301 lines
41 KiB
XML
301 lines
41 KiB
XML
<document id="9693D5A50C3C6B8A4657D44D045075EB" ID-DOI="10.5281/zenodo.14660773" ID-ISSN="1536-9307" ID-Zenodo-Dep="14660773" ID-ZooBank="7A1007F6-0FDB-40D3-A2B0-AF3D40CFCF5C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736980494816" checkinUser="felipe" docAuthor="Euscorpius & No." docDate="2024" docId="03879D78DA228521FED9FF7E471AF82D" docLanguage="en" docName="Euscorpius.401.1-31.pdf" docOrigin="Euscorpius 401" docStyle="DocumentStyle:5E96F6379CE20B0E8E4F9E7DF8E1C0DF.2:Euscorpius.2020-.journal_article" docStyleId="5E96F6379CE20B0E8E4F9E7DF8E1C0DF" docStyleName="Euscorpius.2020-.journal_article" docStyleVersion="2" docTitle="Neobuthus opatovae Euscorpius & No., 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="11" masterDocId="FFBEE500DA29852DFFC3FF94435DFFCD" masterDocTitle="Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XXXIV. Four new species of Neobuthus from Somaliland (Buthidae)" masterLastPageNumber="31" masterPageNumber="1" pageNumber="10" updateTime="1736981056599" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="9A3D328D6E1ED642CFD41F48F3B641B8">Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XXXIV. Four new species of Neobuthus from Somaliland (Buthidae)</mods:title>
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<mods:namePart id="2D21AB1C4017CB99C0D4C72448F08CF1">Euscorpius</mods:namePart>
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<mods:title id="7FA94C8D1E43060CAB8E836A6534B48A">Euscorpius</mods:title>
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<mods:date id="A6C4C617E05A34A16C9A20CBE9D86807">2024</mods:date>
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<taxonomicName id="4C2E57EDDA228526FED9FF7E414CFEC8" authorityName="Euscorpius & No." authorityYear="2024" box="[282,529,234,261]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="opatovae" status="sp. nov.">Neobuthus opatovae</taxonomicName>
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.
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<subSubSection id="C3347FE5DA228526FEE9FE9841DCFEA9" pageId="11" pageNumber="10" type="description">
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<paragraph id="8B912C6EDA228526FEE9FE984059FE89" blockId="11.[113,772,234,356]" pageId="11" pageNumber="10">
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(
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<figureCitation id="131530EBDA228526FEF1FE9842E9FEE9" box="[306,436,268,292]" captionStart-0="Figures 46–47" captionStart-1="Figures 48–56" captionStart-2="Figures 57–62" captionStart-3="Figures 63–75" captionStartId-0="12.[116,193,1697,1719]" captionStartId-1="13.[113,190,1983,2005]" captionStartId-2="14.[116,193,1981,2003]" captionStartId-3="15.[112,189,1962,1984]" captionTargetBox-0="[144,1436,304,1672]" captionTargetBox-1="[113,1474,234,1963]" captionTargetBox-2="[113,1474,220,1963]" captionTargetBox-3="[113,1474,247,1952]" captionTargetId-0="figure-209@12.[113,1474,232,1701]" captionTargetId-1="figure-72@13.[113,1474,234,2003]" captionTargetId-2="figure-55@14.[113,1474,193,2000]" captionTargetId-3="figure-92@15.[113,1474,227,1993]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="15" captionText-0="Figures 46–47: Neobuthus opatovae sp. n., male holotYpe, dorsal (46) and ventral (47) views. Scale bar: 10 mm." captionText-1="Figures 48–56: Neobuthus opatovae sp. n., male holotYpe. Figures 48–49: Carapace and tergites I–IV (48) and coxosternal area and sternites (49). Figures 50–51: Carapace and tergites (50) and coxosternal area and sternites (51) under UV fluorescence. Figures 52–55: Left legs I–IV, retrolateral aspect. Figure. 56. Telson lateral view." captionText-2="Figures 57–62. Neobuthus opatovae sp. n., male holotYpe, metasoma and telson lateral (57, 60), dorsal (58, 61), and ventral (59, 62) views, under UV fluorescence (57–59) and under white light (60–62). Scale bar: 5 mm." captionText-3="Figures 63–75: Neobuthus opatovae sp. n., pedipalp, male holotYpe. Figures 63–73: Chela dorsal (63), external (64), and ventral (65) views, patella dorsal (66), external (67) and ventral (68) views, femur and trochanter internal (69), dorsal (70), and ventral (71) views, movable (72) and fixed (73) finger dentate margin. The trichobothrial pattern is indicated in Figures 63–67, 69–70 (white circles). Figures 74–75: Right pedipalp in dorsal (74) and ventral (75) views under UV fluorescence." figureDoi-0="http://doi.org/10.5281/zenodo.14660788" figureDoi-1="http://doi.org/10.5281/zenodo.14660790" figureDoi-2="http://doi.org/10.5281/zenodo.14660792" figureDoi-3="http://doi.org/10.5281/zenodo.14660794" httpUri-0="https://zenodo.org/record/14660788/files/figure.png" httpUri-1="https://zenodo.org/record/14660790/files/figure.png" httpUri-2="https://zenodo.org/record/14660792/files/figure.png" httpUri-3="https://zenodo.org/record/14660794/files/figure.png" pageId="11" pageNumber="10">Figs. 46–75</figureCitation>
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,
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<figureCitation id="131530EBDA228526FE7DFE9842B5FEE9" box="[446,488,268,292]" captionStart="Figures 160–161" captionStartId="29.[114,191,2011,2033]" captionTargetBox="[113,1474,226,1992]" captionTargetId="figure-9@29.[113,1474,226,2040]" captionTargetPageId="29" captionText="Figures 160–161. Map showing confirmed distribution of Neobuthus spp. (160) with detail for Somaliland (161)." figureDoi="http://doi.org/10.5281/zenodo.14660820" httpUri="https://zenodo.org/record/14660820/files/figure.png" pageId="11" pageNumber="10">160</figureCitation>
