treatments-xml/data/03/87/BB/0387BB49FFB23D13FD7ACCFCFE12FCB1.xml
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<mods:title id="6F7B016C57A454081C415EF966AE5A60">Upper Miocene mammals from Strumyani, South-Western Bulgaria</mods:title>
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<mods:namePart id="4FE9C6256EA8A43B74D3F1CC1C71D3E6">Geraads, Denis</mods:namePart>
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<mods:namePart id="B9A8C8D9E5430EAC9902EFAD0C84436F">Spassov, Nikolaï</mods:namePart>
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<mods:namePart id="1470D75FC7472948BC1CD7AD3C13E81B">Hristova, Latinka</mods:namePart>
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<mods:namePart id="D897A8E9C52B7AB4A9DFB268E2FC880D">Markov, Georgi N.</mods:namePart>
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<mods:namePart id="C891F2FE8B178593DD8A4421E97AADCF">Tzankov, Tzanko</mods:namePart>
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<mods:title id="6F68DF6564F29CC72DFCAFC982CF3EB6">Geodiversitas</mods:title>
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<mods:date id="6D73B449488C7EBDE47A8D98F4BD003A">2011</mods:date>
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<mods:number id="6435631DAF6700B28ECE0B1B64EFEA41">2011-09-30</mods:number>
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<paragraph id="8B910A5FFFB23D12FD7ACCFCFBF2FC10" blockId="10.[752,1134,912,940]" box="[752,1134,912,940]" pageId="10" pageNumber="461">
<heading id="D0D9BD33FFB23D12FD7ACCFCFBF2FC10" box="[752,1134,912,940]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="461" reason="2">
<taxonomicName id="4C2E71DCFFB23D12FD7ACCFCFBF2FC10" authority="(Wehrli, 1941)" baseAuthorityName="Wehrli" baseAuthorityYear="1941" box="[752,1134,912,940]" class="Mammalia" family="Equidae" genus="Hipparion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="10" pageNumber="461" phylum="Chordata" rank="species" species="dietrichi">
<emphasis id="B95AD64DFFB23D12FD7ACCFCFC5AFC17" bold="true" box="[752,966,912,939]" italics="true" pageId="10" pageNumber="461">Hipparion dietrichi</emphasis>
(Wehrli, 1941)
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</heading>
</paragraph>
<paragraph id="8B910A5FFFB23D13FD2FCCBDFEB9FE8E" blockId="10.[674,1212,976,1700]" lastBlockId="11.[111,648,216,781]" lastPageId="11" lastPageNumber="462" pageId="10" pageNumber="461">
There are two relatively well preserved skulls of this species. FM-2029 is a subadult individual, with unerupted I3s, C and M3 just erupted (
<figureCitation id="131516DAFFB23D12FBD2CB7DFB32FB97" box="[1112,1198,1040,1067]" captionStart="FIG" captionStartId="12.[113,124,1122,1139]" captionTargetBox="[113,1209,214,1084]" captionTargetId="figure-262@12.[113,1209,214,1084]" captionTargetPageId="12" captionText="FIG. 6. — Fossil mammals from the Miocene of Strumyani (SW Bulgaria): A, Hippotherium cf. brachypus (Hensel, 1862), lateral view of skull FM-2326; B, Cremohipparion mediterraneum (Roth &amp; Wagner, 1855), lateral view of skull FM-2028;C, Hipparion dietrichi (Wehrli, 1941), lateral view of skull FM-2029. Scale bar: 20 cm." figureDoi="http://doi.org/10.5281/zenodo.5381390" httpUri="https://zenodo.org/record/5381390/files/figure.png" pageId="10" pageNumber="461">Fig. 6C</figureCitation>
). FM-2030 belongs to an adult individual, but is not well-preserved, so that some of the characters described are those of FM-2029. The muzzle is short and broad. The nasal slit ends above the anterior end of P2; the crista facialis reaches the middle of P4; the choanae reach the limit M1-M2; the orbit is above the posterior half of M3. The anteroposteriorly oriented preorbital fossa is small and shallow, without posterior pocket and with a weak anterior rim. It extends from above the anterior part of P3 to above the posterior border of M1. The infraorbital foramen is large, located below the anterior border of the fossa. Tooth row length is about
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for FM-2029 and
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for FM-2030. There is no DP1. The anterostyle of P2 is elongated. The enamel is moderately plicated, with rounded folds whose number ranges from 14 on the premolars to 21 on the molars. The
<emphasis id="B95AD64DFFB23D12FBDDC927FD5EF93F" italics="true" pageId="10" pageNumber="461">pli caballin</emphasis>
is single. The protocon is oval, and lenticular on the less worn teeth. The hypocone is elongated, angular on P4; the hypoconal sinus is comparatively deep in the subadult skull and medium to absent in the adult one.
