay" authorityYear="1819" box="[349,439,1235,1259]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Abraeini">Abraeini</taxonomicName>
are the presence and position of antennal cavities (
<figureCitation id="131E2A2D1704817A4E1BFB52FE2D7E04" box="[323,397,1266,1290]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." figureDoi="http://doi.org/10.5281/zenodo.14516132" httpUri="https://zenodo.org/record/14516132/files/figure.png" pageId="12" pageNumber="13">Fig. 8F</figureCitation>
), emarginate basal pronotal margin receiving anterior projection of mesoventrite (
<figureCitation id="131E2A2D1704817A4DDCFAB2FD587E27" box="[644,760,1298,1322]" captionStart="Figure 8" captionStartId="11.[113,178,1498,1522]" captionTargetBox="[114,1458,144,1471]" captionTargetId="figure-177@11.[114,1458,144,1471]" captionTargetPageId="11" captionText="Figure 8. Palaeabraeus glabrus gen.et sp. nov.. A, dorsal view.B, ventral view. C, mouthparts lateral view.D, head, prosternal process and mesoventrite ventral view.E, frontal view.F, lateral view.Abbreviations:ac, antennal cavity; apm, anterior margin of prosternal process; ees, elytral epipleural stria; ga, galea; lp, labial palpus; lps, lateral stria on prosternal process; ls, labral setae; mms, marginal mesoventral stria; mp, maxillary palpus; pf, prosternal fissures for receiving antennal funicle; pms, postmesocoxal stria; ppm, posterior margin of prosternal process." figureDoi="http://doi.org/10.5281/zenodo.14516132" httpUri="https://zenodo.org/record/14516132/files/figure.png" pageId="12" pageNumber="13">Fig. 8B, D</figureCitation>
). Close relation to
<taxonomicName id="4C254D2B1704817A4E6EFA91FE307E47" authorityName="MacLeay" authorityYear="1819" box="[310,400,1329,1353]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Abraeini">Abraeini</taxonomicName>
was also indicated by all performed phylogenetic analyses. The main differences setting this species apart from recent
<taxonomicName id="4C254D2B1704817A4ECEFAD0FE507E86" authorityName="MacLeay" authorityYear="1819" box="[406,496,1392,1416]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Abraeini">Abraeini</taxonomicName>
genera (
<taxonomicName id="4C254D2B1704817A4D3AFAD0FCA57E86" authority="Leach" authorityName="Leach" authorityYear="1817" box="[610,773,1392,1416]" class="Insecta" family="Histeridae" genus="Abraeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA1704817A4D3AFAD0FD137E86" box="[610,691,1392,1416]" italics="true" pageId="12" pageNumber="13">Abraeus</emphasis>
Leach
</taxonomicName>
,
<taxonomicName id="4C254D2B1704817A4FD8FA2FFEEE7EA8" authority="Moro" authorityName="Moro" authorityYear="1957" box="[128,334,1423,1447]" class="Insecta" family="Histeridae" genus="Spelaeabraeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA1704817A4FD8FA2FFEAC7EA9" box="[128,268,1423,1447]" italics="true" pageId="12" pageNumber="13">Spelaeabraeus</emphasis>
Moro
</taxonomicName>
, and
<taxonomicName id="4C254D2B1704817A4EDAFA2FFDCD7EA9" authority="Portevin" authorityName="Portevin" authorityYear="1929" box="[386,621,1423,1447]" class="Insecta" family="Histeridae" genus="Chaetabraeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA1704817A4EDAFA2FFDAA7EA9" box="[386,522,1423,1447]" italics="true" pageId="12" pageNumber="13">Chaetabraeus</emphasis>
Portevin
</taxonomicName>
) are: oval body shape, relatively long prosternal process, absence of punctuation, and only slightly dilated protibia.
</paragraph>
</subSubSection>
<subSubSection id="C33F65231704817A4FD8F9A8FE787C56" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8B9A36A81704817A4FD8F9A8FE787C56" blockId="12.[128,778,1543,1880]" pageId="12" pageNumber="13">
<emphasis id="B951EABA1704817A4FD8F9A8FF467D11" box="[128,230,1544,1567]" italics="true" pageId="12" pageNumber="13">Diagnosis:</emphasis>
Absence of dorsal striae and punctuation. Marginal mesoventral stria complete, broadly rounded; mesometaventral suture well developed; metaventrite slightly impressed along midline; postmesocoxal stria complete, strongly recurved behind coxa to mesepimeron; lateral metaventral stria not evident. Only
<taxonomicName id="4C254D2B1704817A4E59F904FE7E7DB5" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[257,478,1700,1724]" class="Insecta" family="Histeridae" genus="Pseudacritus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="extinctus">
<emphasis id="B951EABA1704817A4E59F904FE7E7DB5" box="[257,478,1700,1724]" italics="true" pageId="12" pageNumber="13">Pseudacritus extinctus</emphasis>
</taxonomicName>
, described in this study, is similar in general body shape and some characters to this species.
<taxonomicName id="4C254D2B1704817A4FE9F942FEDD7DF4" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[177,381,1762,1786]" class="Insecta" family="Histeridae" genus="Palaeabraeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="glabrus">
<emphasis id="B951EABA1704817A4FE9F942FEDD7DF4" box="[177,381,1762,1786]" italics="true" pageId="12" pageNumber="13">Palaeabraeus glabrus</emphasis>
</taxonomicName>
can be distinguished by the position of antennal cavities (mediolaterally as opposed to
<taxonomicName id="4C254D2B1704817A4D30F8A2FD757C17" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[616,725,1794,1817]" class="Insecta" family="Histeridae" genus="Pseudacritus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="extinctus">
<emphasis id="B951EABA1704817A4D30F8A2FD757C17" box="[616,725,1794,1817]" italics="true" pageId="12" pageNumber="13">P. extinctus</emphasis>
</taxonomicName>
with anteriorly positioned cavities), absence of antescutellar stria and complete absence of punctuation.
</paragraph>
</subSubSection>
<subSubSection id="C33F65231704817A4FD8F8DAFF707CBF" pageId="12" pageNumber="13" type="etymology">
<paragraph id="8B9A36A81704817A4FD8F8DAFF707CBF" blockId="12.[128,777,1914,1969]" pageId="12" pageNumber="13">
<emphasis id="B951EABA1704817A4FD8F8DAFF507C9C" box="[128,240,1914,1938]" italics="true" pageId="12" pageNumber="13">Etymology:</emphasis>
Latin glabrus refers to the smooth cuticle of the species.
</paragraph>
</subSubSection>
</treatment>
</document>