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,
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<tableCitation id="C6AC19D5DA228526FE30FE98411DFEE8" box="[499,576,268,293]" captionStart="Table 1" captionStartId="7.[113,169,974,995]" captionText="Table 1. Comparative measurements of adults of Neobuthus opatovae sp. n. and N. dhobo sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="11" pageNumber="10">Table 1</tableCitation>
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)
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:
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.
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<materialsCitation id="3B462633DA228526FFB2FE194211FE68" box="[113,332,397,421]" collectionCode="LOCALITY, AND" pageId="11" pageNumber="10" specimenCount="1" typeStatus="holotype">
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<materialsCitation id="3B462633DA228526FE96FE1941B2FE09" collectionCode="REPOSITORY" county="HabaaswaYn" latitude="44.052673" location="Somaliland" municipality="Maroodi Jeex Region" pageId="11" pageNumber="10" specimenCount="1" typeStatus="holotype">
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,
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<collectingMunicipality id="6BF5B614DA228526FD64FE184258FE09" pageId="11" pageNumber="10">Maroodi Jeex Region</collectingMunicipality>
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,
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<collectingCounty id="62F054E2DA228526FEDEFE3842F5FE09" box="[285,424,428,452]" pageId="11" pageNumber="10">HabaaswaYn</collectingCounty>
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, 9.3841900°
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E,
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E,
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(
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<typeStatus id="549592CCDA228526FFBBFDD8438AFDA9" box="[120,215,588,612]" pageId="11" pageNumber="10" type="holotype">holotYpe</typeStatus>
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DNA No. 947 and
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<typeStatus id="549592CCDA228526FE7EFDD94144FDA9" box="[445,537,589,612]" pageId="11" pageNumber="10" type="paratype">paratYpe</typeStatus>
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DNA No. 948), leg.
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<collectorName id="26DB49B8DA228526FFB2FDF9425DFD49" box="[113,256,620,644]" pageId="11" pageNumber="10">Hassan Elmi.</collectorName>
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<paragraph id="8B912C6EDA228526FFB2FD394247FCA8" blockId="11.[113,771,684,869]" pageId="11" pageNumber="10">ETYMOLOGY. It is a pleasure to name this species after Vera Opatova, the Czech arachnologist. Her phYlogenetic analYses highlY contribute to the understanding of species diversitY and phYlogenY across various arachnid groups, including scorpions. The name of this species is derived from her last name, Opatova.</paragraph>
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</subSubSection>
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<subSubSection id="C3347FE5DA228526FFB2FC194203F9C9" pageId="11" pageNumber="10" type="diagnosis">
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<paragraph id="8B912C6EDA228526FFB2FC194203F9C9" blockId="11.[113,772,908,1541]" pageId="11" pageNumber="10">
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DIAGNOSIS (
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). Total length of male
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||
, female unknown; carapace with area between anterior median carinae fuscous; tergites with 3 dark stripes, median stripe flanked on either side bY broad longitudinal Yellow bands that maY be broken bY fuscositY extending across anterior tergites; pedipalp relativelY slender, male with femur L/
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, patella L/
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, chela L/
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; chela movable finger with 6 subrows of primarY denticles, 5 external accessorY denticles flanking proximal end of each subrow; dorsoexternal and ventroexternal carinae on pedipalp patella in male present; posterior margins of tergites with 2–3 pairs of macrosetae; pedipalps, legs, metasoma and telson with moderatelY short in male, not verY spiniform, and long, fine setae in females; male with sternites III–VI granulated locallY, sternite VII denselY granulated with 4 granulated carinae; sternite III with 10 macrosetae; metasoma I–III with median lateral carinae present; lateral surface of metasoma V in males granulated, with granules separated; soles of telotarsi with relativelY sparse setation, leg III of adults with 18–21 ventral macrosetae on telotarsus; pectine teeth:
|
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<quantity id="4CD6818BDA228526FF70FA79424FF9C9" box="[179,274,1516,1541]" metricMagnitude="-1" metricUnit="m" metricValue="4.953" metricValueMax="5.08" metricValueMin="4.826" pageId="11" pageNumber="10" unit="in" value="19.5" valueMax="20.0" valueMin="19.0">19–20 in</quantity>
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males.