</paragraph>
</subSubSection>
<subSubSection id="C33459D4FFB33D13FF02CE5AFE12FCB1" pageId="11" pageNumber="462" type="description">
<paragraph id="8B910A5FFFB33D13FF02CE5AFE12FCB1" blockId="11.[111,648,216,781]" pageId="11" pageNumber="462">
The short and broad muzzle, antero-posteriorly oriented oval preorbital fossa without well defined peripheral rims and without posterior pocket, relate this skull to
<taxonomicName id="4C2E71DCFFB33D13FF65CEFAFEF9FE0C" baseAuthorityName="Wehrli" baseAuthorityYear="1941" box="[239,357,406,432]" class="Mammalia" family="Equidae" genus="Hipparion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="11" pageNumber="462" phylum="Chordata" rank="species" species="dietrichi">
<emphasis id="B95AD64DFFB33D13FF65CEFAFEF9FE0C" box="[239,357,406,432]" italics="true" pageId="11" pageNumber="462">H. dietrichi</emphasis>
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, and the tooth features support this conclusion.
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<emphasis id="B95AD64DFFB33D13FEDFCEDBFD83FE73" box="[341,543,437,463]" italics="true" pageId="11" pageNumber="462">Hipparion dietrichi</emphasis>
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is known from many localities in
<collectingCountry id="F3394ACFFFB33D13FEECCEB8FE2EFE4C" box="[358,434,469,496]" name="Greece" pageId="11" pageNumber="462">Greece</collectingCountry>
,
<collectingCountry id="F3394ACFFFB33D13FE30CEB8FD98FE4C" box="[442,516,469,496]" name="Turkey" pageId="11" pageNumber="462">Turkey</collectingCountry>
and
<collectingCountry id="F3394ACFFFB33D13FDBCCEB8FEB8FDAC" name="Macedonia" pageId="11" pageNumber="462">Republic of Macedonia</collectingCountry>
(
<bibRefCitation id="EFBF77AEFFB33D13FEBACE98FE57FDAC" author="KOUFOS G. D." box="[304,459,501,528]" pageId="11" pageNumber="462" pagination="293 - 312" refId="ref20660" refString="KOUFOS G. D. 1987 c. - Study of the Turolian hipparions of the Lower Axios Valley (Macedonia, Greece). 1. Locality &quot; Ravin des Zouaves- 5 &quot; (RZO). Geobios 20: 293 - 312." type="journal article" year="1987">Koufos 1987c</bibRefCitation>
,
<bibRefCitation id="EFBF77AEFFB33D13FE5FCE98FD8CFDAC" author="KOUFOS G. D." box="[469,528,501,528]" pageId="11" pageNumber="462" pagination="15 - 39" refId="ref20705" refString="KOUFOS G. D. 1988. - Study of the Turolian Hipparions of the Lower Axios Valley (Macedonia, Greece). 3 - Localities of Vathylakkos. Paleontologia i Evolucio 22: 15 - 39." type="journal article" year="1988">1988</bibRefCitation>
;
<bibRefCitation id="EFBF77AEFFB33D13FD93CE98FE8FFD8C" author="FORSTEN A. &amp; GAREVSKI R." pageId="11" pageNumber="462" pagination="1 - 204" refId="ref19079" refString="FORSTEN A. &amp; GAREVSKI R. 1989. - Hipparions (Mammalia, Perissodactyla) from Macedonia (Yugoslavia). Geologica macedonica 3 (2): 1 - 204." type="journal article" year="1989">Forstén &amp; Garevski 1989</bibRefCitation>
;