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</paragraph>
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|
||
DESCRIPTION. Total length of adult males
|
||
<quantity id="4CD6818BDA228526FD96F9B941F4F989" box="[597,681,1581,1605]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="10" unit="mm" value="22.0">22 mm</quantity>
|
||
, female unknown; measurements of carapace, telson, segments of metasoma and pedipalps given in
|
||
<tableCitation id="C6AC19D5DA228526FDC2F9F84104F948" box="[513,601,1644,1669]" captionStart="Table 1" captionStartId="7.[113,169,974,995]" captionText="Table 1. Comparative measurements of adults of Neobuthus opatovae sp. n. and N. dhobo sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="11" pageNumber="10">Table 1</tableCitation>
|
||
; positions and distribution of trichobothria of pedipalps shown in
|
||
<figureCitation id="131530EBDA228526FD5DF91843BFF909" captionStart="Figures 63–75" captionStartId="15.[112,189,1962,1984]" captionTargetBox="[113,1474,247,1952]" captionTargetId="figure-92@15.[113,1474,227,1993]" captionTargetPageId="15" captionText="Figures 63–75: Neobuthus opatovae sp. n., pedipalp, male holotYpe. Figures 63–73: Chela dorsal (63), external (64), and ventral (65) views, patella dorsal (66), external (67) and ventral (68) views, femur and trochanter internal (69), dorsal (70), and ventral (71) views, movable (72) and fixed (73) finger dentate margin. The trichobothrial pattern is indicated in Figures 63–67, 69–70 (white circles). Figures 74–75: Right pedipalp in dorsal (74) and ventral (75) views under UV fluorescence." figureDoi="http://doi.org/10.5281/zenodo.14660794" httpUri="https://zenodo.org/record/14660794/files/figure.png" pageId="11" pageNumber="10">Figs. 63– 67, 69–70</figureCitation>
|
||
; trichobothrium
|
||
<emphasis id="B95AF07CDA228526FE5BF93842F8F909" box="[408,421,1708,1732]" italics="true" pageId="11" pageNumber="10">d</emphasis>
|
||
2 present on pedipalp femur and patella; base color Yellow to light orange with variable fuscous pigmentation and extensive patterns of dark maculation on mesosoma, metasoma, partiallY on pedipalps and legs; chelicerae Yellow with dark reticulation on anterior manus, dentition reddish.
|
||
<emphasis id="B95AF07CDA228526FEF0F8D842A2F8A9" box="[307,511,1868,1892]" italics="true" pageId="11" pageNumber="10">Sexual dimorphism</emphasis>
|
||
unknown.
|
||
</paragraph>
|
||
<paragraph id="8B912C6EDA228526FFB2F8F84796FEEE" blockId="11.[113,772,1580,2020]" lastBlockId="11.[816,1475,234,2019]" pageId="11" pageNumber="10">
|
||
<emphasis id="B95AF07CDA228526FFB2F8F8438BF849" bold="true" box="[113,214,1900,1924]" pageId="11" pageNumber="10">Pedipalp</emphasis>
|
||
(
|
||
<figureCitation id="131530EBDA228526FF28F8F8422FF849" box="[235,370,1900,1924]" captionStart="Figures 63–75" captionStartId="15.[112,189,1962,1984]" captionTargetBox="[113,1474,247,1952]" captionTargetId="figure-92@15.[113,1474,227,1993]" captionTargetPageId="15" captionText="Figures 63–75: Neobuthus opatovae sp. n., pedipalp, male holotYpe. Figures 63–73: Chela dorsal (63), external (64), and ventral (65) views, patella dorsal (66), external (67) and ventral (68) views, femur and trochanter internal (69), dorsal (70), and ventral (71) views, movable (72) and fixed (73) finger dentate margin. The trichobothrial pattern is indicated in Figures 63–67, 69–70 (white circles). Figures 74–75: Right pedipalp in dorsal (74) and ventral (75) views under UV fluorescence." figureDoi="http://doi.org/10.5281/zenodo.14660794" httpUri="https://zenodo.org/record/14660794/files/figure.png" pageId="11" pageNumber="10">Figs. 63–75</figureCitation>
|
||
). Pedipalp mostlY sparselY hirsute; finelY granulated in males; femur with five conspicuouslY granulose carinae developed in male; patella with seven granulose carinae, well developed; chela movable finger with 6 subrows of primarY denticles, 5 external accessorY denticles flanking proximal end of each subrow.
|
||
</paragraph>
|
||
<paragraph id="8B912C6EDA228526FCF3FEBE470CFDAF" blockId="11.[816,1475,234,2019]" pageId="11" pageNumber="10">
|
||
<emphasis id="B95AF07CDA228526FCF3FEBE40C3FE8F" bold="true" box="[816,926,298,322]" pageId="11" pageNumber="10">Carapace</emphasis>
|
||
(
|
||
<figureCitation id="131530EBDA228526FC75FEBF4716FE8F" box="[950,1099,298,323]" captionStart="Figures 48–56" captionStartId="13.[113,190,1983,2005]" captionTargetBox="[113,1474,234,1963]" captionTargetId="figure-72@13.[113,1474,234,2003]" captionTargetPageId="13" captionText="Figures 48–56: Neobuthus opatovae sp. n., male holotYpe. Figures 48–49: Carapace and tergites I–IV (48) and coxosternal area and sternites (49). Figures 50–51: Carapace and tergites (50) and coxosternal area and sternites (51) under UV fluorescence. Figures 52–55: Left legs I–IV, retrolateral aspect. Figure. 56. Telson lateral view." figureDoi="http://doi.org/10.5281/zenodo.14660790" httpUri="https://zenodo.org/record/14660790/files/figure.png" pageId="11" pageNumber="10">Figs. 48, 50</figureCitation>
|
||
). StronglY trapezoidal (narrower anteriorlY), wider than long (L/
|
||
<geoCoordinate id="EE1A4AA9DA228526FB58FEDF47AFFEAE" box="[1179,1266,331,355]" degrees="0.90" direction="west" orientation="longitude" pageId="11" pageNumber="10" precision="555" value="-0.9">W 0.90</geoCoordinate>
|
||
); posterior median postocular area flat, anterior median preocular area gentlY sloped downwards towards anterior margin; lateral flanks steeplY sloped; ocular tubercle broad, prominent, located slightlY anterior of carapace; anterior margin straight, finelY microdenticulate, with coarser granules overlapping edge, bearing 6 macrosetae; anterior median carinae present, coarselY granular, other carinae indistinct; dense granulation covering most of carapace.
|
||
</paragraph>
|
||
<paragraph id="8B912C6EDA228526FCF3FDFE476FFCAE" blockId="11.[816,1475,234,2019]" pageId="11" pageNumber="10">
|
||
<emphasis id="B95AF07CDA228526FCF3FDFE40C3FD4F" bold="true" box="[816,926,618,642]" pageId="11" pageNumber="10">Chelicera</emphasis>
|
||
. Fingers with tYpical buthid dentition (
|
||
<bibRefCitation id="EFBF519FDA228526FAA9FDFF4035FD6E" author="VACHON" pageId="11" pageNumber="10" pagination="161 - 166" refId="ref10978" refString="VACHON, M. 1963. De l'utilite, en sYstematique, d'une nomenclature des dents de cheliceres chez les scorpions. Bulletin du Museum National d'Histoire Naturelle, Paris, (2), 35 (2): 161 - 166." type="journal article" year="1963">Vachon, 1963</bibRefCitation>
|
||
; Lowe & KovařÍk, 2016;
|
||
<bibRefCitation id="EFBF519FDA228526FB53FD1F462BFD6E" author="KOVARIK" box="[1168,1398,651,677]" pageId="11" pageNumber="10" pagination="1 - 82" refId="ref10578" refString="KOVARIK, F., G. LOWE, A. I. AWALE, H. SH A. ELMI & A. A. HURE. 2018. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XVII. Revision of Neobuthus, with description of seven new species from Ethiopia, KenYa and Somaliland (Buthidae). Euscorpius, 271: 1 - 82." type="journal article" year="2018">KovařÍk et al., 2018</bibRefCitation>
|
||
); fixed finger with large distal denticle, 1 subdistal denticle and 2 basal denticles fused into bicusp, single denticle on ventral surface at level of bicusp; dorsal margin of movable finger with 5 denticles: 1 large distal denticle, medium-sized subdistal and medial, and 2 small, partiallY fused basal denticles; ventral margin with 2 denticles.