<bibRefCitation id="EFBF77AEFFB33D13FEABCD78FDABFD8C" author="VLACHOU T. &amp; KOUFOS G. D." box="[289,567,533,560]" pageId="11" pageNumber="462" pagination="215 - 263" refId="ref22546" refString="VLACHOU T. &amp; KOUFOS G. D. 2002. - The hipparions (Mammalia, Perissodactyla) from theTurolian locality &quot; Nikiti- 2 &quot; (NIK), Macedonia, N. Greece. Annales de Paleontologie 88: 215 - 263." type="journal article" year="2002">Vlachou &amp; Koufos 2002</bibRefCitation>
, 2006, 2009;
<bibRefCitation id="EFBF77AEFFB33D13FF33CD59FE54FDF3" author="KOUFOS G. D. &amp; VLACHOU T. D." box="[185,456,564,591]" pageId="11" pageNumber="462" pagination="633 - 705" refId="ref20797" refString="KOUFOS G. D. &amp; VLACHOU T. D. 2005. - Equidae (Mammalia, Perissodactyla) from the late Miocene of AkkasdaGI, Turkey. Geodiversitas 27 (4): 633 - 705." type="journal article" year="2005">Koufos &amp; Vlachou 2005</bibRefCitation>
). From its muzzle length the skull from Str-2 is close to
<taxonomicName id="4C2E71DCFFB33D13FD86CD38FD1BFDD2" baseAuthorityName="Wehrli" baseAuthorityYear="1941" box="[524,647,596,622]" class="Mammalia" family="Equidae" genus="Hipparion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="11" pageNumber="462" phylum="Chordata" rank="species" species="dietrichi">
<emphasis id="B95AD64DFFB33D13FD86CD38FD1BFDD2" box="[524,647,596,622]" italics="true" pageId="11" pageNumber="462">H. dietrichi</emphasis>
</taxonomicName>
from AkkasdaĞI, but in cheek tooth length it is intermediate between the Perivolaki and Vathylakkos samples on the one side and the AkkasdaĞI form on the other (
<bibRefCitation id="EFBF77AEFFB33D13FF57CDBEFEF1FD52" author="KOUFOS G. D." box="[221,365,723,750]" pageId="11" pageNumber="462" pagination="15 - 39" refId="ref20705" refString="KOUFOS G. D. 1988. - Study of the Turolian Hipparions of the Lower Axios Valley (Macedonia, Greece). 3 - Localities of Vathylakkos. Paleontologia i Evolucio 22: 15 - 39." type="journal article" year="1988">Koufos 1988</bibRefCitation>
;
<bibRefCitation id="EFBF77AEFFB33D13FEFCCDBEFD1CFD52" author="KOUFOS G. D. &amp; VLACHOU T. D." box="[374,640,723,750]" pageId="11" pageNumber="462" pagination="633 - 705" refId="ref20797" refString="KOUFOS G. D. &amp; VLACHOU T. D. 2005. - Equidae (Mammalia, Perissodactyla) from the late Miocene of AkkasdaGI, Turkey. Geodiversitas 27 (4): 633 - 705." type="journal article" year="2005">Koufos &amp; Vlachou 2005</bibRefCitation>
;
<bibRefCitation id="EFBF77AEFFB33D13FFFBCD9FFE1EFCB1" author="VLACHOU T. D. &amp; KOUFOS G. D." box="[113,386,754,781]" pageId="11" pageNumber="462" pagination="81 - 119" refId="ref22591" refString="VLACHOU T. D. &amp; KOUFOS G. D. 2006. - The late Miocene vertebrate locality of Perivolaki, Thessaly, Greece. 6. Equidae. Palaeontographica 276: 81 - 119." type="journal article" year="2006">Vlachou &amp; Koufos 2006</bibRefCitation>
).
</paragraph>
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