|
||
</paragraph>
|
||
<paragraph id="8B912C6EDA228526FCF3FCFF40A0F9AF" blockId="11.[816,1475,234,2019]" pageId="11" pageNumber="10">
|
||
<emphasis id="B95AF07CDA228526FCF3FCFF40F5FC4E" bold="true" box="[816,936,875,899]" pageId="11" pageNumber="10">Mesosoma</emphasis>
|
||
(
|
||
<figureCitation id="131530EBDA228526FC7FFCFF4718FC4F" box="[956,1093,874,899]" captionStart="Figures 48–56" captionStartId="13.[113,190,1983,2005]" captionTargetBox="[113,1474,234,1963]" captionTargetId="figure-72@13.[113,1474,234,2003]" captionTargetPageId="13" captionText="Figures 48–56: Neobuthus opatovae sp. n., male holotYpe. Figures 48–49: Carapace and tergites I–IV (48) and coxosternal area and sternites (49). Figures 50–51: Carapace and tergites (50) and coxosternal area and sternites (51) under UV fluorescence. Figures 52–55: Left legs I–IV, retrolateral aspect. Figure. 56. Telson lateral view." figureDoi="http://doi.org/10.5281/zenodo.14660790" httpUri="https://zenodo.org/record/14660790/files/figure.png" pageId="11" pageNumber="10">Figs. 48–51</figureCitation>
|
||
). Tergites I–VI bear three carinae of which the lateral pair maY be less conspicuous mainlY on tergites I–IV; tergite VII bears five indicated carinae; tergites denselY granular, with coarser granules on posterior lateral areas; tergite; sternites III–VI granulated locallY, posterior margins of tergites with 2–3 pairs of macrosetae; sternite VII denselY granulated with 4 granulated carinae; sternite III with 10 macrosetae; sternum
|
||
<typeStatus id="549592CCDA228526FB59FBDF4795FBAF" box="[1178,1224,1099,1122]" pageId="11" pageNumber="10">tYpe</typeStatus>
|
||
1, triangular in shape, smooth, with deep posteromedian invagination; genital opercula smooth; genital papillae present; pectines extending to around end of sternite IV in males; pectine teeth
|
||
<quantity id="4CD6818BDA228526FAA1FB3F469FFB0E" box="[1378,1474,1195,1219]" metricMagnitude="-1" metricUnit="m" metricValue="4.953" metricValueMax="5.08" metricValueMin="4.826" pageId="11" pageNumber="10" unit="in" value="19.5" valueMax="20.0" valueMin="19.0">19–20 in</quantity>
|
||
males; combs with 3 marginal lamellae and 6 middle lamellae; marginal lamellae, middle lamellae and fulcra with dense cover of short dark reddish macrosetae; fulcra with 2–4 setae.
|
||
<emphasis id="B95AF07CDA228526FCF3FABF4038FA8E" bold="true" box="[816,869,1323,1347]" pageId="11" pageNumber="10">Legs</emphasis>
|
||
(
|
||
<figureCitation id="131530EBDA228526FCB7FABF40AAFA8F" box="[884,1015,1322,1347]" captionStart="Figures 48–56" captionStartId="13.[113,190,1983,2005]" captionTargetBox="[113,1474,234,1963]" captionTargetId="figure-72@13.[113,1474,234,2003]" captionTargetPageId="13" captionText="Figures 48–56: Neobuthus opatovae sp. n., male holotYpe. Figures 48–49: Carapace and tergites I–IV (48) and coxosternal area and sternites (49). Figures 50–51: Carapace and tergites (50) and coxosternal area and sternites (51) under UV fluorescence. Figures 52–55: Left legs I–IV, retrolateral aspect. Figure. 56. Telson lateral view." figureDoi="http://doi.org/10.5281/zenodo.14660790" httpUri="https://zenodo.org/record/14660790/files/figure.png" pageId="11" pageNumber="10">Figs. 52–55</figureCitation>
|
||
). Coxa, femora, patella and tibia of all legs bearing variable numbers of short to medium length, straight, dark-reddish macrosetae; tarsi with mix of short and longer, dark-reddish macrosetae; basitarsi I–III slightlY compressed with flat retrolateral surfaces, with bristle combs consisting of retrosuperior series of longer macrosetae, plus retroinferior and proinferior series of shorter macrosetae; telotarsi with two rows of short macrosetae on ventral aspect, 18–21 macrosetae on telotarsus III; strong tibial spurs present on leg IV and reduced on leg. III.
|
||
</paragraph>
|
||
<paragraph id="8B912C6EDA228526FCF3F9FF46E7F80E" blockId="11.[816,1475,234,2019]" pageId="11" pageNumber="10">
|
||
<emphasis id="B95AF07CDA228526FCF3F9FF4777F94E" bold="true" box="[816,1066,1643,1667]" pageId="11" pageNumber="10">Metasoma and telson</emphasis>
|
||
(
|
||
<figureCitation id="131530EBDA228526FBF8F9FF479FF94F" box="[1083,1218,1642,1667]" captionStart-0="Figures 48–56" captionStart-1="Figures 57–62" captionStartId-0="13.[113,190,1983,2005]" captionStartId-1="14.[116,193,1981,2003]" captionTargetBox-0="[113,1474,234,1963]" captionTargetBox-1="[113,1474,220,1963]" captionTargetId-0="figure-72@13.[113,1474,234,2003]" captionTargetId-1="figure-55@14.[113,1474,193,2000]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="Figures 48–56: Neobuthus opatovae sp. n., male holotYpe. Figures 48–49: Carapace and tergites I–IV (48) and coxosternal area and sternites (49). Figures 50–51: Carapace and tergites (50) and coxosternal area and sternites (51) under UV fluorescence. Figures 52–55: Left legs I–IV, retrolateral aspect. Figure. 56. Telson lateral view." captionText-1="Figures 57–62. Neobuthus opatovae sp. n., male holotYpe, metasoma and telson lateral (57, 60), dorsal (58, 61), and ventral (59, 62) views, under UV fluorescence (57–59) and under white light (60–62). Scale bar: 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14660790" figureDoi-1="http://doi.org/10.5281/zenodo.14660792" httpUri-0="https://zenodo.org/record/14660790/files/figure.png" httpUri-1="https://zenodo.org/record/14660792/files/figure.png" pageId="11" pageNumber="10">Figs. 56–62</figureCitation>
|
||
). Metasoma and telson sparselY hirsute, macrosetae moderatelY short, not verY spiniform, and long in male; metasomal segments I–III with 10 carinae, IV with 8 carinae, V with 2 carinae; all carinae relativelY well developed; segment V with strong, granulate to dentate-lobate ventrolateral carinae; segments I–IV with dense granulation on all intercarinal surfaces; segment V denselY granular on lateral and ventral surfaces, more coarselY so on ventral surface; telson smooth, ventral surface sparselY, weaklY granular; vesicle slightlY elongated; aculeus stout, shorter than vesicle, tip of aculeus almost verticallY directed.
|
||
</paragraph>
|
||
<paragraph id="8B912C6EDA228526FCF3F85F473AF82E" blockId="11.[816,1475,234,2019]" box="[816,1127,1995,2019]" pageId="11" pageNumber="10">
|
||
<emphasis id="B95AF07CDA228526FCF3F85F408AF82E" bold="true" box="[816,983,1995,2019]" pageId="11" pageNumber="10">Measurements</emphasis>
|
||
. See
|
||
<tableCitation id="C6AC19D5DA228526FBD2F85F473CF82E" box="[1041,1121,1995,2019]" captionStart="Table 1" captionStartId="7.[113,169,974,995]" captionText="Table 1. Comparative measurements of adults of Neobuthus opatovae sp. n. and N. dhobo sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="11" pageNumber="10">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF517CE6DA258521FFB7F93547EFF97B" ID-DOI="http://doi.org/10.5281/zenodo.14660788" ID-Zenodo-Dep="14660788" box="[116,1202,1697,1719]" httpUri="https://zenodo.org/record/14660788/files/figure.png" pageId="12" pageNumber="11" startId="12.[116,193,1697,1719]" targetBox="[144,1436,304,1672]" targetPageId="12" targetType="figure">
|
||
<paragraph id="8B912C6EDA258521FFB7F93547EFF97B" blockId="12.[116,1202,1697,1719]" box="[116,1202,1697,1719]" pageId="12" pageNumber="11">
|
||
<emphasis id="B95AF07CDA258521FFB7F935425EF97B" bold="true" box="[116,259,1697,1719]" pageId="12" pageNumber="11">Figures 46–47</emphasis>
|
||
:
|
||
<taxonomicName id="4C2E57EDDA258521FECCF9354289F97A" authorityName="Euscorpius & No." authorityYear="2024" box="[271,468,1697,1719]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="opatovae" status="sp. nov.">
|
||
<emphasis id="B95AF07CDA258521FECCF9354289F97A" box="[271,468,1697,1719]" italics="true" pageId="12" pageNumber="11">Neobuthus opatovae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2694D07DA258521FE19F9354152F97B" box="[474,527,1697,1718]" pageId="12" pageNumber="11" rank="species">
|
||
<emphasis id="B95AF07CDA258521FE19F93542ACF97B" bold="true" box="[474,497,1697,1718]" pageId="12" pageNumber="11">sp</emphasis>
|
||
.
|
||
<emphasis id="B95AF07CDA258521FE3EF9354157F97B" bold="true" box="[509,522,1697,1718]" pageId="12" pageNumber="11">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, male holotYpe, dorsal (46) and ventral (47) views. Scale bar: 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B912C6EDA258521FFB2F97C471AF82D" blockId="12.[113,772,1768,2016]" lastBlockId="12.[816,1475,1768,2016]" pageId="12" pageNumber="11">
|
||
AFFINITIES. The described features distinguish
|
||
<taxonomicName id="4C2E57EDDA258521FDBEF97D405EF8CD" authorityName="Euscorpius & No." authorityYear="2024" box="[637,771,1769,1792]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="opatovae" status="sp. n.">
|
||
<emphasis id="B95AF07CDA258521FDBEF97D405EF8CD" box="[637,771,1769,1792]" italics="true" pageId="12" pageNumber="11">N. opatovae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2694D07DA258521FFB2F89C43F3F8D2" box="[113,174,1800,1824]" pageId="12" pageNumber="11" rank="species">
|
||
<emphasis id="B95AF07CDA258521FFB2F89C43D7F8ED" bold="true" box="[113,138,1800,1824]" pageId="12" pageNumber="11">sp</emphasis>
|
||
.
|
||
<emphasis id="B95AF07CDA258521FF5BF89C43FAF8ED" bold="true" box="[152,167,1800,1824]" pageId="12" pageNumber="11">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
from all other species of the genus. TheY are confirmed bY DNA phYlogenY (paper in preparation) but we did not include it in the keY below because the female is unknown. The new species is most similar according to DNA analYsis to
|
||
<taxonomicName id="4C2E57EDDA258521FF53F81D426DF86D" box="[144,304,1928,1952]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="amoudensis">
|
||
<emphasis id="B95AF07CDA258521FF53F81D426DF86D" box="[144,304,1928,1952]" italics="true" pageId="12" pageNumber="11">N. amoudensis</emphasis>
|
||
</taxonomicName>
|
||
, which has pedipalp chela movable finger with 6 subrows (
|
||
<taxonomicName id="4C2E57EDDA258521FEE7F83D42F8F80D" authorityName="Euscorpius & No." authorityYear="2024" box="[292,421,1961,1984]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="opatovae" status="sp. n.">
|
||
<emphasis id="B95AF07CDA258521FEE7F83D42F8F80D" box="[292,421,1961,1984]" italics="true" pageId="12" pageNumber="11">N. opatovae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2694D07DA258521FE68F83C42B5F872" box="[427,488,1960,1984]" pageId="12" pageNumber="11" rank="species">
|
||
<emphasis id="B95AF07CDA258521FE68F83C4299F80D" bold="true" box="[427,452,1960,1984]" pageId="12" pageNumber="11">sp</emphasis>
|
||
.
|
||
<emphasis id="B95AF07CDA258521FE11F83C42BCF80D" bold="true" box="[466,481,1960,1984]" pageId="12" pageNumber="11">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
has movable finger with 5 subrows of primarY denticles).
|
||
<typeStatus id="549592CCDA258521FE75F85C42B6F812" box="[438,491,1992,2015]" pageId="12" pageNumber="11">TYpe</typeStatus>
|
||
localitY of
|
||
<taxonomicName id="4C2E57EDDA258521FD9CF85D4182F82D" authorityName="Euscorpius & No." authorityYear="2024" box="[607,735,1993,2016]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="opatovae" status="sp. n.">
|
||
<emphasis id="B95AF07CDA258521FD9CF85D4182F82D" box="[607,735,1993,2016]" italics="true" pageId="12" pageNumber="11">N. opatovae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2694D07DA258521FD20F85C401BF932" box="[739,838,1768,2016]" pageId="12" pageNumber="11" rank="species">
|
||
<emphasis id="B95AF07CDA258521FD20F85C41A1F82D" bold="true" box="[739,764,1992,2016]" pageId="12" pageNumber="11">sp</emphasis>
|
||
.
|
||
<emphasis id="B95AF07CDA258521FCF3F97C4062F8CD" bold="true" box="[816,831,1768,1792]" pageId="12" pageNumber="11">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is relativelY near to localities of
|
||
<taxonomicName id="4C2E57EDDA258521FB5DF97D464DF8CD" box="[1182,1296,1768,1792]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="factorio">
|
||
<emphasis id="B95AF07CDA258521FB5DF97D464DF8CD" box="[1182,1296,1768,1792]" italics="true" pageId="12" pageNumber="11">N. factorio</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2E57EDDA258521FA81F97D469FF8CD" authorityName="KovarIk" authorityYear="2019" box="[1346,1474,1768,1792]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="solegladi">
|
||
<emphasis id="B95AF07CDA258521FA81F97D469FF8CD" box="[1346,1474,1768,1792]" italics="true" pageId="12" pageNumber="11">N. solegladi</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<figureCitation id="131530EBDA258521FCA0F89C409CF8ED" box="[867,961,1800,1824]" captionStart="Figures 160–161" captionStartId="29.[114,191,2011,2033]" captionTargetBox="[113,1474,226,1992]" captionTargetId="figure-9@29.[113,1474,226,2040]" captionTargetPageId="29" captionText="Figures 160–161. Map showing confirmed distribution of Neobuthus spp. (160) with detail for Somaliland (161)." figureDoi="http://doi.org/10.5281/zenodo.14660820" httpUri="https://zenodo.org/record/14660820/files/figure.png" pageId="12" pageNumber="11">Fig. 160</figureCitation>
|
||
).
|
||
<taxonomicName id="4C2E57EDDA258521FC16F89D4714F8ED" box="[981,1097,1800,1824]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="factorio">
|
||
<emphasis id="B95AF07CDA258521FC16F89D4714F8ED" box="[981,1097,1800,1824]" italics="true" pageId="12" pageNumber="11">N. factorio</emphasis>
|
||
</taxonomicName>
|
||
has pedipalp chela movable finger with 6 subrows of primarY denticles and 5 external accessorY denticles versus 5 subrows of primarY denticles and 4 external accessorY denticles in
|
||
<taxonomicName id="4C2E57EDDA258521FBECF8FD47E8F84D" authorityName="Euscorpius & No." authorityYear="2024" box="[1071,1205,1897,1920]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="opatovae" status="sp. n.">
|
||
<emphasis id="B95AF07CDA258521FBECF8FD47E8F84D" box="[1071,1205,1897,1920]" italics="true" pageId="12" pageNumber="11">N. opatovae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2694D07DA258521FB03F8FC465FF8B2" box="[1216,1282,1896,1920]" pageId="12" pageNumber="11" rank="species">
|
||
<emphasis id="B95AF07CDA258521FB03F8FC4784F84D" bold="true" box="[1216,1241,1896,1920]" pageId="12" pageNumber="11">sp</emphasis>
|
||
.
|
||
<emphasis id="B95AF07CDA258521FB2FF8FC47A6F84D" bold="true" box="[1260,1275,1896,1920]" pageId="12" pageNumber="11">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
<taxonomicName id="4C2E57EDDA258521FACEF8FD46CEF84D" authorityName="KovarIk" authorityYear="2019" box="[1293,1427,1896,1920]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="solegladi">
|
||
<emphasis id="B95AF07CDA258521FACEF8FD46CEF84D" box="[1293,1427,1896,1920]" italics="true" pageId="12" pageNumber="11">N. solegladi</emphasis>
|
||
</taxonomicName>
|
||
has leg III of adults with 9–13 ventral macrosetae on telotarsus (
|
||
<taxonomicName id="4C2E57EDDA258521FCFAF83D40E0F80D" authorityName="Euscorpius & No." authorityYear="2024" box="[825,957,1961,1984]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="opatovae" status="sp. n.">
|
||
<emphasis id="B95AF07CDA258521FCFAF83D40E0F80D" box="[825,957,1961,1984]" italics="true" pageId="12" pageNumber="11">N. opatovae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2694D07DA258521FC05F83C475BF872" box="[966,1030,1960,1984]" pageId="12" pageNumber="11" rank="species">
|
||
<emphasis id="B95AF07CDA258521FC05F83C4082F80D" bold="true" box="[966,991,1960,1984]" pageId="12" pageNumber="11">sp</emphasis>
|
||
.
|
||
<emphasis id="B95AF07CDA258521FC33F83C40A2F80D" bold="true" box="[1008,1023,1960,1984]" pageId="12" pageNumber="11">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
has leg III of adults with 18–21 ventral macrosetae on telotarsus).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |